Twenty-two new species in the genus Hyphantrophaga Townsend (Diptera: Tachinidae) from Area de Conservacion Guanacaste, with a key to the species of Mesoamerica Fleming, AJ Wood, D. Monty Smith, M. Alex Dapkey, Tanya Hallwachs, Winnie Janzen, Daniel Biodiversity Data Journal 2019 7 29553 29553 urn:lsid:zoobank.org:act:E866E684-19A6-4A47-A070-0935DFA78F75 Fleming & Wood Insecta Tachinidae Hyphantrophaga CoL Animalia Hyphantrophaga nigricauda Diptera 0 29553 Arthropoda species nigricauda  Materials  Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0027908; recordedBy: D.H. Janzen, W. Hallwachs & Leonel Siezar; individualID: DHJPAR0027908; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYE145-08, 08-SRNP-70578, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Leonel; verbatimElevation: 510; verbatimLatitude: 10.9964; verbatimLongitude: -85.4019; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9964; decimalLongitude: -85.4019; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 28-Jun-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007291; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0007291; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT063-06, 05-SRNP-32268, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0193; decimalLongitude: -85.41; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 20-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0042693; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0042693; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYH451-11, 11-SRNP-31067, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9888; decimalLongitude: -85.4234; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Crambidae, Eulepte Solis15; verbatimEventDate: 20-May-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007293; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0007293; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT065-06, 05-SRNP-32270, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0193; decimalLongitude: -85.41; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 20-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007290; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0007290; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT062-06, 05-SRNP-32248, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0193; decimalLongitude: -85.41; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 16-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0036438; recordedBy: D.H. Janzen, W. Hallwachs & Guillermo Pereira; individualID: DHJPAR0036438; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYD1629-09, 09-SRNP-14037, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Santa Rosa; locality: Area de Conservacion Guanacaste; verbatimLocality: Camino Borrachos; verbatimElevation: 295; verbatimLatitude: 10.8429; verbatimLongitude: -85.6161; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8429; decimalLongitude: -85.6161; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Crambidae, Conchylodesplatinalis; verbatimEventDate: 01-Aug-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035694; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0035694; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYD1075-09, 09-SRNP-41180, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Juntas; verbatimElevation: 400; verbatimLatitude: 10.9066; verbatimLongitude: -85.2878; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9066; decimalLongitude: -85.2878; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 01-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0010356; recordedBy: D.H. Janzen, W. Hallwachs & Minor Carmona; individualID: DHJPAR0010356; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAS187-06, 06-SRNP-42821, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Guaca; verbatimElevation: 400; verbatimLatitude: 10.9061; verbatimLongitude: -85.2828; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9061; decimalLongitude: -85.2828; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Crambidae, Pleuroptya Solis01; verbatimEventDate: 31-Aug-2006; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0011461; recordedBy: D.H. Janzen, W. Hallwachs & Jose Perez; individualID: DHJPAR0011461; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAQ848-06, 01-SRNP-5433, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Quebrada Escondida; verbatimElevation: 420; verbatimLatitude: 10.8993; verbatimLongitude: -85.2749; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.8993; decimalLongitude: -85.2749; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 19-Aug-2001; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0042303; recordedBy: D.H. Janzen, W. Hallwachs & Edwin Apu; individualID: DHJPAR0042303; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYH067-11, 11-SRNP-69487, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9408; decimalLongitude: -85.3177; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 07-Mar-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007294; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0007294; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT066-06, 05-SRNP-32262, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0193; decimalLongitude: -85.41; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 17-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0027944; recordedBy: D.H. Janzen, W. Hallwachs & Roster Moraga; individualID: DHJPAR0027944; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYE181-08, 08-SRNP-21716, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0137; decimalLongitude: -85.4253; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 17-Jul-2008; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0035815; recordedBy: D.H. Janzen, W. Hallwachs & Noe Castillo; individualID: DHJPAR0035815; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYD1196-09, 09-SRNP-69335, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9408; decimalLongitude: -85.3177; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 13-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007295; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0007295; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT067-06, 05-SRNP-32276, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0193; decimalLongitude: -85.41; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 14-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0042702; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0042702; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYH460-11, 11-SRNP-31063, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9888; decimalLongitude: -85.4234; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Crambidae, Eulepte Solis15; verbatimEventDate: 19-May-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0036606; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0036606; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYE1517-09, 09-SRNP-30852, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Mismo; verbatimElevation: 680; verbatimLatitude: 10.9876; verbatimLongitude: -85.4197; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9876; decimalLongitude: -85.4197; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Thyrididae, Dysodia Janzen10; verbatimEventDate: 24-Sep-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0042558; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0042558; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYH316-11, 11-SRNP-31057, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9888; decimalLongitude: -85.4234; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Crambidae, Eulepte Solis15; verbatimEventDate: 16-May-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007289; recordedBy: D.H. Janzen, W. Hallwachs & Lucia Rios; individualID: DHJPAR0007289; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT061-06, 05-SRNP-22093, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0137; decimalLongitude: -85.4253; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 01-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0045600; recordedBy: D.H. Janzen, W. Hallwachs & Freddy Quesada; individualID: DHJPAR0045600; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ACGAZ789-11, 11-SRNP-31069, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Sendero Laguna; verbatimElevation: 680; verbatimLatitude: 10.9888; verbatimLongitude: -85.4234; verbatimCoordinateSystem: Decimal; decimalLatitude: 10.9888; decimalLongitude: -85.4234; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Crambidae, Eulepte Solis15; verbatimEventDate: 18-May-2011; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007292; recordedBy: D.H. Janzen, W. Hallwachs & Calixto Moraga; individualID: DHJPAR0007292; individualCount: 1; sex: female; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT064-06, 05-SRNP-32272, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste; verbatimLocality: Pasmompa; verbatimElevation: 440; verbatimLatitude: 11.0193; verbatimLongitude: -85.41; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0193; decimalLongitude: -85.41; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 16-Jul-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0037607; recordedBy: D.H. Janzen, W. Hallwachs & Elieth Cantillano; individualID: DHJPAR0037607; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASHYC4352-10, 09-SRNP-21421, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Puente Mena; verbatimElevation: 280; verbatimLatitude: 11.0456; verbatimLongitude: -85.4574; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0456; decimalLongitude: -85.4574; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 23-Jul-2009; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0007313; recordedBy: D.H. Janzen, W. Hallwachs & Elieth Cantillano; individualID: DHJPAR0007313; individualCount: 1; sex: male; lifeStage: adult; preparations: pinned; otherCatalogNumbers: ASTAT085-06, 05-SRNP-22072, BOLD:AAA1929; Taxon: scientificName: Hyphantrophaganigricauda; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Hyphantrophaga; specificEpithet: nigricauda; scientificNameAuthorship: Fleming & Wood, 2017; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Del Oro; locality: Area de Conservacion Guanacaste; verbatimLocality: Monte Cristo; verbatimElevation: 525; verbatimLatitude: 11.0137; verbatimLongitude: -85.4253; verbatimCoordinateSystem: Decimal; decimalLatitude: 11.0137; decimalLongitude: -85.4253; Identification: identifiedBy: AJ Fleming; dateIdentified: 2017; Event: samplingProtocol: Reared from the larva of the Geometridae, Cyclomiadisparilis; verbatimEventDate: 28-Jun-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen  Description Male (Fig. 22). Length: 4-7 mm. Head (Fig. 22b): vertex 1/5 of head width; two reclinate upper orbital setae; ocellar setae arising beside anterior ocellus; ocellar triangle gold, concolorous with fronto-orbital plate; fronto-orbital plate gold over 90% of surface, sparsely setulose, setulae not extending below lowest frontal seta; parafacial shiny silver and bare; facial ridge bare; pedicel black, concolorous with postpedicel; arista black, very minutely pubescent, distinctly thickened on basal 1/3-1/4; palpus yellow and haired. Thorax (Fig. 22a, c): gold tomentose dorsally, grey tomentose laterally; four prominent dorsal vittae, outermost two broken across suture, innermost pair unbroken across suture, reaching 3rd postsutural dorsocentral seta; postpronotum with 3-4 setae arranged in a triangle; chaetotaxy: acrostichal setae 3:3; dorsocentral setae 3:4; intra-alar setae 2:3; supra-alar setae 2:3; three katepisternal setae, basal seta extremely weak and anterior to suture; lateral scutellar setae subequal in length to apical setae, curving inwards medially; lateral scutellar setae subequal in length to basal setae; subapical setae strongest and longest of marginal scutellar setae, these not strongly divergent; apical scutellar setae short and relatively weak, erect and crossed midway; one pair of discal scutellar setae; scutellum with a darkened crescent across basal 30%, remainder concolorous with scutum. Legs (Fig. 22c): black throughout; hind coxa setose. Wing (Fig. 22a): pale grey translucent, slightly infuscate, veins dark amber brown; vein R4+5 with two setulae at base. Abdomen (Fig. 22a, c): ground colour black; mid-dorsal depression on ST1+2 reaching hind margin; abdominal tomentum present as gold bands spanning anterior 50% of T3-T4; T5 gold tomentose with black apex; median marginal setae present on ST1+2-T3; a complete row of marginal setae present on T4; discal setae present on T3-T5; sex patch absent. Terminalia (Fig. 22d, e, f): anterior margin of sternite 5 (Fig. 22f) with a slight curved medial depression, posterior edge with a deeply excavated median cleft, smoothly U-shaped; posterior lobes of sternite rounded apically and completely devoid of setae; apical plate shorter than posterior lobes. Unsclerotised "window" anterior to median cleft with central rectangular gap as wide as median cleft, branched on either side, with two thin arms reaching around base of posterior lobes. Cerci (Fig. 22d) in posterior view subrectangular and slightly shorter than surstyli, blunt and squared off towards apex, completely separate medially, diverging at tips; in lateral view (Fig. 22e) with a strong, almost 90 degree bend 1/3 along their length apically; densely setulose almost to apex dorsally, setulose ventrally along basal 2/3 (visible in lateral view). Surstylus in lateral view almost parallel-sided along its length, ending in a slightly downcurved apex, making the structure appear blade-like; surstylus appearing fused with epandrium; when viewed dorsally, surstyli appearing strongly divergent and wide open. Pregonite broad and well-developed, bent basally, apical 2/3 cinched, giving it a petiolate appearance, apically rounded, devoid of setulae. Postgonite elongate, slender, very slightly hooked at tip, subequal in length to pregonite. Distiphallus cone-shaped, apically flared, with a slender median longitudinal sclerotised reinforcement on its posterior surface and a broad, anterolateral, sclerotised acrophallus on each side, joining the plate of opposite side on anterior surface near apex; approximately two times as long as basiphallus. Female. Length: 5-7 mm. As male, differing only by the presence of two pairs of proclinate orbital setae.  Diagnosis  Hyphantrophaga nigricaudasp. n. can be distinguished from all other Hyphantrophagaspecies by the following combination of traits: pedicel and arista concolorous, dark brown/black, four postsutural dorsocentral setae, four thoracic vittae, inner pair solid across suture, hind coxa setose, discal setae present on T3-T5 and T5 gold tomentose with a black apex.  Etymology From the Latin adjective "nigrum"meaning black and the Latin noun "cauda" meaning tail, in reference to the apically black T5.  Distribution Costa Rica, ACG, Alajuela and Guanacaste Provinces, 280-680 m elevation.  Ecology  Hyphantrophaga nigricaudasp. n. has been reared 22 times from five species of Lepidopterain the families Geometridae, ( Cyclomia disparilisSchaus, 1911; Crambidae: PleuroptyaSolis02, EulepteSolis15 and Conchylodes platinalis( Guenee, 1854)); and Thyrididae, ( DysodiaJanzen10); in rain forest and dry-rain lowland intergrade. 2008-06-28 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Leonel Siezar Costa Rica 10.9964 Area de Conservacion Guanacaste -85.4019 0 29553 DHJPAR 0027908 1 1 Holotype 2005-07-20 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Calixto Moraga Costa Rica 11.0193 Area de Conservacion Guanacaste -85.41 0 29553 DHJPAR 0007291 1 1 Paratype 2011-05-20 Reared from the larva of the Crambidae, Eulepte Solis 15 CNC, Insects D. H. Janzen, W. Hallwachs & Freddy Quesada Costa Rica 10.9888 Area de Conservacion Guanacaste -85.4234 0 29553 DHJPAR 0042693 1 1 Paratype 2005-07-20 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Calixto Moraga Costa Rica 11.0193 Area de Conservacion Guanacaste -85.41 0 29553 DHJPAR 0007293 1 1 Paratype 2005-07-16 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Calixto Moraga Costa Rica 11.0193 Area de Conservacion Guanacaste -85.41 0 29553 DHJPAR 0007290 1 1 Paratype 2009-08-01 Reared from the larva of the Crambidae, Conchylodesplatinalis CNC, Insects D. H. Janzen, W. Hallwachs & Guillermo Pereira Costa Rica 10.8429 Area de Conservacion Guanacaste -85.6161 0 29553 DHJPAR 0036438 1 1 Paratype 2009-07-01 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Jose Perez Costa Rica 10.9066 Area de Conservacion Guanacaste -85.2878 0 29553 DHJPAR 0035694 1 1 Paratype 2006-08-31 Reared from the larva of the Crambidae, Pleuroptya Solis 01 CNC, Insects D. H. Janzen, W. Hallwachs & Minor Carmona Costa Rica 10.9061 Area de Conservacion Guanacaste -85.2828 0 29553 DHJPAR 0010356 1 1 Paratype 2001-08-19 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Jose Perez Costa Rica 10.8993 Area de Conservacion Guanacaste -85.2749 0 29553 DHJPAR 0011461 1 1 Paratype 2011-03-07 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Edwin Apu Costa Rica 10.9408 Area de Conservacion Guanacaste -85.3177 0 29553 DHJPAR 0042303 1 1 Paratype 2005-07-17 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Calixto Moraga Costa Rica 11.0193 Area de Conservacion Guanacaste -85.41 0 29553 DHJPAR 0007294 1 1 Paratype 2008-07-17 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Roster Moraga Costa Rica 11.0137 Area de Conservacion Guanacaste -85.4253 0 29553 DHJPAR 0027944 1 1 Paratype 2009-07-13 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Noe Castillo Costa Rica 10.9408 Area de Conservacion Guanacaste -85.3177 0 29553 DHJPAR 0035815 1 1 Paratype 2005-07-14 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Calixto Moraga Costa Rica 11.0193 Area de Conservacion Guanacaste -85.41 0 29553 DHJPAR 0007295 1 1 Paratype 2011-05-19 Reared from the larva of the Crambidae, Eulepte Solis 15 CNC, Insects D. H. Janzen, W. Hallwachs & Freddy Quesada Costa Rica 10.9888 Area de Conservacion Guanacaste -85.4234 0 29553 DHJPAR 0042702 1 1 Paratype 2009-09-24 Reared from the larva of the Thyrididae, Dysodia Janzen 10 CNC, Insects D. H. Janzen, W. Hallwachs & Calixto Moraga Costa Rica 10.9876 Area de Conservacion Guanacaste -85.4197 0 29553 DHJPAR 0036606 1 1 Paratype 2011-05-16 Reared from the larva of the Crambidae, Eulepte Solis 15 CNC, Insects D. H. Janzen, W. Hallwachs & Freddy Quesada Costa Rica 10.9888 Area de Conservacion Guanacaste -85.4234 0 29553 DHJPAR 0042558 1 1 Paratype 2005-07-01 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Lucia Rios Costa Rica 11.0137 Area de Conservacion Guanacaste -85.4253 0 29553 DHJPAR 0007289 1 1 Paratype 2011-05-18 Reared from the larva of the Crambidae, Eulepte Solis 15 CNC, Insects D. H. Janzen, W. Hallwachs & Freddy Quesada Costa Rica 10.9888 Area de Conservacion Guanacaste -85.4234 0 29553 DHJPAR 0045600 1 1 Paratype 2005-07-16 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Calixto Moraga Costa Rica 11.0193 Area de Conservacion Guanacaste -85.41 0 29553 DHJPAR 0007292 1 1 Paratype 2009-07-23 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Elieth Cantillano Costa Rica 11.0456 Area de Conservacion Guanacaste -85.4574 0 29553 DHJPAR 0037607 1 1 Paratype 2005-06-28 Reared from the larva of the Geometridae, Cyclomiadisparilis CNC, Insects D. H. Janzen, W. Hallwachs & Elieth Cantillano Costa Rica 11.0137 Area de Conservacion Guanacaste -85.4253 0 29553 DHJPAR 0007313 1 1 Paratype