Tonnoiriella paveli Ježek, 1999 The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae) Wagner, Rüdiger Withers, Phil Zootaxa 2020 2020-01-22 4728 2 183 210 7CHTT Jezek, 1999 Jezek 1999 [151,530,552,579] Insecta Psychodidae Tonnoiriella Animalia Diptera 6 189 Arthropoda species paveli  ( Figs. 10, 11)    Tonnoiriella paveli Ježek, 1999: Časopis Národniho Muzea v Praze Řada přirodovedna 168: 10.   Material:  1♂ holotype, slide mounted: head, antennae (1), palpi (2), thorax (2), wings (2), halter (1), legs (4), ab- domen (1), styli (2), aedeagus complex (1), epandrium and epandrial processes (1); left side white label: Morocco, 8278, High Atlas Mts., a stream near a wet rocky wall S of Tizi-n-Testapproximately  1700 ma.s.l.,  2.–3. June 1985, Chvojkaleg.; near forest brook with close undergrowth. Rightside red label:  Tonnoiriella paveliJežek 1989; holo- type Cat. No.33802, 1996 male. 2♂(nos 8279, 8280; Cat. Nos. 33803, 33804) with holotype.  1♂ paratype, left side white label: Morocco, 8283, High Atlas Mts., Toubkai Massif, a streamlet, rocks NW of LacIfni (  2.500 ma.s.l.),  3.October 1994, Chvojkaleg.; right side red label:  Tonnoiriella paveliJežek 1989, paratype, Cat. No33807, 1999 male.  1♂ paratype, left side white label: Morocco, 8285; NE of Anto-Atlas Mts, a streamlet of a semidesert area;  1.800 ma.s.l.N of Tizi-n-Bachkoun,  1.June 1985 Chvojkaleg. Right side red label:  Tonnoiriella paveliJežek 1989, paratype, Cat. No33808, 1999 male (all in collection NMPC).   Description:Head round, eyebridge of 3 facet rows; distance between eyes 4 facet diameters; postocular bristles present. Palpus segments absolute length: 0.06-0.07 - 0.096 -0.166mm; relative length: 30-35-48-83. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.042 -0.054-0.04-0.04-0.04-0.04-0.04-0.044-0.046-0.044- 0.036-0.034-0.03-0.02 -0.024 - 0.028mm; relative length 21-27-20-20-20-20-20-22-23-22- 18-17-15-10-12-14; terminal flagellomere with apiculus. Pairs of digitate ascoids on flagellomeres 5–11. Wing 3.18 times longer than wide; length 1.84 mm, width 0.58 mm. Male terminalia ( Fig. 10) with hypandrium medially broadened. Gonocoxites are approximately 2.5x longer than wide with a long seta at about middle. Gonostyli about as long as gonocoxites, almost straight. Parameres difficult to identify, small. Aedeagus with dorsal lobe of the basiphallus terminating in a broad sclerite laterally with elongate spine on one side, and a less sclerotized area on the other side ( Fig. 11). Ventral arm bilobed, short lobe connecting at about middle to the lateral side with the distiphallus, long lobe almost straight, tip at about tip of gonocoxite. The ventral distiphallus is a large strongly S-shaped sclerite, margin of baso-lateral end runs parallel to the margin of the dorsal basiphallus lobe; laterally with a tip, in the basal third with a small lobe connecting to the ventral lobe of the basiphallus; the long cross arm almost straight with blunt tips. The dorsal distiphallus has two slightly sinuous lateral lobes with sharp tips; aedeagal sheath with numerous wrinkles. Epandrium is rectangular; epandrial processes almost straight longer than epandrium with 11 tenacula, length of latter decreasing apically.   Distribution: North Africa, Morocco.   Remarks:  T. paveliis a close relative of  T. pulchradue to the shape of the ventral distiphallus, bilobed on both sides; the dorsal distiphallus has two large lateral acute tips. The shape of the median part of the hypandrium was not clearly discernable and may differ. 2556505434 1985-06-02 1985-06-03 1985-06-02 Chvojka & Right & Cat. No. Morocco 1700 High Atlas Mts. 6 189 1 1 holotype 2556505450 1994-10-03 Chvojka Morocco Cat. No 3 High Atlas Mts. Toubkai Massif 6 189 2 1 paratype 2556505428 1985-06-01 NMPC Chvojka Morocco 2 Tizi-n-Bachkoun Cat. No 6 189 2 1 paratype