Lycosa impedita Simon, 1909: 187–188 Rainbow 1911: 269 Bonnet 1957: 2646 McKay 1973: 379 Allocosa impedita (Simon) Roewer 1955: 206 Trochosa expolita impedita (Simon) McKay 1979: 288 McKay 1985: 86 Artoria impedita (Simon) Framenau 2005: 283–286 Taxonomy of the wolf spider genus Artoria in Western Australia (Araneae, Lycosidae, Artoriinae) Do Prado, André W. Baptista, Renner Luiz Cerqueira Framenau, Volker W. Zootaxa 2024 2024-12-09 5547 1 1 81 H2BV (Simon, 1909) Gooseberry Hill (WA Simon 1909 [151,531,151,178] Arachnida Lycosidae Artoria Animalia Araneae 33 34 Arthropoda species impedita   Figs 15–17      Lycosa impedita Simon, 1909: 187–188, fig. 6;  Rainbow 1911: 269;  Bonnet 1957: 2646;  McKay 1973: 379.    Allocosa impedita(Simon)—  Roewer 1955: 206.    Trochosa expolita impedita(Simon)—  McKay 1979: 288(misidentification);  McKay 1985: 86.    Artoria impedita(Simon)—  Framenau 2005: 283–286, fig. 9A–F.   Type material.  holotypefemale, Gooseberry Hill, Western Australia, AUSTRALIA, 31º57’S, 116º03’E, 1905, W. Michaelsen, R. Hartmeyer, Hamburg SW Australian Expedition, Station152 ( MNHN 11076). Examined.  Other material examined ( 27 males, 24 females, 1 femalewith egg sac, 5 juv.in 32 records).  AUSTRALIA: Western Australia:  2 males, Bringo Cutting, 28º45’S, 114º50’E( WAM 98/2143–4);  7 males, 1 female, Buntine Nature Reserve, 29º58’16’’S, 116º35’05’’E( WAM T47374);  1 male, Dulyalbin Rock,  25.9 kmSE of Bodallin, 31º34’21’’S, 118º59’25’’E( WAM T108173);  1 male, Fitzgerald River National Park, St Mary River, 5 kminland, 34º10’S, 119º32’E( WAM T69961);  1 female, near Holt Rock, 32º40’S, 119º24’E( WAM T51402); 1 male, N of Koorda-Bullfinch Road, W of Warrachupin Road, 31º00’35’’S, 118º42’02’’E( WAMT75885) ; 1 female, 1 juv., KwolyinHill, 31º51’28’’S, 117º46’54’’E( WAM T58487) ; 1 female, ca.  15 kmNW Kwolyin, Munyard Road, 31º51’56’’S, 117º36’39’’E( WAM T58485) ; 1 female, same locality ( WAM T162499) ; 3 males, 3 females, 1 juv., Lake Ninan Shire Reserve, East, 30º57’9’’S, 116º39’29’’E( WAM T47367) ; 1 male, Mackie Creek Reserve, 31º59’33’’S, 117º01’19’’E( WAM T47370) ; 3 females, 1 juv., McDermid Rock, 32º01’S, 120º44’E( WAM T65114) ; 1 female, same locality ( WAM T65112) ; 1 male, Moorajin Soak Nature Reserve, 30º45’17’’S, 117º14’30’’E( WAM T47369) ; 1 male, 1 female, Mt Caroline Nature Reserve, 31º47’36’’S, 117º38’34’’E( WAM T58484) ; 1 female, Mt Caroline Nature Reserve, 31º47’36’’S, 117º38’34’’E( WAM T58493) ; 1 female, Mt Hampton Nature Reserve Dam, 31º45’40’’S, 119º04’21’’E( WAM T47372) ; 1 female, 1 juv., Mt Sterling Nature Reserve, 31º49’59’’S, 117º35’45’’E( WAM T99227) ; 1 female, Mt Sterling Nature Reserve, 31º50’06’’S, 117º35’42’’E( WAM T77197) ; 1 female, Mt Sterling Nature Reserve, 31º49’59’’S, 117º35’41’’E( WAM T58486) ; 2 females, Nokaning Rock, N of Nukarni West Road, ca.  26 kmNW Merredin, 31º19’52’’S, 118º04’25’’E( WAM T58489) ; 1 male, 1 female, same locality ( WAM T58490) ; 1 male, gully E of Red Hill, 31º51’S, 116º06’E( WAM T70255) ; 1 male, Rosewood Hills, 35 milesfrom Borden, “Weddehoon”, 34º04’S, 118º16’E( WAM T51484) ; 1 male, 2 females, Totadgin Rock Conservation Reserve, 31º34’15’’S, 118º12’58’’E( WAM T58488) ; 1 male, 1 juv., Wambyn Nature Reserve, 31º53’46’’S, 116º37’25’’E( WAM T47373) ; 1 male, Yarragin Rock, 31º02’S, 117º57’E( WAM T69982) ; 1 male, Yenyenning Lakes, 32º13’38’’S, 117º13’08’’E( WAM T47368) ; 1 male, same locality ( WAM T47371) ; 1 male, 1 female, Yorkrakine Rock, 31º25’S, 117º30’E( WAM T58460) ; 1 female, same locality ( WAM T58461).   Diagnosis.  A. impeditais similar to  A. aculeata  sp. nov.Males share a pedipalp with a gaping tegular apophysis, but it has a serrated distal edge in  A. impeditawhereas it has a smooth distal lob in  A. aculeata  sp. nov.( Figs 15Cvs. 2C). Females of  A. impeditashare with those of  A. aculeata  sp. nov.the presence of a scape-like process in the epigyne; however, in  A. impeditait is broader, long (folding over itself) and posteriorly truncated ( Fig 16C, D), while in  A. aculeata  sp. nov.it is shorter and more narrowed posteriorly ( Fig. 3C).   Description.For detailed description of both sexes of  A. impeditasee Framenau (2005)(see also Figs 15A–E, 16A–D).  Life history and habitat preferences.Mature males and females of  A. impeditawere largely found between April and October, suggesting reproductive activity largely in the cooler months of the year.Where habitat preferences were given on labels they included ‘granite outcrop’, ‘under granite slab’ or ‘under rock’. This suggests this species to be a habitat specialist on granite outcrops similar to  Tapetosa darwini Framenau, Main, Harvey & Waldock, 2009( Framenau et al.2009).   Distribution.Throughout the south-western Western AustraliaWheatbelt ( Fig. 17). 5282678649 MNHN W. Michaelsen & R. Hartmeyer Australia -31.95 Gooseberry Hill 1213 116.05 34 35 MNHN 11076 1 Western Australia holotype 5282679229 Australia Western Australia 34 35 1 Western Australia 5282679549 [388,1116,1873,1900] WAM Australia -28.75 Bringo Cutting 1230 114.833336 34 35 WAM 98 2 2 Western Australia 5282679426 WAM Australia -29.971111 Buntine Nature Reserve 20 116.584724 34 35 T47374 8 1 7 Western Australia 5282678833 WAM Australia -31.5725 Dulyalbin Rock 20 118.99027 34 35 T108173 1 1 Western Australia 5282678373 WAM Australia -34.166668 St Mary River 1200 119.53333 Fitzgerald River National Park 34 35 T69961 1 1 Western Australia 5282678431 [568,1278,1981,2008] WAM Australia -32.666668 Holt Rock 1209 119.4 34 35 T51402 1 1 Western Australia 5282678329 WAM -31.857779 Kwolyin 20 117.78167 35 36 T58487 2 1 5282678317 WAM -31.865557 Munyard Road 20 117.61083 35 36 T58485 1 1 5282678601 [655,1162,222,249] WAM -31.865557 Munyard Road 20 117.61083 35 36 T162499 1 1 5282678586 WAM -30.952501 Lake Ninan Shire Reserve 20 116.65806 East 35 36 T47367 7 3 3 5282679174 WAM -31.9925 Mackie Creek Reserve 20 117.02195 35 36 T47370 1 1 5282678976 [635,1432,294,321] WAM -32.016666 McDermid Rock 1213 120.73333 35 36 T65114 4 3 5282679334 WAM -32.016666 McDermid Rock 1213 120.73333 35 36 T65112 1 1 5282679531 WAM -30.754723 Moorajin Soak Nature Reserve 20 117.24166 35 36 T47369 1 1 5282678937 [251,1315,366,393] WAM -31.793333 Mt Caroline Nature Reserve 20 117.64278 35 36 T58484 2 1 1 5282679068 WAM -31.793333 Mt Caroline Nature Reserve 20 117.64278 35 36 T58493 1 1 5282678533 WAM -31.76111 Mt Hampton Nature Reserve Dam 20 119.0725 35 36 T47372 1 1 5282679562 WAM -31.833055 Mt Sterling Nature Reserve 20 117.59583 35 36 T99227 2 1 5282679214 [492,1432,474,501] WAM -31.835001 Mt Sterling Nature Reserve 20 117.595 35 36 T77197 1 1 5282678380 WAM -31.833055 Mt Sterling Nature Reserve 20 117.59473 35 36 T58486 1 1 5282678396 WAM -31.331112 Nokaning Rock 20 118.07361 Nukarni West Road 35 36 T58489 2 2 5282678789 WAM -31.331112 Nokaning Rock 20 118.07361 Nukarni West Road 35 36 T58490 2 1 1 5282678359 [491,1218,582,609] WAM -31.85 Red Hill 1214 116.1 35 36 T70255 1 1 5282679430 WAM -34.066666 Rosewood Hills 1201 118.26667 Borden 35 36 T51484 1 1 5282679105 WAM -31.570835 Totadgin Rock Conservation Reserve 20 118.21611 35 36 T58488 3 2 1 5282679133 WAM -31.89611 Wambyn Nature Reserve 20 116.62361 35 36 T47373 2 1 5282678609 [646,1338,690,717] WAM -31.033333 Yarragin Rock 1218 117.95 35 36 T69982 1 1 5282678451 WAM -32.227222 Yenyenning Lakes 20 117.21889 35 36 T47368 1 1 5282679226 [868,1317,727,753] WAM -32.227222 Yenyenning Lakes 20 117.21889 35 36 T47371 1 1 5282679323 WAM -31.416666 Yorkrakine Rock 1216 117.5 35 36 T58460 2 1 1 5282678853 [844,1306,762,789] WAM -31.416666 Yorkrakine Rock 1216 117.5 35 36 T58461 1 1