A taxonomic revision of the genus Noronhia Stadtm. ex Thouars (Oleaceae) in Madagascar and the Comoro Islands Hong-Wa, Cynthia Boissiera 2016 2016-10-21 70 1 292 8WYRV Hong-Wa 2016 [503,748,380,408] Magnoliopsida Oleaceae Noronhia Plantae Lamiales 233 233 Tracheophyta species rollandii    Typus: MADAGASCAR. Prov. Toamasina: Alaotra-Mangoro, Moramanga, Ambatovy, Ampitabe, piste direction S du campement Dynatec, 18°51’34”S 48°17’13”E,  1080 m, 18. I .2005,  Ranaivojaonaet al. 1151(holo-: MO-6615552!; iso-: G[ G00341615]!, K!, P!, TAN!).   Diagnosis   Noronhia rollandiiHong-Wa can be distinguished from other members of the genus by its non-woody petioles, its opposite to verticillate, oblong leaf blades, its urceolate, yellow flowers, and its ovoid, punctate fruits.   Description  Treesto 15 mtall, trunk to 24 cmdiameter; young twigs cylindrical to somewhat flattened, 1.1-2.3 mmdiameter, glabrous; bark dark gray, smooth, with scattered lenticels. Leavesopposite, verticillate at the tips of the branches, persistent; bud scales deciduous; blades light green above, darker below, oblong, 4-10 3 1.5-3 cm, coriaceous, glabrous, domatia absent, base attenuate, margin flat, apex retuse, midrib slightly sunken above, distinctly raised below, secondary veins inconspicuous, 7-12 per side, 5-15 mmapart, looping 1-3.5 mmfrom the margin; petiole brown, 4-10 3 1-3 mm, not woody or only up to mid-length, glabrous. Thyrsesgeminate, multiflorous, compact; peduncle 5-10 mmlong, moderately pubescent; pedicel 2-8 mmlong, moderately pubescent; calyx moderately pubescent outside, glabrous inside, lobes triangular, 0.8-1.5 3 1-1.8 mm; corolla yellow tinged pinkish outside, yellow inside, urceolate, 3-4.5 mmlong, glabrous on both sides, the tube 1-2 mmlong, lobes widely ovate, apex acute; corona present, 1.7-2.3 mmlong, undivided; stamens 2-2.5 mmlong, anthers oblong to obovate, 1.5-1.8 mmlong; pistil 2-2.7 mmlong, stigma capitate. Fruitingpedicel 2-8 3 1-2.5 mm; young fruits green, brownish to blackish when mature, ovoid, 12-24 3 8-17 mm, covered with white dots, apex bluntly pointed to apiculate; dry pericarp 0.3-1.4 mmthick; endocarp woody; seed 9-15.5 3 7-11.5 mm.   Etymology This name honors the late Rolland Ranaivojaona, a botanist at the Parc Botanique et Zoologique de Tsimbazaza(TAN), for collecting the typespecimen and for sharing his knowledge of the Malagasyflora.   Distribution, ecology and phenology   Noronhia rollandiioccurs in mid- to high-elevation humid forests in the east, from Anjanaharibe-Sud to Mangoro ( Fig. 49). It produces flowers year-round except in December.   Conservation status The assessment included 29 collections representing 28 localities and yielded an EOO of 38,636 km 2, an AOO of 84 km 2, and 13 subpopulations representing 11 locations, of which seven occur within protected areas (Anjanaharibe-Sud, Corridor Ankeniheny-Zahamena, Makira, Mantadia, Masoala, Torotorofotsy, and Zahamena). Although the estimated AOO is rather small, the actual AOO is likely much larger and may well exceed 2,000 km 2. That, along with a large EOO, a high number of locations, and good representation within the network of protected areas, indicates a preliminary status of “Least Concern” for  N. rollandii.   Notes   Noronhia rollandiiis similar to  N. obtusifolia, but can be distinguished by its opposite to verticillate (vs. entirely opposite), oblong (vs. obovate) leaves, urceolate (vs. campanulate), yellow tinged pinkish (vs. white) flowers, the presence (vs. absence) of a corona, and its punctate (vs. smooth to ribbed) fruits. The new species can be recognized by its non-woody petioles, opposite to verticillate, oblong leaf blades, urceolate, yellow flowers, and ovoid, punctate fruits.    Paratypi   MADAGASCAR. Prov. Antsiranana: Andapa, Anjanaharibe-SudRS, 14°46’15”S 49°28’E,  1161-1424 m,  21.III-7.IV.1994,  Ravelonarivo et al.91( K, MO, TAN);  ibid. loc., 14°44’42”S 49°27’42”E,  1185-1335 m,  3.XI.1994,  Ravelonarivo & Rabesonina484( G, K, MO, P, TAN).  Prov. Fianarantsoa: IvohibeRS, 22°25’18”S 46°53’54”E,  1200 m,  10.XI.1997,  Rakotomalaza1526( G, MO).  Prov. Mahajanga: Befandriana-Nord, Matsoandakana, Ambotaharanana, Andranomena, 15°08’16”S 49°20’55”E,  1077 m,  12.II.2008,  Bernard770( G, MO, P, TAN);  ibid. loc., Andasinanantsomanga, Amparihy, 14°55’34”S 49°24’59”E,  25.II.2008,  Bernard938( G, MO, P, TAN);  ibid. loc., Ambotaharanana, Andranomena, forêt d’Anjiabe, 15°08’19”S 49°21’50”E,  1208 m,  14.II.2008,  Ravelonarivo et al.2831( MO, P, TAN);  ibid. loc.,  15.II.2008,  Ravelonarivo et al.2865( MO, P, TAN);  Tsaravilona, Amparihy, Androva, 14°55’36”S 49°25’40”E,  1154-1178 m,  22.II.2008,  Ravelonarivo2893( MO, P, TAN).  Prov. Toamasina: Antanandava, Zahamena PN, 17°29’47”S 48°45’35”E,  1000-1912 m,  2.II.2002,  Andrianjafyet al. 289( CNARP, K, MO, P, TEF);  Maroantsetra, Mahalevona, MasoalaPN, 15°25’47”S 49°57’33”E,  1198 m,  8.X.2003,  Antilahimena et al.2152( MO, P);  Ambinanitelo, Ambalamahogo, Maimborondro, 15°18’39”S 49°22’43”E,  1100 m,  6.IX.2004,  Antilahimena2835( MO, P);  Moramanga, Ambatovy, 18°49’19”S 48°20’08”E,  1094 m,  3.II.2005,  Antilahimena et al.3299( MO, P, TAN);  ibid. loc., 18°48’24”S 48°19’51”E,  1102 m,  10.XI.2005,  Antilahimena & Edmond4182( G, MO, P, TAN);  ibid. loc., 18°51’34”S 48°18’00”E,  1129 m,  8.VIII.2008,  Antilahimena et al.6425( MO, P, TAN);  Brickaville, Maroseranana, Ambodilendemy, Ankerana, 18°25’56”S 48°47’11”E,  854 m,  18.III.2011,  Antilahimena & Randrimitantsoa7637( G, MO, P, TAN);  Ambatovy, 18°50’30”S 48°18’11”E,  1111 m,  19.V.2010,  Bernard et al.1573( MO, P, TAN);  AnalamazaotraRS, 18°49’32”S 48°26’04”E,  941 m,  3.V.2010,  Hong-Wa & Ortiz663( MO, P, TAN);  Ambatovy, 18°51’17”S 48°19’02”E,  1117 m,  8.IX.2008,  Miandrimanana et al.320( G, MO, P, TAN);  ibid. loc., 18°51’48”S 48°18’26”E,  1022 m,  27.I.2008,  Phillipson & Antilahimena6094( G, MO);  Moramanga, Antanambao, Analamay, 18°49’26”S 48°20’16”E,  1000 m,  26.I.1997,  Rakotomalaza et al.998( MO, P);  ibid. loc., 18°51’22”S 48°19’34”E,  1080 m,  29.VI.1997,  Rakotomalaza et al.1364( G, MO, TAN);  Ambatovy, 18°51’08”S 48°17’48”E,  1127 m,  16.II.2005,  Rakotovao et al.1191( G, MO, P, TAN);  ZahamenaRNI, à  1 kmau S du village d’Antenina, 17°30’30”S 48°46’15”E,  900 m,  13.VIII.1994,  Randrianjanaka & Zafy190A( G, K, MO, P, TAN);  Manakambahiny-Est, ZahamenaRNI, 17°40’17”S 48°45’13”E,  1350 m,  16.IX.2002,  S. Randrianasolo et al.304( G, MO, P, TEF);  ibid. loc., 17°40’17”S 48°45’13”E,  1350 m,  16.IX.2002,  S. Randrianasolo et al.307( G, MO, P, TEF);  Brickaville, Maroseranana, Ambodilendemy, 18°25’57”S 48°46’52”E,  727 m,  14.III.2011,  Ravelonarivo & Edmond3645( G, MO, P, TAN);  ibid. loc., 18°25’24”S 48°47’06”E,  896 m,  22.III.2011,  Ravelonarivo et al.3787( G, MO, P, TAN);  Analamay, 18°50’49”S 48°18’30”E,  1124 m,  10.II.2005,  Razafindraibe & Antilahimena76( MO, P, TAN). 4023241078 S Madagascar Typus Moramanga Alaotra-Mangoro 233 233 1 Toamasina 4023240895 [701,1400,652,679] I Madagascar 1080 -18.859446 Dynatec 21 48.286945 233 233 1 Toamasina 4023241050 G, K, P, TAN Madagascar Ranaivojaona 233 233 G00341615 1 Toamasina 4023240954 1994-03-21 1994-04-07 1994-03-21 K, MO, TAN Ravelonarivo et al. Madagascar 1293 -14.770833 Anjanaharibe-Sud 894 49.466667 Andapa 235 235 1 Prov. Antsiranana 4023240712 1994-11-03 1994-04-07 1994-03-21 G, K, MO, P, TAN Ravelonarivo & Rabesonina Madagascar 1260 -14.770833 894 49.466667 235 235 1 Antsiranana 4023240864 1997-11-10 1994-04-07 1994-03-21 G, MO Rakotomalaza Madagascar Prov. 1200 -14.770833 Ivohibe 894 49.466667 Prov. 235 235 1 Prov. Fianarantsoa 4023241111 2008-02-12 1994-04-07 1994-03-21 G, MO, P, TAN Bernard Madagascar Prov. 1077 -14.770833 Matsoandakana 894 49.466667 Befandriana-Nord 235 235 1 Prov. Mahajanga 4023241125 2008-02-25 1994-04-07 1994-03-21 G, MO, P, TAN Bernard Madagascar Prov. 1293 -14.770833 Amparihy 894 49.466667 Andasinanantsomanga 235 235 1 Antsiranana 4023241524 2008-02-14 1994-04-07 1994-03-21 MO, P, TAN Ravelonarivo et al. Madagascar Ambotaharanana 1208 -14.770833 Andranomena 894 49.466667 Andranomena 235 235 1 Antsiranana 4023241553 2008-02-15 MO, P, TAN Ravelonarivo et al. Madagascar Tsaravilona 1208 -15.138611 Androva 21 49.363888 Amparihy 235 235 1 4023240496 2008-02-22 MO, P, TAN Ravelonarivo Madagascar Tsaravilona 1166 -14.926667 Androva 21 49.42778 Amparihy 235 235 1 Toamasina 4023240840 2002-02-02 CNARP, K, MO, P, TEF Zahamena PN & Andrianjafy Madagascar 1456 -14.926667 Antanandava 21 49.42778 Prov. 235 235 1 Toamasina 4023241286 2003-10-08 MO, P Antilahimena et al. Madagascar Ambinanitelo 1198 -14.926667 Mahalevona 21 49.42778 Maroantsetra 235 235 1 Toamasina 4023240391 2004-09-06 MO, P Antilahimena Madagascar Ambinanitelo 1100 -14.926667 Maimborondro 21 49.42778 Ambalamahogo 235 235 1 Toamasina 4023241102 2005-02-03 MO, P, TAN Antilahimena et al. Madagascar 1094 -15.310834 Ambatovy 21 49.37861 Moramanga 235 235 1 4023240535 2005-11-10 G, MO, P, TAN Antilahimena & Edmond Madagascar 1102 -15.310834 21 49.37861 Moramanga 235 235 1 4023241556 2008-08-08 MO, P, TAN Antilahimena et al. Madagascar Brickaville 1129 -18.806665 Ambodilendemy 21 48.330833 Maroseranana 235 235 1 4023240413 2011-03-18 G, MO, P, TAN Antilahimena & Randrimitantsoa Madagascar Brickaville 854 -18.806665 Ambodilendemy 21 48.330833 Maroseranana 235 235 1 4023240520 2010-05-19 MO, P, TAN Bernard et al. Madagascar Brickaville 1111 -18.806665 Ambatovy 21 48.330833 Maroseranana 235 235 1 4023241042 2010-05-03 MO, P, TAN Hong-Wa & Ortiz Madagascar Brickaville 941 -18.806665 Analamazaotra 21 48.330833 Maroseranana 235 235 1 4023240337 2008-09-08 G, MO, P, TAN Miandrimanana et al. Madagascar 1117 -18.854723 Ambatovy 21 48.317223 235 235 1 4023241082 2008-01-27 G, MO Phillipson & Antilahimena Madagascar Moramanga 1022 -18.854723 Ambatovy 21 48.317223 Antanambao 235 235 1 4023241280 1997-01-26 MO, P Rakotomalaza et al. Madagascar Moramanga 1000 -18.854723 Analamay 21 48.317223 Antanambao 235 235 1 4023241339 1997-06-29 G, MO, TAN Rakotomalaza et al. Madagascar Moramanga 1080 -18.854723 Ambatovy 21 48.317223 Antanambao 235 235 1 4023241328 1994-08-13 2005-02-16 1994-08-13 G, MO, P, TAN, K Rakotovao et al. Madagascar 1127 -18.856112 Ambatovy 21 48.32611 Manakambahiny-Est 235 235 1 4023241116 1994-08-13 2005-02-16 1994-08-13 G, MO, P, TAN, K Randrianjanaka & Zafy Madagascar 1127 -18.856112 Ambatovy 21 48.32611 Manakambahiny-Est 235 235 1 4023241405 2002-09-16 G, MO, P, TEF S. Randrianasolo et al. Madagascar 1350 -18.856112 Zahamena 21 48.32611 Manakambahiny-Est 235 235 1 4023240938 2002-09-16 G, MO, P, TEF S. Randrianasolo et al. Madagascar Brickaville 1350 -18.856112 Ambatovy 21 48.32611 Manakambahiny-Est 235 235 1 4023241071 2011-03-14 G, MO, P, TAN Ravelonarivo & Edmond Madagascar Brickaville 727 -18.856112 Ambodilendemy 21 48.32611 Maroseranana 235 235 1 4023240870 2011-03-22 G, MO, P, TAN Ravelonarivo et al. Madagascar Brickaville 896 -18.856112 Ambatovy 21 48.32611 Manakambahiny-Est 235 235 1 4023241347 2005-02-10 MO, P, TAN Razafindraibe & Antilahimena Madagascar Brickaville 1124 -18.856112 Analamay 21 48.32611 Manakambahiny-Est 235 235 1