Pinodytes Horn, 1880: 248 Catops cryptophagoides Mannerheim, 1852 Catopocerus P. capizzii ( Hatch, 1957: 21 ) P. chandleri Peck and Cook, 2011: 22 P. colorado Peck and Cook, 2011: 20 P. constrictus Peck and Cook, 2011: 23 P. contortus Peck and Cook, 2011: 23 P. cryptophagoides ( Mannerheim, 1852: 333 ) Horn 1880: 249 Pinodytes Hatch 1928: 72 1957: 20 Catopocerus Silfverberg 1987: 49 Horn (1880: 249) Leng 1920: 87 P. delnorte Peck and Cook, 2011: 44 P. eldorado Peck and Cook, 2011: 24 P. fresno Peck and Cook, 2011: 25 P. garibaldi Peck and Cook, 2011: 45 P. gibbosus Peck and Cook, 2011: 25 P. haidagwaii Peck and Cook, 2011: 45 P. humboldtensis Peck and Cook, 2011: 27 P. idaho Peck and Cook, 2011: 28 P. imbricatus ( Hatch, 1957: 20 ) P. isabella Peck and Cook, 2011: 46 P. klamathensis Peck and Cook, 2011: 28 P. losangeles Peck and Cook, 2011: 29 P. marinensis Peck and Cook, 2011: 30 P. minutus Peck and Cook, 2011: 31 P. monterey Peck and Cook, 2011: 31 P. newelli ( Hatch, 1957: 20 ) P. newtoni Peck and Cook, 2011: 17 P. orca Peck and Cook, 2011: 47 P. ovatus Hatch, 1957: 21 P. parvus Peck and Cook, 2011: 32 P. punctatus Peck and Cook, 2011: 19 P. pusio Horn, 1892: 45 Hatch 1928: 72 Catopocerus P. rothi Hatch, 1957: 21 P. sanjacinto Peck and Cook, 2011 P. sequoia Peck and Cook, 2011: 34 P. setosus Peck and Cook, 2011: 48 P. shasta Peck and Cook, 2011: 35 P. shoshone Peck and Cook, 2011: 48 P. sinuatus Peck and Cook, 2011: 49 P. spinus Peck and Cook, 2011: 36 P. subterraneus ( Hatch, 1935: 115 ) Hatch 1957: 19 Catopocerus P. tehama Peck and Cook, 2011: 36 P. tibialis ( Hatch, 1957: 20 ) P. tuolumne Peck and Cook, 2011: 37 An Annotated Catalog of the Leiodidae (Coleoptera) of the Nearctic Region (Continental North America North of Mexico) Peck, Stewart B. Newton, Alfred F. The Coleopterists Bulletin 2017 2017-06-27 71 2 211 258 8VRZ6 Horn, 1880 Horn 1880 [857,1115,1564,1585] Insecta Leiodidae Pinodytes Animalia Coleoptera 5 215 Arthropoda genus      Pinodytes Horn, 1880: 248. Typespecies:  Catops cryptophagoides Mannerheim, 1852(by monotypy). Portevin 1922: 2 (as synonym of  Catopocerus); Peck and Cook 2011: 14 (resurrected as valid genus).   Biology.All species are eyeless and flightless soil inhabitants (edaphobites), probably feed on subterranean fungi, and are sometimes collected in caves.   Distribution.W Nearctic (unglaciated lands west of the MississippiRiver) (42 species). =  Homeosoma Austin, 1880: 16(attributed to Horn; preoccupied, not Curtis, 1833). Typespecies:   Catops cryptophagoides Mannerheim, 1852(by monotypy). Hatch 1928: 72(synonymy).  Note.As  Homaeosomain Neave (1939–40) =  Typhloleiodes Hatch, 1935: 116. Typespecies:   Typhloleiodes subterraneus Hatch, 1935(by origi- nal designation). Hatch 1957: 19(synonymy).   P. angulatus Peck and Cook, 2011: 43. Holotypemale in CMNC. Type locality: Green Creek, Forest Road 1177, 14 miN and  22 miE Sweet Home, Willamette National Forest, Linn County, Oregon. North Americandistribution:  USA:OR.   P. borealis Peck and Cook, 2011: 18. Holotypemale in CMNC. Type locality: Chena Ridge,  5 miW Fairbanks, Alaska. North Americandistribution:  USA:AK; occurring north of the limits of Pleistoceneglaciations as a probable relict of a wider Tertiarydistribution.      P. capizzii(  Hatch, 1957: 21)( Catopocerus); Peck and Cook 2011: 41. Holotypemale in USNM. Type locality: Humbug Mt., Curry County, Oregon. North Americandistribution:  USA:ID, OR, WA. Biology: Fogeland Peck(1975).   P. chandleri Peck and Cook, 2011: 22. Holotypemale in FMNH. Type locality:  6 miW Log Springs, Mendocino National Forest, Tehama County, California. North Americandistribution:  USA:CA.   P. colorado Peck and Cook, 2011: 20. Holotypemale in CMNC. Type locality:  10 miN Wolcott, Eagle County, Colorado. North Americandistribution:  USA:CO.   P. constrictus Peck and Cook, 2011: 23. Holotypemale in FMNH. Type locality: 2.0 km off on Forest Road2N46, Angeles Crest Highway, Los Angeles County, California. North Americandistribution:  USA:CA.   P. contortus Peck and Cook, 2011: 23. Holotypemale in CDAE. Type locality:  5 miE Placervilleon highway 50, El Dorado County, California. North Americandistribution:  USA:CA.     P. cryptophagoides( Mannerheim, 1852: 333)( Catops);  Horn 1880: 249(  Pinodytes);  Hatch 1928: 72,  1957: 20(  Catopocerus); Peck and Cook 2011: 38 (distribution restricted). Lectotypein MCZC (designated by Peck and Cook 2011: 38); paralectotypein MZHF (in Mannerheim coll.) (  Silfverberg 1987: 49). Type locality: “Sitkha” {Sitka], Alaska. North American distribution:  Canada:BC.  USA:AK, WA (misidentified from Washington, DC by  Horn (1880: 249)and from Californiaby (  Leng 1920: 87), testePeck and Cook (2011)). Biology: Fogel and Peck (1975); Peck and Cook (2011).   P. delnorte Peck and Cook, 2011: 44. Holotypemale in CMNC. Type locality:  10 kmE Crescent City, Jed Smith Redwood State Park, Del Norte County, California. North Americandistribution:  USA:CA.   P. eldorado Peck and Cook, 2011: 24. Holotypemale in CDAE. Type locality: 1.6 mi W Quintette, El Dorado County, California. North Americandistribution:  USA:CA.   P. fresno Peck and Cook, 2011: 25. Holotypemale in FMNH. Type locality: Tamarack Ridge, 3.4 mi SE Highway168, Sierra National Forest, Fresno County, California. North Americandistribution:  USA:CA.   P. garibaldi Peck and Cook, 2011: 45. Holotypemale in OSUO. Type locality:  2 miN Garibaldi, Tillamook County, Oregon. North Americandistribution:  USA:OR (not WA).   P. gibbosus Peck and Cook, 2011: 25. Holotypemale in CMNC. Type locality:  20 kmSW Lake Elsinore, Route74, Orange County, California. North Americandistribution:  USA:CA.   P. haidagwaii Peck and Cook, 2011: 45. Holotypemale in CNCI. Type locality: Kiusta, Graham Island, Queen Charlotte Island, British Columbia. North Americandistribution:  Canada:BC.   P. humboldtensis Peck and Cook, 2011: 27. Holotypemale in CMNC. Type locality: Orick, Lady Byrd Johnson Grove, Redwood National Park, Humboldt County, California. North Americandistribution:  USA:CA.   P. idaho Peck and Cook, 2011: 28. Holotypemale in WFBM. Type locality:  10 miE Slate Creek, Slate Creek Campground, Idaho County, Idaho. North Americandistribution:  USA:ID.     P. imbricatus(  Hatch, 1957: 20)( Catopocerus); Peck and Cook 2011: 40. Holotypefemale in USNM. Type locality:  6 miN Buxton, Washington County, Oregon. North Americandistribution:  USA:OR.   P. isabella Peck and Cook, 2011: 46. Holotypemale in WFBM. Type locality: Isabella Creek, Clearwater County, Idaho. North Americandistribution:  USA:ID.   P. klamathensis Peck and Cook, 2011: 28. Holotypemale in CMNC. Type locality:  18 kmNE Crescent City, Myrtle Creek Botanical Reserve, Del Norte County, California. North Americandistribution:  USA:CA, OR.   P. losangeles Peck and Cook, 2011: 29. Holotypemale in USNM. Type locality: Noprecise location, Los Angeles County, California. North Americandistribution:  USA:CA.   P. marinensis Peck and Cook, 2011: 30. Holotypemale in FMNH. Type locality: Tocaloma, Marin County, California. North Americandistribution:  USA:CA.   P. minutus Peck and Cook, 2011: 31. Holotypemale in FMNH. Type locality: 4.9 mi W Pine Grove, Amador County, California. North Americandistribution:  USA:CA.   P. monterey Peck and Cook, 2011: 31. Holotypemale in CDAE. Type locality: 2.2 mi W Bottchers Gap, Monterey County, California. North Americandistribution:  USA:CA.     P. newelli(  Hatch, 1957: 20)( Catopocerus); Peck and Cook 2011: 41. Holotypemale in USNM. Type locality: Coos Head, Coos County, Oregon. North Americandistribution:  USA:OR, WA.   P. newtoni Peck and Cook, 2011: 17. Holotypemale in UADE. Type locality: Queen Wilhelmina State Park, Polk County, Arkansas. North Americandistribution:  USA:AR, MO, TX.   P. orca Peck and Cook, 2011: 47. Holotypemale in CMNC. Type locality: Oregon Cave, Oregon Caves National Monument, Josephine County, Oregon. North Americandistribution:  USA:OR.     P. ovatus  Hatch, 1957: 21( Catopocerus); Peck and Cook 2011: 21. Holotypemale in USNM. Type locality:  5 miN Gold Beach, Oregon. Peckand Cook2011: 21. North Americandistribution:  USA:OR. Biology: Fogeland Peck(1975).   P. parvus Peck and Cook, 2011: 32. Holotypemale in CDAE. Type locality:  5 miS Gasquet, Del Norte County, California. North Americandistribution:  USA:CA.   P. punctatus Peck and Cook, 2011: 19. Holotypemale in FMNH. Type locality: Mt. Spokane, Spokane County, Washington. North Americandistribution:  USA:ID, WA.     P. pusio Horn, 1892: 45;  Hatch 1928: 72(  Catopocerus); Peck and Cook 2011: 21. Lectotypemale in MCZC (designated by Peck and Cook 2011: 21). Type locality: Alameda County, California.North American distribution:  USA:CA.     P. rothi  Hatch, 1957: 21( Catopocerus); Peck and Cook 2011: 42. Holotypemale in USNM. Type locality:  6 miN Buxton, Washington County, Oregon. North Americandistribution:  USA:CA, OR. Biology: Fogeland Peck(1975).   P. sanjacinto Peck and Cook, 2011; 33. Holotypemale in CMNC. Type locality: Lake Fulmor,  15 kmNW Idyllwild, San Jacinto Mountains, Riverside County, California. North Americandistribution:  USA:CA.   P. sequoia Peck and Cook, 2011: 34. Holotypemale in CMNC. Type locality: Carmoe Crevice Cave, Tulare County, California. North Americandistribution:  USA:CA.   P. setosus Peck and Cook, 2011: 48. Holotypemale in FMNH. Type locality: 6.7 mi NE Brookings, Curry County, Oregon. North Americandistribution:  USA:CA, OR.   P. shasta Peck and Cook, 2011: 35. Holotypemale in FMNH. Type locality Buckhorn Pass, Highway299, Shasta-Trinity Countyline, California. North Americandistribution:  USA:CA.   P. shoshone Peck and Cook, 2011: 48. Holotypemale in WFBM. Type locality: Hobo Cedar Grove, Shoshone County, Idaho. North Americandistribution:  USA:ID.   P. sinuatus Peck and Cook, 2011: 49. Holotypemale in CMNC. Type locality: 7 miS, 13 miE, Ashland,  Soda MountainRoad, Jackson County, Oregon. North Americandistribution:  USA:OR.   P. spinus Peck and Cook, 2011: 36. Holotypemale in CDAE. Type locality:  5 miNE Butte Meadows, Butte County, California. North Americandistribution:  USA:CA.     P. subterraneus( Hatch, 1935: 115)( Typhloleiodes);  Hatch 1957: 19(  Catopocerus); Peck and Cook 2011: 39 (distribution restricted, not BC). Holotypemale in USNM. Type locality: Seaside, Clatsop County, Oregon. North American distribution:  USA:OR. Biology: Fogel and Peck (1975).   P. tehama Peck and Cook, 2011: 36. Holotypemale in DENH. Type locality:  5 miSE Manton, Bluff Springs, Tehama County; California. North Americandistribution:  USA:CA.     P. tibialis(  Hatch, 1957: 20)( Catopocerus); Peck and Cook 2011: 43. Holotypemale in USNM. Type locality: Gold Beach, Curry County, Oregon. North Americandistribution:  USA:OR.   P. tuolumne Peck and Cook, 2011: 37. Holotypemale in CMNC. Type locality: Harden Flat, Tuolumne County, California. North Americandistribution:  USA:CA. [287,538,556,577] CMNC United States of America Linn County Green Creek North American 6 216 1 Oregon holotype [273,534,690,711] CMNC United States of America Tertiary Chena Ridge Pleistocene 6 216 1 Alaska holotype [267,432,851,872] USNM United States of America Curry County Humbug Mt. Peck 6 216 1 Oregon holotype [284,538,986,1007] FMNH United States of America Tehama County Mendocino National Forest North American 6 216 1 California holotype [278,536,1118,1139] CMNC United States of America Eagle County North American 6 216 1 Colorado holotype [293,540,1225,1246] FMNH United States of America Los Angeles County Forest Road North American 6 216 1 California holotype [283,537,1359,1380] CDAE United States of America El Dorado County North American 6 216 1 California holotype [809,1070,273,294] CMNC United States of America Del Norte County Jed Smith Redwood State Park North American 6 216 1 California holotype [812,1070,406,427] CDAE W Quintette United States of America El Dorado County North American 6 216 1 California holotype [780,1023,512,533] FMNH Tamarack Ridge & SE Highway United States of America Fresno County Sierra National Forest North American 6 216 1 California holotype [814,1071,645,666] OSUO United States of America Tillamook County North American 6 216 1 Oregon holotype [812,1070,752,773] CMNC United States of America Orange County Route North American 6 216 1 California holotype [845,1119,858,879] CNCI Canada North American Kiusta Queen Charlotte Island 6 216 1 British Columbia holotype [868,1123,991,1012] CMNC United States of America Humboldt County Orick North American 6 216 1 California holotype [775,1021,1123,1144] WFBM United States of America North American Slate Creek Campground County 6 216 1 Idaho holotype [823,979,1230,1251] USNM United States of America County North American 6 216 1 Washington holotype [791,1026,1336,1357] WFBM United States of America Clearwater County Isabella Creek North American 6 216 1 Idaho holotype [853,1105,1442,1463] CMNC United States of America Del Norte County Myrtle Creek Botanical Reserve North American 6 216 1 California holotype [824,1073,1575,1596] USNM United States of America Los Angeles County No North American 6 216 1 California holotype [828,1074,1681,1702] FMNH United States of America Marin County Tocaloma North American 6 216 1 California holotype [296,556,247,268] FMNH W Pine Grove United States of America Amador County North American 7 217 1 California holotype [303,558,353,374] CDAE W Bottchers Gap United States of America Monterey County North American 7 217 1 California holotype [287,453,459,480] USNM United States of America Coos County Coos Head North American 7 217 1 Oregon holotype [294,556,565,586] UADE United States of America Polk County Queen Wilhelmina State Park North American 7 217 1 Arkansas holotype [248,487,672,693] CMNC United States of America Josephine County Cave North American 7 217 1 Oregon holotype [268,419,778,799] USNM United States of America Peck Peck Fogel 7 217 1 Oregon holotype [270,511,911,932] CDAE United States of America Del Norte County North American 7 217 1 California holotype [307,558,1017,1038] FMNH United States of America Spokane County Mt. Spokane North American 7 217 1 Washington holotype [256,412,1256,1277] USNM United States of America Peck County Fogel 7 217 1 Washington holotype [311,522,1389,1410] CMNC United States of America Riverside County Lake Fulmor North American 7 217 1 California holotype [278,513,1522,1543] CMNC United States of America Tulare County Carmoe Crevice Cave North American 7 217 1 California holotype [273,512,1628,1649] FMNH NE Brookings United States of America Curry County North American 7 217 1 Oregon holotype [801,1044,194,215] FMNH United States of America Shasta-Trinity County Buckhorn Pass North American 7 217 1 California holotype [838,1093,300,321] WFBM United States of America Shoshone County Hobo Cedar Grove North American 7 217 1 Idaho holotype [831,1091,406,427] CMNC United States of America Jackson County Ashland North American 7 217 1 Oregon holotype [802,1044,512,533] CDAE United States of America Butte County North American 7 217 1 California holotype [810,1047,805,826] DENH United States of America Tehama County Bluff Springs North American 7 217 1 California holotype [818,986,911,932] USNM United States of America Curry County Gold Beach North American 7 217 1 Oregon holotype [839,1093,1017,1038] CMNC United States of America Tuolumne County Harden Flat North American 7 217 1 California holotype