Matsumura, 1912 : 123 Arotes moiwanus Uchida, 1928 : 34 moiwanus Arotes albicinctus Uchida, 1953 : 128 Iwata, 1960 : 150 Arotes albicinctus Iwata, 1958 : 68 Konishi & Yamamoto, 2000 : 749 Castillo et al ., 2011 : 85 Revision of the genus Arotes Gravenhorst (Hymenoptera: Ichneumonidae: Acaenitinae) from Japan Ito, Masato Watanabe, Kyohei Maeto, Kaoru Zootaxa 2014 3893 2 196 208 Matsumura, 1912 Matsumura 1912 [151,608,1045,1072] Insecta Ichneumonidae Arotes Animalia Hymenoptera 8 204 Arthropoda species moiwanus     Phaenolobus( Acoenitus) moiwanus  Matsumura, 1912: 123. [ Typelocality: Japan, Hokkaido, Sapporo]    Arotes moiwanus var. alboannulatus  Uchida, 1928: 34. [ Typelocality: Japan, Hokkaido] Synonymized with the variety  moiwanusby Townes et al. (1965)    Arotes albicinctus var. moiwanus:  Uchida, 1953: 128;  Iwata, 1960: 150.    Arotes albicinctusf.  moiwanus:  Iwata, 1958: 68.    Arotes albicinctus moiwanus:  Konishi & Yamamoto, 2000: 749.    Arotes moiwanus:  Castillo et al., 2011: 85-86.   Female(n=18). Body length: 13.5–18.0 mm. Head. Clypeus 0.6–0.7 times as long as wide. Face 0.6–0.7 times as long as wide, strongly punctate and weakly convex. MSL 0.9 times as long as BWM. Lower tooth of mandible longer than upper tooth. OOL/OD=1.6–1.8. POL/OD= 1.2–1.4. Antenna with 35–38 flagellomeres; first flagellomere 1.2–1.4 times as long as second flagellomere. Mesosoma. Lateral area of pronotum densely punctate and longitudinally striate. Lateral part of collar densely punctate. Subalar prominence sparsely punctate. Metapleuron extensively punctate ( Fig. 5). Fore wing: length 11.5–15.5 mm, with vein rs-mcross-vein distad of vein 2 m-cu( Fig. 11). Hindfemur 3.4–3.7 times as long as maximum depth in lateral view, with distinct convexity ventrally ( Fig. 8). Hindtibia 10.8–12.8 times as long as maximum depth in lateral view. Hindfirst tarsomere 2.4–2.7 times as long as second hind tarsomere and 3.2–3.8 times as long as longer hind tibial spur. Metasoma. T1 2.6–3.2 times as long as maximum width, 2.2–2.3 times as long as T2. T2 0.7–0.8 times as long as maximum width. T1and T2 sparsely punctate. Other tergites densely punctate. Ovipositor sheath 2.4–2.5 times as long as hind tibia. Coloration. Body black. Antennal flagellum usually without a white band (color form A), but occasionally with a white band in Hokkaido (color form B) ( Figs 19, 20). Inner margin of eye, tegula, hind corner of pronotum, mesopleuron in part, scutellum, postscutellum, propodeum, apices of T1 and T2 light yellow. Front and middle leg, trochanter, trochantellus, apex of hind tarsus tinged with yellowish brown. Fore wing lacking dark marks ( Fig. 11).  Male(n=6; genitalia, n=3). Similar to female. Frons sometimes with strong oblique striations extending from medial ocellus toward eye and densely punctate near antenna ( Fig. 21). MSL 0.8 times as long as BWM. OOL/ OD= 1.4–1.7 OD. POL/OD= 1.1–1.3. Antenna with 39–42 flagellomeres; first flagellomere 1.1–1.2 times as long as second flagellomere. Hindfemur 3.2–3.6 times as long as maximum depth in lateral view. Hindfirst tarsomere 2.1–2.6 times as long as second hind tarsomere and 2.8–3.6 times as long as longer hind tibial spur. Face and clypeus light yellow. Subgenital plate pentagonal with basal angle obtuse ( Fig. 16), its length ca. 0.7 times as long as aedeagus. Paramere short, basal part strongly projected toward the base of subgenital plate ( Figs 13, 14). Aedeagus gently curved, its penis valve ca. 2.0 times as long as basal apodeme ( Fig. 15).   FIGURE 19–21.  A. moiwanus. 19, antennal flagellum, dorsal view, female (color form B, Hokkaido); 20, antennal flagellum, dorsal view, female (color form A, Fukui Pref.); 21, head, dorsal view, male (Ehime Pref.).  Specimens examined. Color form A: Japan: 1F ( holotypeof  Phaenolobus moiwanus), Mt. Moiwasan, Sapporo-shi, Hokkaido, 18. June 1904, Ishida (SEHU); 1M, Jozan-kei, Sapporo-shi, Hokkaido, 1. July 1912, S. Mastumura (SEHU); 1F, Maruyama, Sapporo-shi, Hokkaido, 29. July 2009, K. Watanabe (KPMNH) [AmHk]; 1F, Hinoemata-mura, Fukushima Pref. 19. July-1. August 2004, H. Makihara (MsT) (KPMNH); 1F, Mt. Hotaka-san, Katashina-mura, Gumma Pref., 1. August, T. Ishizaki (KPMNH); 1F, Mt. Hotaka-san, Katashina-mura, Gunma Pref., 1. August 2007; 1F, same locality, 2. August 2007, K. Watanabe (KPMNH) [AmHn1]; 1F, Kinunuma, Kuriyama-mura, Tochigi Pref., 19. July-1. August 2004, H. Makihara (Malaise Trap) (KPMNH); 1F, Mt. Hakkaisan, Outaki-mura, Nagano Pref., 8. August 2010, K. Watanabe (KPMNH); 1F, Mt. Hakkaisan, Outaki-mura, Nagano Pref., 2. August 2012, M. Ito (NSMT); 1F, Masutomi, Hokuto-shi, Yamanashi Pref., 7. August 2007, K. Watanabe (KPMNH); 1M, Hikawa-rindo, Yamanashi Pref., 16. June 2007, H. Katahira (KPMNH); 1F, Daibosatsutoge, Yamanashi Pref., 3. August 2010, T. Maeda (KPMNH); 1F, Gozaishi Spa., Nirasaki-shi, Yamanashi Pref., 8–17. July 2005, K. Hosoda (Malaise Trap) (OMNH); 1M, Kaminegori, Obama City, Fukui Pref., 8. June 2013, M. Adachi (NSMT) [AmHn2]; 1F, Noka-dani, Ooi Town, Fukui Pref., 9. June 2013, S. Fujie (SEHU) [AmHn3]; 1F, Biwako Balley, Otsu-shi, Shiga Pref., 29. June 2011, M. Ito (SEHU); 1M, Seizui, Togouchi-cho, Hiroshima Pref., 16. May 1997, A. Shimizu (KPMNH); 1M, Tsuchigoya, Omogo-mura, Ehime Pref., 12. July 1998, R. Matsumoto (OMNH); 2M, Mt. Ibukiyama, Saijo-shi, Ehime Pref., 23. June 2004, R. Matsumoto (OMNH).  Color form B: Japan: 1F ( lectotypeof  Arotes moiwanus var. alboannulatus), T. Uchida (SEHU); 1F, Mt. Daisetsu-zan, Hokkaido, 4–10. August 1926, S. Matsumura (SEHU); 1F, Mt. Teine-kanayama, Sapporo-shi, Hokkaido, 18. July 2012, M. Ito (OMNH) [Ama1]; 1F, Kannonzawa, Sapporo-shi, Hokkaido, 20. August 2007, T. Yoshida (KPMNH) [Ama2]; 1F, Yuni-cho, Sorachi, Hokkaido, 2–12. July 2007, N. Kuhara (MsT) (NIAES).   Distribution. Japan(Hokkaido, Honshu, Shikoku and Kyushu) and Taiwan.   Remarks.We found no other morphological differences between the color form A (including the typeof  A. moiwanus) and the color form B (including the typeof A. m. var. alboannulatus). As shown by Castillo et al. (2011), this species can be distinguished from  A. albicinctusby the following character states of females: frons with weak striations or striations absent between medial ocellus and eye (densely and coarsely punctate in  A. albicinctus); orbit with moderately fine, sparse punctures near antenna (strong oblique striations in  A. albicinctus); yellow marking on scutellum (black in  A. albicinctus). But males of both species cannot be distinguished from each other except for the following body coloration: scutellum, propodeum and mesopleuron each with a light yellow (black in  A. albicinctus). It can also be distinguished from two Japanese congeners,  A. japonicusand  A. sugiharai, by the fore wing lacking dark markings and the long ovipositor sheath, 2.4–2.5 times as long as the hind tibia (1.9–2.0 times in other species).