Noronhia emarginata var. edentata H. Perrier in Mém. Inst. Sci. Madagascar , Sér. B, Biol. Vég. 2: 296. 1949
A taxonomic revision of the genus Noronhia Stadtm. ex Thouars (Oleaceae) in Madagascar and the Comoro Islands
Hong-Wa, Cynthia
Boissiera
2016
2016-10-21
70
1
292
8WYR4
(H. Perrier)
H. Perrier
[503,903,380,408]
Magnoliopsida
Oleaceae
Noronhia
Plantae
Lamiales
104
104
Tracheophyta
species
edentata
≡ Noronhia emarginatavar. edentataH. Perrierin Mém. Inst. Sci. Madagascar, Sér. B, Biol. Vég. 2: 296. 1949.
Lectotypus(designated here): MADAGASCAR. Prov. Toamasina: Forêt d’Analamazaotra, [ 18°55’48”S 48°25’48”E], 800 m, s.d., Perrier de la Bâthie8809( P[ P00791237]!; isolecto-: K[ K000233192, K000233193] image seen; P[ P03558849, P03558850]!). Syntypi: MADAGASCAR. Prov. Toamasina: Analamazaotra RS, 2.IV.1905, Thouvenot 58( K[ K000233191] image seen, P[ P00701239, P00701240]!); ibid. loc., comm. 3.XII.1934, Ursch 82( P[ P03558851]!).
Description Treesto 25 mtall, trunk to 35 cmdiameter; young twigs cylindrical, 1-6 mmdiameter, glabrous; bark medium to light gray, smooth to rugose. Leavesopposite, persistent; bud scales persistent; blades dark green above, medium green below, elliptic to obovate, 5-17 3 2-7.5 cm, coriaceous, glabrous, domatia absent, base acute to attenuate, margin slightly revolute, apex rounded to slightly acuminate, the acumen 0-5(-15) mm long, midrib sunken above, raised below, secondary veins conspicuous, 7-17 per side, 7-25 mmapart, looping 1.5-7 mmfrom the margin; petiole medium gray, 6-38 3 1.3-4 mm, entirely woody, glabrous. Thyrsesfasciculate, pauciflorous, compact; peduncle 8-9 mmlong, moderately pubescent; pedicel 1.5-8 mmlong, moderately pubescent; calyx moderately pubescent outside, glabrous inside, lobes triangular, 1-1.5 3 0.8-2 mm; corolla white sometimes tinged greenish, urceolate, 3-5.5 mmlong, glabrous on both sides, the tube 1.2-3 mmlong, lobes widely ovate, apex obtuse; corona present, 1.4-2.2 mmlong, undivided; stamens 1.8-2.5 mmlong, anthers widely oblong, 1.1-1.9 mmlong; pistil 1.4-2.5 mmlong, stigma capitate. Fruitingpedicel 2-9 3 1.8-3.6 mm; young fruits green, reddish black when mature, ovoid to subglobose, 12-31 3 8.5-26 mm, surface smooth, sometimes covered with a white pellicle, apex flat to bluntly pointed or apiculate; dry pericarp 1.3-4.2 mmthick; endocarp woody; seed 11.5-21.5 3 7-18 mm.
Distribution, ecology and phenology Noronhia edentataoccurs in low- to high-elevation humid forests in the east, from Sambava to Ambondrombe ( Fig. 19). It produces flowers and fruits from September to May.
Conservation status The assessment was based on 27 collections representing 19 localities, resulting in an EOO of 38,642 km 2, an AOO of 68 km 2, and 13 subpopulations representing 12 locations, of which six occur within protected areas (Analamazaotra, Betampona, Makira, Makirovana, Masoala, and Zahamena). Being widespread and present in many protected areas, Noronhia edentatais assigned a preliminary status of “Least Concern”.
Notes Noronhia edentatacan be recognized by its leaves, which have broadly elliptic blades and long, woody petioles, its white greenish flowers and its large fruits, with thick pericarp. There is a slight leaf heteroblasty in this species. In particular, the leaf base is narrower and the acumen longer (up to 15 mmlong) in seedlings and saplings than in adult plants. This taxon was previously recognized as a variety of N. emarginata, from which it differs by its habit, leaf shape, and flower color and shape. Moreover, these two entities belong to separate clades (HONG- WA & BESNARD, 2013, 2014). PERRIER DE LA BâTHIE (1949)described N. emarginatavar. edentatabased on the following syntypes: Perrier de la Bâthie8809, Thouvenot58and Ursch820; the latter is a typographical mistake for Ursch82. Theoverall quality and completeness of Perrier de la Bâthie8809justify the selection of one of the specimens at P as the lectotype.
Additional specimens examined MADAGASCAR. Prov. Antsiranana: Ambatobiribiry, [ 14°11’S 50°05’E], 50-345 m, 29-30.XI.1950, Humbert & Capuron24423( G, MO, P). Prov. Toamasina: Maroantsetra, Ambinanitelo, Ankirindro forest, 15°17’35”S 49°23’59”E, 664 m, 16.I.2003, Antilahimena et al.1741( G, MO, P); Manambolo, Amparihibe forest, 15°02’26”S 49°34’59”E, 800 m, 22.II.2003, Antilahimena1919( MO); Moramanga, Sahaevo, 18°50’38”S 48°16’54”E, 1006 m, 11.XI.2006, Antilahimena & Edmond4944( G, MO, P, TAN, TEF); Ambatovy, 18°51’43”S 48°17’39”E, 1049 m, 9.XII.2006, Antilahimena & Edmond5092( MO, P, TAN, TEF); ibid. loc., 18°50’28”S 48°18’16”E, 1085 m, 13.V.2010, Bernard1566( MO, P, TAN); AnalamazaotraRS, 18°56’12”S 48°25’09”E, 953 m, 2.V.2010, Hong-Wa & Ortiz650( MO, P, TAN); BetamponaRNI, 17°51’S 49°12’E, 275-650 m, 30.IX.1993, Lewis711( K, MO, P, TAN); AnalamazaotraRS, [ 18°56’S 48°26’E], 25.VII, Louvel88( P); Andasibe, 18°55’50”S 48°25’04”E, 963 m, 5.XII.2008, Rakotondrafara et al.932( MO, P, TAN); Moango, ZahamenaRNI, 17°33’32”S 48°53’55”E, 800 m, 4.V.2003, Rakotondrajaona et al.274( CNARP, G, MO, P, TEF); Lakato, Ambodigavo, forêt d’Analanjahana, 19°07’57”S 48°24’00”E, 794 m, 31.V.2007, Razanatsima et al.266( G, MO, P, TAN); 32 kmE of Moramanga, 18°57’S 48°27’E, 870 m, 29.I.1993, Schatz3452( MO, P, TAN); Maroantsetra, Farankaraina, [ 15°25’S 49°52’E], 20-25 m, 16.IX.1952, Service Forestier6151( P, TEF); Périnet, [ 18°56’S 48°26’E], 1000 m, 1.VI.1954, Service Forestier10374( P, TEF); Masoala Peninsula, Ambanizana, 15°40’24”S 49°57’51”E, 110-260 m, 28.X.1994, Vasey & Velo83( MO, P, TAN).
4023240881
P, K
Perrier de la Bathie
Madagascar
800
-18.93
Foret d'Analamazaotra
21
48.43
104
104
P00791237, K000233192, K000233193, P03558849, P03558850
1
Toamasina
lectotype
4023240409
1905-04-02
1934-12-03
1905-04-02
M, K, P
Analamazaotra RS
Madagascar
800
-18.93
Foret d'Analamazaotra
21
48.43
104
104
K000233191, P00701239, P00701240, P03558851
1
Toamasina
syntype
4023241036
1950-11-29
1950-11-30
1950-11-29
G, MO, P
Humbert & Capuron
Madagascar
Prov.
198
-14.183333
Ambatobiribiry
1289
50.083332
Maroantsetra
107
107
1
Prov. Antsiranana
4023241533
2003-01-16
1950-11-30
1950-11-29
G, MO, P
Antilahimena et al.
Madagascar
Prov.
664
-14.183333
Ambinanitelo
1289
50.083332
Maroantsetra
107
107
1
Prov. Toamasina
4023240953
2003-02-22
MO
Antilahimena
Madagascar
800
-15.040556
Amparihibe forest
21
49.583054
Manambolo
107
107
1
4023240540
2006-11-11
G, MO, P, TAN, TEF
Antilahimena & Edmond
Madagascar
1006
-18.84389
Sahaevo
21
48.281666
Moramanga
107
107
1
4023241552
2006-12-09
MO, P, TAN, TEF
Antilahimena & Edmond
Madagascar
1049
-18.84389
Ambatovy
21
48.281666
Moramanga
107
107
1
4023241131
2010-05-13
MO, P, TAN
Bernard
Madagascar
1085
-18.84389
Sahaevo
21
48.281666
Moramanga
107
107
1
4023240844
2010-05-02
MO, P, TAN
Hong-Wa & Ortiz
Madagascar
953
-18.84389
Analamazaotra
21
48.281666
Moramanga
107
107
1
4023240955
1993-09-30
K, MO, P, TAN
Lewis
Madagascar
463
-18.930555
Betampona
21
48.417778
107
107
1
4023240863
1993-09-30
P
Louvel
Madagascar
953
-18.930555
Analamazaotra
21
48.417778
107
107
1
4023241434
2008-12-05
MO, P, TAN
Rakotondrafara et al.
Madagascar
963
-18.930555
Andasibe
21
48.417778
Moango
107
107
1
4023240719
2003-05-04
CNARP, G, MO, P, TEF
Rakotondrajaona et al.
Madagascar
Lakato
800
-18.930555
Zahamena
21
48.417778
Moango
107
107
1
4023240899
2007-05-31
G, MO, P, TAN
Razanatsima et al.
Madagascar
Lakato
794
-18.930555
Ambodigavo
21
48.417778
Ambodigavo
107
107
1
4023241232
1993-01-29
MO, P, TAN
Schatz
Madagascar
870
-18.95
Moramanga
1273
48.45
Maroantsetra
107
107
1
4023240502
1952-09-16
P, TEF
Service Forestier
Madagascar
23
-18.95
Farankaraina
1273
48.45
Maroantsetra
107
107
1
4023241520
[454,1473,1500,1527]
1954-06-01
P, TEF
Service Forestier
Madagascar
1000
-18.95
Perinet
1273
48.45
Maroantsetra
107
107
1
4023240667
1994-10-28
MO, P, TAN
Vasey & Velo
Madagascar
185
-15.673334
Ambanizana
21
49.96417
Masoala Peninsula
107
107
1