A taxonomic revision of the genus Noronhia Stadtm. ex Thouars (Oleaceae) in Madagascar and the Comoro Islands
Hong-Wa, Cynthia
Boissiera
2016
2016-10-21
70
1
292
8WYRV
Hong-Wa
2016
[503,748,380,408]
Magnoliopsida
Oleaceae
Noronhia
Plantae
Lamiales
233
233
Tracheophyta
species
rollandii
Typus: MADAGASCAR. Prov. Toamasina: Alaotra-Mangoro, Moramanga, Ambatovy, Ampitabe, piste direction S du campement Dynatec, 18°51’34”S 48°17’13”E, 1080 m, 18. I .2005, Ranaivojaonaet al. 1151(holo-: MO-6615552!; iso-: G[ G00341615]!, K!, P!, TAN!).
Diagnosis Noronhia rollandiiHong-Wa can be distinguished from other members of the genus by its non-woody petioles, its opposite to verticillate, oblong leaf blades, its urceolate, yellow flowers, and its ovoid, punctate fruits.
Description Treesto 15 mtall, trunk to 24 cmdiameter; young twigs cylindrical to somewhat flattened, 1.1-2.3 mmdiameter, glabrous; bark dark gray, smooth, with scattered lenticels. Leavesopposite, verticillate at the tips of the branches, persistent; bud scales deciduous; blades light green above, darker below, oblong, 4-10 3 1.5-3 cm, coriaceous, glabrous, domatia absent, base attenuate, margin flat, apex retuse, midrib slightly sunken above, distinctly raised below, secondary veins inconspicuous, 7-12 per side, 5-15 mmapart, looping 1-3.5 mmfrom the margin; petiole brown, 4-10 3 1-3 mm, not woody or only up to mid-length, glabrous. Thyrsesgeminate, multiflorous, compact; peduncle 5-10 mmlong, moderately pubescent; pedicel 2-8 mmlong, moderately pubescent; calyx moderately pubescent outside, glabrous inside, lobes triangular, 0.8-1.5 3 1-1.8 mm; corolla yellow tinged pinkish outside, yellow inside, urceolate, 3-4.5 mmlong, glabrous on both sides, the tube 1-2 mmlong, lobes widely ovate, apex acute; corona present, 1.7-2.3 mmlong, undivided; stamens 2-2.5 mmlong, anthers oblong to obovate, 1.5-1.8 mmlong; pistil 2-2.7 mmlong, stigma capitate. Fruitingpedicel 2-8 3 1-2.5 mm; young fruits green, brownish to blackish when mature, ovoid, 12-24 3 8-17 mm, covered with white dots, apex bluntly pointed to apiculate; dry pericarp 0.3-1.4 mmthick; endocarp woody; seed 9-15.5 3 7-11.5 mm.
Etymology This name honors the late Rolland Ranaivojaona, a botanist at the Parc Botanique et Zoologique de Tsimbazaza(TAN), for collecting the typespecimen and for sharing his knowledge of the Malagasyflora.
Distribution, ecology and phenology Noronhia rollandiioccurs in mid- to high-elevation humid forests in the east, from Anjanaharibe-Sud to Mangoro ( Fig. 49). It produces flowers year-round except in December.
Conservation status The assessment included 29 collections representing 28 localities and yielded an EOO of 38,636 km 2, an AOO of 84 km 2, and 13 subpopulations representing 11 locations, of which seven occur within protected areas (Anjanaharibe-Sud, Corridor Ankeniheny-Zahamena, Makira, Mantadia, Masoala, Torotorofotsy, and Zahamena). Although the estimated AOO is rather small, the actual AOO is likely much larger and may well exceed 2,000 km 2. That, along with a large EOO, a high number of locations, and good representation within the network of protected areas, indicates a preliminary status of “Least Concern” for N. rollandii.
Notes Noronhia rollandiiis similar to N. obtusifolia, but can be distinguished by its opposite to verticillate (vs. entirely opposite), oblong (vs. obovate) leaves, urceolate (vs. campanulate), yellow tinged pinkish (vs. white) flowers, the presence (vs. absence) of a corona, and its punctate (vs. smooth to ribbed) fruits. The new species can be recognized by its non-woody petioles, opposite to verticillate, oblong leaf blades, urceolate, yellow flowers, and ovoid, punctate fruits.
Paratypi MADAGASCAR. Prov. Antsiranana: Andapa, Anjanaharibe-SudRS, 14°46’15”S 49°28’E, 1161-1424 m, 21.III-7.IV.1994, Ravelonarivo et al.91( K, MO, TAN); ibid. loc., 14°44’42”S 49°27’42”E, 1185-1335 m, 3.XI.1994, Ravelonarivo & Rabesonina484( G, K, MO, P, TAN). Prov. Fianarantsoa: IvohibeRS, 22°25’18”S 46°53’54”E, 1200 m, 10.XI.1997, Rakotomalaza1526( G, MO). Prov. Mahajanga: Befandriana-Nord, Matsoandakana, Ambotaharanana, Andranomena, 15°08’16”S 49°20’55”E, 1077 m, 12.II.2008, Bernard770( G, MO, P, TAN); ibid. loc., Andasinanantsomanga, Amparihy, 14°55’34”S 49°24’59”E, 25.II.2008, Bernard938( G, MO, P, TAN); ibid. loc., Ambotaharanana, Andranomena, forêt d’Anjiabe, 15°08’19”S 49°21’50”E, 1208 m, 14.II.2008, Ravelonarivo et al.2831( MO, P, TAN); ibid. loc., 15.II.2008, Ravelonarivo et al.2865( MO, P, TAN); Tsaravilona, Amparihy, Androva, 14°55’36”S 49°25’40”E, 1154-1178 m, 22.II.2008, Ravelonarivo2893( MO, P, TAN). Prov. Toamasina: Antanandava, Zahamena PN, 17°29’47”S 48°45’35”E, 1000-1912 m, 2.II.2002, Andrianjafyet al. 289( CNARP, K, MO, P, TEF); Maroantsetra, Mahalevona, MasoalaPN, 15°25’47”S 49°57’33”E, 1198 m, 8.X.2003, Antilahimena et al.2152( MO, P); Ambinanitelo, Ambalamahogo, Maimborondro, 15°18’39”S 49°22’43”E, 1100 m, 6.IX.2004, Antilahimena2835( MO, P); Moramanga, Ambatovy, 18°49’19”S 48°20’08”E, 1094 m, 3.II.2005, Antilahimena et al.3299( MO, P, TAN); ibid. loc., 18°48’24”S 48°19’51”E, 1102 m, 10.XI.2005, Antilahimena & Edmond4182( G, MO, P, TAN); ibid. loc., 18°51’34”S 48°18’00”E, 1129 m, 8.VIII.2008, Antilahimena et al.6425( MO, P, TAN); Brickaville, Maroseranana, Ambodilendemy, Ankerana, 18°25’56”S 48°47’11”E, 854 m, 18.III.2011, Antilahimena & Randrimitantsoa7637( G, MO, P, TAN); Ambatovy, 18°50’30”S 48°18’11”E, 1111 m, 19.V.2010, Bernard et al.1573( MO, P, TAN); AnalamazaotraRS, 18°49’32”S 48°26’04”E, 941 m, 3.V.2010, Hong-Wa & Ortiz663( MO, P, TAN); Ambatovy, 18°51’17”S 48°19’02”E, 1117 m, 8.IX.2008, Miandrimanana et al.320( G, MO, P, TAN); ibid. loc., 18°51’48”S 48°18’26”E, 1022 m, 27.I.2008, Phillipson & Antilahimena6094( G, MO); Moramanga, Antanambao, Analamay, 18°49’26”S 48°20’16”E, 1000 m, 26.I.1997, Rakotomalaza et al.998( MO, P); ibid. loc., 18°51’22”S 48°19’34”E, 1080 m, 29.VI.1997, Rakotomalaza et al.1364( G, MO, TAN); Ambatovy, 18°51’08”S 48°17’48”E, 1127 m, 16.II.2005, Rakotovao et al.1191( G, MO, P, TAN); ZahamenaRNI, à 1 kmau S du village d’Antenina, 17°30’30”S 48°46’15”E, 900 m, 13.VIII.1994, Randrianjanaka & Zafy190A( G, K, MO, P, TAN); Manakambahiny-Est, ZahamenaRNI, 17°40’17”S 48°45’13”E, 1350 m, 16.IX.2002, S. Randrianasolo et al.304( G, MO, P, TEF); ibid. loc., 17°40’17”S 48°45’13”E, 1350 m, 16.IX.2002, S. Randrianasolo et al.307( G, MO, P, TEF); Brickaville, Maroseranana, Ambodilendemy, 18°25’57”S 48°46’52”E, 727 m, 14.III.2011, Ravelonarivo & Edmond3645( G, MO, P, TAN); ibid. loc., 18°25’24”S 48°47’06”E, 896 m, 22.III.2011, Ravelonarivo et al.3787( G, MO, P, TAN); Analamay, 18°50’49”S 48°18’30”E, 1124 m, 10.II.2005, Razafindraibe & Antilahimena76( MO, P, TAN).
4023241078
S
Madagascar
Typus
Moramanga
Alaotra-Mangoro
233
233
1
Toamasina
4023240895
[701,1400,652,679]
I
Madagascar
1080
-18.859446
Dynatec
21
48.286945
233
233
1
Toamasina
4023241050
G, K, P, TAN
Madagascar
Ranaivojaona
233
233
G00341615
1
Toamasina
4023240954
1994-03-21
1994-04-07
1994-03-21
K, MO, TAN
Ravelonarivo et al.
Madagascar
1293
-14.770833
Anjanaharibe-Sud
894
49.466667
Andapa
235
235
1
Prov. Antsiranana
4023240712
1994-11-03
1994-04-07
1994-03-21
G, K, MO, P, TAN
Ravelonarivo & Rabesonina
Madagascar
1260
-14.770833
894
49.466667
235
235
1
Antsiranana
4023240864
1997-11-10
1994-04-07
1994-03-21
G, MO
Rakotomalaza
Madagascar
Prov.
1200
-14.770833
Ivohibe
894
49.466667
Prov.
235
235
1
Prov. Fianarantsoa
4023241111
2008-02-12
1994-04-07
1994-03-21
G, MO, P, TAN
Bernard
Madagascar
Prov.
1077
-14.770833
Matsoandakana
894
49.466667
Befandriana-Nord
235
235
1
Prov. Mahajanga
4023241125
2008-02-25
1994-04-07
1994-03-21
G, MO, P, TAN
Bernard
Madagascar
Prov.
1293
-14.770833
Amparihy
894
49.466667
Andasinanantsomanga
235
235
1
Antsiranana
4023241524
2008-02-14
1994-04-07
1994-03-21
MO, P, TAN
Ravelonarivo et al.
Madagascar
Ambotaharanana
1208
-14.770833
Andranomena
894
49.466667
Andranomena
235
235
1
Antsiranana
4023241553
2008-02-15
MO, P, TAN
Ravelonarivo et al.
Madagascar
Tsaravilona
1208
-15.138611
Androva
21
49.363888
Amparihy
235
235
1
4023240496
2008-02-22
MO, P, TAN
Ravelonarivo
Madagascar
Tsaravilona
1166
-14.926667
Androva
21
49.42778
Amparihy
235
235
1
Toamasina
4023240840
2002-02-02
CNARP, K, MO, P, TEF
Zahamena PN & Andrianjafy
Madagascar
1456
-14.926667
Antanandava
21
49.42778
Prov.
235
235
1
Toamasina
4023241286
2003-10-08
MO, P
Antilahimena et al.
Madagascar
Ambinanitelo
1198
-14.926667
Mahalevona
21
49.42778
Maroantsetra
235
235
1
Toamasina
4023240391
2004-09-06
MO, P
Antilahimena
Madagascar
Ambinanitelo
1100
-14.926667
Maimborondro
21
49.42778
Ambalamahogo
235
235
1
Toamasina
4023241102
2005-02-03
MO, P, TAN
Antilahimena et al.
Madagascar
1094
-15.310834
Ambatovy
21
49.37861
Moramanga
235
235
1
4023240535
2005-11-10
G, MO, P, TAN
Antilahimena & Edmond
Madagascar
1102
-15.310834
21
49.37861
Moramanga
235
235
1
4023241556
2008-08-08
MO, P, TAN
Antilahimena et al.
Madagascar
Brickaville
1129
-18.806665
Ambodilendemy
21
48.330833
Maroseranana
235
235
1
4023240413
2011-03-18
G, MO, P, TAN
Antilahimena & Randrimitantsoa
Madagascar
Brickaville
854
-18.806665
Ambodilendemy
21
48.330833
Maroseranana
235
235
1
4023240520
2010-05-19
MO, P, TAN
Bernard et al.
Madagascar
Brickaville
1111
-18.806665
Ambatovy
21
48.330833
Maroseranana
235
235
1
4023241042
2010-05-03
MO, P, TAN
Hong-Wa & Ortiz
Madagascar
Brickaville
941
-18.806665
Analamazaotra
21
48.330833
Maroseranana
235
235
1
4023240337
2008-09-08
G, MO, P, TAN
Miandrimanana et al.
Madagascar
1117
-18.854723
Ambatovy
21
48.317223
235
235
1
4023241082
2008-01-27
G, MO
Phillipson & Antilahimena
Madagascar
Moramanga
1022
-18.854723
Ambatovy
21
48.317223
Antanambao
235
235
1
4023241280
1997-01-26
MO, P
Rakotomalaza et al.
Madagascar
Moramanga
1000
-18.854723
Analamay
21
48.317223
Antanambao
235
235
1
4023241339
1997-06-29
G, MO, TAN
Rakotomalaza et al.
Madagascar
Moramanga
1080
-18.854723
Ambatovy
21
48.317223
Antanambao
235
235
1
4023241328
1994-08-13
2005-02-16
1994-08-13
G, MO, P, TAN, K
Rakotovao et al.
Madagascar
1127
-18.856112
Ambatovy
21
48.32611
Manakambahiny-Est
235
235
1
4023241116
1994-08-13
2005-02-16
1994-08-13
G, MO, P, TAN, K
Randrianjanaka & Zafy
Madagascar
1127
-18.856112
Ambatovy
21
48.32611
Manakambahiny-Est
235
235
1
4023241405
2002-09-16
G, MO, P, TEF
S. Randrianasolo et al.
Madagascar
1350
-18.856112
Zahamena
21
48.32611
Manakambahiny-Est
235
235
1
4023240938
2002-09-16
G, MO, P, TEF
S. Randrianasolo et al.
Madagascar
Brickaville
1350
-18.856112
Ambatovy
21
48.32611
Manakambahiny-Est
235
235
1
4023241071
2011-03-14
G, MO, P, TAN
Ravelonarivo & Edmond
Madagascar
Brickaville
727
-18.856112
Ambodilendemy
21
48.32611
Maroseranana
235
235
1
4023240870
2011-03-22
G, MO, P, TAN
Ravelonarivo et al.
Madagascar
Brickaville
896
-18.856112
Ambatovy
21
48.32611
Manakambahiny-Est
235
235
1
4023241347
2005-02-10
MO, P, TAN
Razafindraibe & Antilahimena
Madagascar
Brickaville
1124
-18.856112
Analamay
21
48.32611
Manakambahiny-Est
235
235
1