Achradocera insignis Parent, 1933: 247 Taxonomic revision of Achradocera Becker (Diptera: Dolichopodidae), with description of two new species Quevedo, Lucas Capellari, Renato S. Lamas, Carlos José E. Zootaxa 2024 2024-10-09 5519 3 301 344 89TRT Parent, 1933 Parent 1933 [151,575,151,178] Insecta Dolichopodidae Achradocera Animalia Diptera 29 330 Arthropoda species insignis  ( Figs 17, 18, 26)      Achradocera insignis Parent, 1933: 247, figs 4–5.  Type-locality: Tapikiolé, Argentina. Lectotypemale, SNMS.   Diagnosis (male).Postpedicel longer than arista-like stylus; lower postoccipital setae dense. Legs mostly yellow, except for FIII, dorsal surface of FI and FII, TII at apical half, and distal tarsi, which are all brown infuscated; TII ventrally arched near the middle, and apex bearing two rows of about 4 setae each at av and pv surfaces; fore and hind legs simple. Hypopygium small; surstylus subtriangular, tapered and bearing 1 apical and 1 subapical strong seta; postgonite enlarged and covered by small setae.   Redescription.Similar to  A. angustifacies  stat. rev., except as noted. Male.Body length: 2.15–2.25 mm. Wing: 2.0– 2.1 mmlong, 0.64–0.67 mmwide. Antenna.Postpedicel slightly longer than arista-like stylus. FAr ratio, 1.12. Thorax.5 ac and 5 dc pairs of setae. Wings.CuAx: 0.55; RMx: 1.33. Lower calypter yellow with fan of about 6–7 brownish setae; halter pale yellow. Leg I.38, 36, 15, 6, 4, 3, 4. F with posterior row of about 4 setae on apical 1/4; T bearing ventral row with distinct setae and distinct dorsal seta at basal 1/4. Leg II.39, 40, 20, 9, 6, 4, 3. F bearing pv row of about 3 conspicuous setae near apex; T brownish infuscated at apical half, ventrally arched near middle, and apex bearing 2 rows of about 4 setae each at av and pv surfaces ( Fig. 17D). Leg III.44, 50, 15, 9, 6, 4, 3. F with about 2 anterior and 2 posterior setae near apex; T yellow, bearing 3 pd, 2 ad, and 2 apical setae (1 av, 1 v); t 3–4simple. Hypopygium.As in Fig. 18A–D, similar to  A. angustifacies  stat. rev.  Female.Unknown.   Distribution. Argentina, Paraguay, Brazil, and Bolivia( Fig. 26).   Type material examined.   LECTOTYPE[here designated] 1♂, labelled: “Tapikiolé-Arg. |  XI.25–1.26Lind | D. Chaco-Exped”, “  Achradocera insignis n. sp.| O Parent. Type” [handwritten], “SMNS_Dip_ | 007101” [ SMNS].   PARALECTOTYPE: 1♀: “ VIII.25.Trini- | dad b. Asuncion| D. Chaco-Exped”, “  Achradocera insignisPar.” [handwritten], “SMNS_Dip_ | 007134” [ SMNS].  Additional material examined.   Brazil. Goiás. 1♂, “ JATAI, Goiaz,  I.1955| M. Carrera, A. Machado| F. S. Pereira, E. Dente| Milgar Loureirocol”;  1♂, “BRASIL, GO, Bom Jesusde Goiás| Fazenda Sta. Rosa do Meia Ponte| 18º19′42.7″S 49º36′08.3″W| Malaise trap;  20–22.xi.2021| PR Riccardileg.” [all in MZUSP].  Bolivia. 1♂, “ BOLIVIA: Santa Cruz Prov.| Campo Guairuy 24 kmS | Camiri,  18.VIII–2.IX 2000| S20º11.81′; W63º28.55′| malaise trap,  870 m| M.E. Irwin& M. Hauser”. [ MTEC].   Remarks.According to the data collected thus far,  A. insignisappears to have a restricted distribution along the eastern South American’s Dry Diagonal ( Luebert 2021), occurring in Chacoand Cerrado areas, which contrasts with most other  Achradoceraspeciesthat appear to be quite generalists for environments, occurring over larger biogeographical areas. Another noteworthy feature is that  A. insignishas a sympatric distribution with other  Achradoceraspecies, such as  A. gimli  sp. nov.and  A. tuberculata, making female association nearly impossible based only on morphology. Because of this issue, we have chosen to highlight the females which are not readily associated with males in a separate section below (see “Unassociated females”) to avoid mistakes. The female paratypeof  A. insigniswas not described or considered in the distribution map despite its type status because it is from a different area than the male holotypeand we are uncertain that it is conspecific. 4953412425 1925-11 SMNS D. Chaco-Exped 30 331 1 1 lectotype 4953412388 SMNS D. Chaco-Exped Asuncion 30 331 1 1 Asuncion paralectotype 4953412411 1955-01 2021-11-22 1955-01 JATAI, MZUSP M. Carrera & A. Machado & F. S. Pereira & E. Dente & Milgar Loureiro & Riccardi Brazil Goiaz -18.32853 Fazenda Sta. Rosa do Meia Ponte 1 -49.602303 Bom Jesus 30 331 2 2 Goias 4953412312 1955-01 2021-11-22 1955-01 JATAI, MZUSP M. Carrera & A. Machado & F. S. Pereira & E. Dente & Milgar Loureiro & Riccardi Brazil Goiaz -18.32853 Fazenda Sta. Rosa do Meia Ponte 1 -49.602303 Bom Jesus 30 331 2 2 Goias 4953412427 2000-08-18 2000-09-02 2000-08-18 MTEC M. E. Irwin & M. Hauser Bolivia Goiaz 870 true -20.196833 Camiri 12 -63.475834 Campo Guairuy 30 331 1 1 Santa Cruz