New taxonomic and faunistic data on the funnel-weavers (Araneae, Agelenidae) of Turkiye and the Caucasus, with five new species Zamani, Alireza Kaya, Rahşen S. Marusik, Yuri M. ZooKeys 2024 2024-11-21 1218 251 286 Zamani & Kaya & Marusik 2024 Arachnida Agelenidae Maimuna Animalia Araneae Arthropoda species antalyensis sp. nov.   Figs 1 A, B, 2 A – C, 3 A – D, 4 A – D   Type material.    Holotype• ♂(  ZMUT), Turkiye: Antalya Prov.: Alanya, env. Kestel, Dim Valley,  36°32'34.5"N, 32°06'17.5"E,  110 m, pine and oak forest,  2.01.2013( Y. M. Marusik).   Paratypes: • 5 ♀(  ZMUT), same data as for the holotype; •  1 ♀(  ZMUT), Asmaca,  36°36'32.3"N, 32°03'12.4"E,  686 m, pine and oak forest,  3.01.2013( Y. M. Marusik); •  2 ♀(  ZMUT), Elikesik rd.,  36°33'55.6"N, 31°55'30.3"E,  24 m, maquis on S exposed slope,  8.01.2013( Y. M. Marusik); •  2 ♀(  ZMUT), slopes of Alanya Castle, Damlataş side,  36°32'11.6"N, 31°59'30.3"E,  50 m, pine forest, under stones and in litter,  7.01.2013( Y. M. Marusik).   Comparative material.   Maimuna vestita(C. L. Koch, 1841):  Turkiye: Bursa Prov.: • 1 ♀(  ZMUU), Bursa Uludağ University campus area,  30.11.1999( R. S. Kaya); •  2 ♀(  ZMUU), same,  16.05.2000( R. S. Kaya); •  1 ♀(  ZMUU), same,  3.03.2003( R. S. Kaya); •  1 ♀(  ZMUU), same,  10.09.2005( R. S. Kaya); •  1 ♂ 1 ♀(  ZMUU), same,  20.04.2012( R. S. Kaya); •  1 ♂ 1 ♀(  ZMUU), same,  21.05.2012( R. S. Kaya); •  5 ♀(  ZMUU), same,  4.05.2023( R. S. Kaya); •  2 ♂ 3 ♀(  ZMUU), same,  4.01.2024( R. S. Kaya); •  1 ♂ 2 ♀(  ZMUU), same,  11.07.2024( R. S. Kaya); •  5 ♀(  ZMUU), Lake Uluabat, Halilbey Island,  8.04.2001( R. S. Kaya); •  1 ♂ 3 ♀(  ZMUU), Lake Uluabat, Terzioğlu Island,  14.10.2004( R. S. Kaya); •  5 ♀(  ZMUU), same,  25.04.2005( R. S. Kaya); •  1 ♂ 2 ♀(  ZMUU), Lake Uluabat, Manastır Island,  28.09.2005( R. S. Kaya); •  1 ♂(  ZMUU), same,  29.09.2005( R. S. Kaya); •  2 ♂ 1 ♀(  ZMUU), Lake Uluabat, Halilbey Island,  15.12.2005( R. S. Kaya); •  1 ♀(  ZMUU), Karacabey, Boğaz,  4.05.2005( R. S. Kaya); •  3 ♀(  ZMUU), same,  8.08.2007( R. S. Kaya); •  2 ♀(  ZMUU), same,  5.06.2018( R. S. Kaya); •  2 ♀(  ZMUU), same,  11.06.2021( R. S. Kaya); •  1 ♂ 1 ♀(  ZMUU), Kaplıkaya,  29.03.2007( R. S. Kaya); •  2 ♂ 2 ♀(  ZMUU), same,  10.12.2008( R. S. Kaya); •  1 ♀(  ZMUU), Görükle Vill.,  20.04.2012( R. S. Kaya); •  1 ♀(  ZMUU), Nilüfer Metro station,  26.05.2012( R. S. Kaya); •  1 ♂(  ZMUU), Orhangazi Dist.,  4.05.2014( R. S. Kaya);  Çanakkale Prov.: • 1 ♀(  ZMUU), Gökçeada Island, Lake salt area,  4.05.2004( R. S. Kaya).   Diagnosis. The new species is similar to  M. cariaeBrignoli, 1978 inthe overall shape of its copulatory organs. The male differs by having a shorter tip of the cymbium (as long as the palpal tibia, vs longer), and by a different shape of the conductor and the median process (cf. Fig. 3 B – D and Dimitrov 2022: fig. 26). The female of the new species has a hexagonal epigynal fovea, in contrast to the subtriangular fovea of  M. cariae(cf. Fig. 2 A, Band Dimitrov 2022: fig. 17).   Figure 3. Male palp of  Maimuna antalyensis sp. nov. Aventral view Bretroventral view Cfemur, ventral view Dretrolateral view. Abbreviations:  Eb– base of the embolus,  Em– embolus,  Cn– conductor,  Mp– median process, Vc– retroventral crest of the tibia. Scale bars: 0.2 mm.   Description.  Male.Habitus as in Fig. 1 A. Total length 5.40. Carapace 2.60 long, 1.90 wide. Eye sizes: AME: 0.10, ALE: 0.13, PME: 0.18, PLE: 0.13. Carapace, sternum, labium, and maxillae pale brown; carapace with darker submedian bands; ocular region black. Legs yellowish brown, with annulations. Abdomen dorsally dark greyish with paler chevrons, pale greyish ventrally. Spinnerets pale greyish, darker basally. Measurements of legs: I: 5.81 + missing Ta(1.74, 0.82, 1.43, 1.82, missing), II: 6.73 (1.77, 0.82, 1.37, 1.73, 1.04), III: 6.64 (1.74, 0.76, 1.34, 1.74, 1.06), IV: 8.53 (2.19, 0.85, 1.80, 2.55, 1.14). Palp as in Figs 3 A – D, 4 A – D; femur modified – with retroventral bump in mid part, ~ 3 × longer than wide, shorter than cymbium; dorsal length of patella same as in tibia; tibia ~ 1.4 × wider than long (Fig. 3 C, D), lacking prominent apophysis but with retroventral crest ( Vc) (Fig. 3 A, B); cymbium ~ 1.4 × longer than wide, with tip ~ 1 / 3 of cymbial length; bulb as long as wide; conductor (  Cn) massive, strongly sclerotized prolateral arm lacking, terminal part with two claw-like projections ( Cc; Fig. 4 D); median process (  Mp) massive, roundly bent in lateral view (Fig. 4 B, C); embolus (  Em) filiform, weakly sclerotized, with small base (  Eb).   Figure 4. Dissected bulb of  Maimuna antalyensis sp. nov. Aproximal view Bventral view Cproventral view Dretroventral view. Abbreviation: Cc– claw-like projections of the conductor. Scale bars: 0.2 mm.   Figure 5. Epigyne of  Tegenaria hamid. Aintact, ventral view Bmacerated, anteroventral view Cvulva, dorsal view Dvulva, anterodorsal view. Scale bars: 0.2 mm.  Female.Habitus as in Fig. 1 B. Total length 7.50. Carapace 3.28 long, 2.07 wide. Eye sizes: AME: 0.12, ALE: 0.17, PME: 0.20, PLE: 0.15. Coloration as in male. Measurements of legs: I: 7.25 (1.94, 1.02, 1.44, 1.76, 1.09), II: 7.24 (1.85, 1.05, 1.47, 1.73, 1.14), III: 7.37 (2.00, 0.96, 1.42, 1.94, 1.05), IV: 9.58 (2.45, 1.11, 2.13, 2.66, 1.23). Epigyne as in Fig. 2 A – C; epigynal plate slightly wider than long; fovea (  Fo) hexagonal, approximately as long as wide, located anteriorly (Fig. 2 C); copulatory ducts gradually turning to receptacles and approximately as wide as receptacles; copulatory ducts converging and contiguous, receptacles diverging (Fig. 2 A, B).   Comment. In the examined comparative female specimens of  M. vestita, we observed noticeable variation in both body size and epigyne morphology. In particular, in smaller individuals, the shape of the epigynal fovea can vary significantly. A similar pattern is observed in  M. antalyensis sp. nov., where smaller females also exhibit variation in the shape of the epigynal fovea, ranging from hexagonal to nearly circular in some specimens.   Note. For comments on the homology of the structure referred to here as the “ median process, ” see Kaya et al. (2023).   Distribution. Known from the listed localities in Antalya Province, southwestern Turkiye.   Etymology. The specific epithet refers to the type locality of the species in Antalya, Turkiye. 4981553347 pine and oak forest 2013-01-02 ZMUT Y. M. Marusik Turkey Alanya 110 36.542915 Dim Valley 1 32.10486 env. Kestel 1 1 Antalya Prov. holotype 4981553352 pine and oak forest 2013-01-02 ZMUT Y. M. Marusik Turkey Alanya 110 36.542915 Dim Valley 1 32.10486 Kestel 5 5 Antalya paratype 4981553312 pine and oak forest 2013-01-03 ZMUT Y. M. Marusik Turkey 686 36.60897 Asmaca 1 32.053444 1 1 Antalya paratype 4981553346 maquis on S exposed slope 2013-01-08 ZMUT Y. M. Marusik Turkey 24 36.565445 Elikesik rd. 1 31.925083 2 2 Antalya paratype 4981553316 pine forest, under stones and in litter 2013-01-07 ZMUT Y. M. Marusik Turkey 50 36.536556 slopes of Alanya Castle, Damlatas side 1 31.99175 Damlatas 2 2 Antalya paratype 4981553325 1999-11-30 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 1 1 Bursa Prov. 4981553317 2000-05-16 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 2 2 Bursa 4981553303 2003-03-03 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 1 1 Bursa 4981553304 2005-09-10 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 1 1 Bursa 4981553345 2012-04-20 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 2 1 1 Bursa 4981553348 2012-05-21 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 2 1 1 Bursa 4981553309 2023-05-04 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 5 5 Bursa 4981553361 2024-01-04 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 5 3 2 Bursa 4981553375 2024-07-11 ZMUU R. S. Kaya Turkey Bursa Uludag University campus area 3 2 1 Bursa 4981553335 2001-04-08 ZMUU R. S. Kaya Turkey Halilbey Island Lake Uluabat 5 5 Bursa 4981553353 2004-10-14 ZMUU R. S. Kaya Turkey Terzioglu Island Lake Uluabat 4 3 1 Bursa 4981553356 2005-04-25 ZMUU R. S. Kaya Turkey Terzioglu Island Lake Uluabat 5 5 Bursa 4981553330 2005-09-28 ZMUU R. S. Kaya Turkey Manastir Island Lake Uluabat 3 2 1 Bursa 4981553323 2005-09-29 ZMUU R. S. Kaya Turkey Manastir Island Lake Uluabat 1 1 Bursa 4981553350 2005-12-15 ZMUU R. S. Kaya Turkey Halilbey Island Lake Uluabat 3 1 2 Bursa 4981553326 2005-05-04 ZMUU R. S. Kaya Turkey Bogaz Karacabey 1 1 Bursa 4981553315 2007-08-08 ZMUU R. S. Kaya Turkey Bogaz Karacabey 3 3 Bursa 4981553314 2018-06-05 ZMUU R. S. Kaya Turkey Bogaz Karacabey 2 2 Bursa 4981553354 2021-06-11 ZMUU R. S. Kaya Turkey Bogaz Karacabey 2 2 Bursa 4981553302 2007-03-29 ZMUU R. S. Kaya Turkey Kaplikaya 2 1 1 Bursa 4981553313 2008-12-10 ZMUU R. S. Kaya Turkey Kaplikaya 4 2 2 Bursa 4981553332 2012-04-20 ZMUU R. S. Kaya Turkey Gorukle Vill. 1 1 Bursa 4981553322 2012-05-26 ZMUU R. S. Kaya Turkey Nilufer Metro station 1 1 Bursa 4981553351 2014-05-04 ZMUU R. S. Kaya Turkey Orhangazi Dist. 1 1 Canakkale 4981553336 2004-05-04 ZMUU R. S. Kaya Turkey Lake salt area Gokceada Island 1 1 Canakkale Prov.