Lema semipunctata Lilioceris semipunctata Lema dehaanii Crioceris rufimembris A review of the semipunctata species group within the genus Lilioceris Reitter, 1913 (Coleoptera, Chrysomelidae) Xu, Yuan Qiao, Gexia Liang, Hongbin ZooKeys 2024 2024-03-20 1195 337 381 96E405E9-2055-5B33-9D8C-73AD64410A82 (Fabricius, 1801) Fabricius 1801 Insecta Chrysomelidae Lilioceris CoL Animalia Lilioceris semipunctata Coleoptera 0 337 Arthropoda species semipunctata   Figs 21 , 22 , 35A-D , 45A-C , 50A-D , 62A-C , 67 , 73A, B   Lema semipunctataFabricius, 1801: 472 (Sumatra).  Lilioceris semipunctata: Kimoto and Gressitt 1961: 58.  Lema dehaaniiGuerin-Meneville, 1844: 261 (Java). Synonymized by Kimoto and Gressitt 1979: 231.  Crioceris rufimembrisPic, 1921: 2 (China: Yunnan, syntype, male), syn. nov.  Type material examined.  Lectotype(here selected) of  Lilioceris semipunctata(ZMUC, photo), L. semipunctata, Sumatra, Daldorff / TYPE / zmuc 00031030 [other two syntypes: zmuc 00031032 and zmuc 00031031 are identical to  Lilioceris latissima]. Syntypeof  Lilioceris rufimembris(MNHN, photo): Yunnan / Lilioceris rufimembris, Pic / type / Museum Paris, Coll. M. Pic / Type / Syntype / Syntype, Lilioceris rufimembris(Pic, 1921) / MNHN, Paris, EC15769.  Other material examined. Total  451 specimens.  China: Yunnan:  2♀, Xiaomengyang,  1000 m/ 1957.V.6, Dahua Liucoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.VII.9, Yiran Zhangcoll.;  1♀, YiwutoMenglun,  650 m/ 1964.IV.14, Baolin Zhangcoll.;  1♀, Xishuangbanna, Menghai  1200-1600 m/ 1958.VII.25, Shuyong Wangcoll.;  1♀, Xishuangbanna, Jinghong, 900-  710 m/ 1958.IV.29, Yiran Zhangcoll.;  1♀, Xishuangbanna, Damenglong,  650 m/ 1958.VII.11;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.19, Fuji Pucoll.;  2♀, Xishuangbanna,  750 m/ 1958.V.31, Chunpei Hongcoll., Lilioceris semipunctata, det. Peiyu Yu;  2♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.IV.30, Shuyong Wangcoll.;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.IV.30, Fuji Pucoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1958.VIII.31;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.8, Fuji Pucoll.;  1♀, Xishuangbanna, Damenglong,  650 m;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.10, Shuyong Wangcoll.;  1♀, Xishuangbanna, Damenglong,  650 m;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.V.6, Fuji Pucoll.;  1♀, Xishuangbanna, Mengzhe,  1890 m/ 1958.VII.7, Fuji Pucoll.;  1♀, Xiaomengyang,  850 m/ 1957.VI.21, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.VII.9, Yiran Zhangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1958.VII.20, Yiran Zhangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.VII.10, Yiran Zhangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.VI.6, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.19, Facai Zhangcoll.;  1♀, Xishuangbanna, Mengzhe,  620-650 m/ 1959.V.8, Fuji Pucoll.;  1♀, Xishuangbanna, Mengzhe,  850 m/ 1958.VII.3, Fuji Pucoll.;  1♀, Xishuangbanna, Mengzhe,  850 m/ 1958.VII.4, Shuyong Wangcoll.;  2♀, Xishuangbanna, Mengzhe,  1200 m/ 1958.VI.14, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengzhe,  875 m/ 1958.VII.4, Zhizi Chencoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.16, Facai Zhangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.VI.9, Shuyong Wangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.VI.25, Shuyong Wangcoll.;  2♀, Xishuangbanna, Mengzhe,  870 m/ 1958.IX.3, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengzhe,  1200 m/ 1958.VI.14, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengzhe,  875 m/ 1958.VII.1, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.12, Facai Zhangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.22, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.VI.29, Facai Zhangcoll.;  1♀, Xishuangbanna, Mengsong,  1600 m/ 1958.VII.25, Leyi Zhengcoll.;  1♀, Xishuangbanna, Mengsong,  1600 m/ 1958.IX.24, Leyi Zhengcoll.;  1♀, Xishuangbanna, Mengzhe,  870 m/ 1958.VII.11, Fuji Pucoll.;  1♀, Menghai,  1060 m, 1980.5.1, Zuyun Guocoll.;  1♀, Xishuangbanna, Menghun,  750 m, 1958.5.30;  1♀, Xishuangbanna, Menghun,  750-950 m, 1958.VI.3;  1♀, Xishuangbanna, Menga,  800 m/ 1958.VI.1, Shuyong Wangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.IX.3, Shuyong Wangcoll.;  1♀, Xishuangbanna, Damenglong,  650 m, 1958.IV.21, Xuwu Meng;  1♀, Ruili, Nongdao, Wudian, 23.98207°N, 97.61129°E/ 2012.XI.  4, 990 m, Xinlei Huangleg. Inst.of Zoology, CAS.;  1♀, Nabanhe Reserve, 2008.VIII. 6, 727 m/ 22.16713°N, 100.65887°E, Jingxin Liucoll.;  1♀, Fugong, Shangpa Town, 26.86203°N, 98.87142°E/  1177 m, 2005.8.22, Liang HBand Zhang JFcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.12, Facai Zhangcoll.;  1♀, Xishuangbanna, Mengla,  1050-1080 m/ 1958.VIII.15, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  1050-1080 m/ 1958.VIII.20, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  1050-1080 m/ 1958.VIII.16, Shuyong Wangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.VI.18, Shuyong Wangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.VII.17, Shuyong Wangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.VII.12, Shuyong Wangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.VII.9, Shuyong Wangcoll.;  1♀, Xishuangbanna, Xiaomengyang,  850 m/ 1957.VI.27, Lingchao Zangcoll.;  1♀, Xishuangbanna, Menglun,  650 m/ 1964.IV.14, Baolin Zhangcoll.;  1♀, Xishuangbanna, Ganlanba,  540 m, 1957.IV.17, Fuji Pucoll.;  1♀, Baoshanto Yongping, 1955.V.28, Tianrong Huangcoll.;  1♀, Wenshan/ 1958.VIII.2;  1♀, Xishuangbanna, Menghun,  1200-1400 m, 1958.X.28, Shuyong Wangcoll.;  2♀, Xishuangbanna, Menghun,  750 m, 1958.VI.1, Shuyong Wangcoll.;  3♀, Xishuangbanna, Menghun,  750 m/ 1958.VI.5, Chunpei Hongcoll.;  1♀, Xishuangbanna, Menga,  800 m/ 1958.IX.3, Shuyong Wangcoll.;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.16, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1958.VII.9, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1958.V.30, Facai Zhangcoll.;  1♀, Xishuangbanna, Damenglong,  650 m/ 1958.IV.7;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.31, Suofu Licoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.8, Fuji Pucoll./ host plant: Smilaxsp.;  1♀, Ruili, 1979.IX.2, Mengqiao, Leyi Zhengcoll./ 1400 m/ Lilioceris semipunctata, det. Liang H.B.;  1♀, Tengchong, Jietou, Wujia Jidi, 1929.IV.2,  1530 m, Xiaohong Oucoll.;  1♀, Xishuangbanna, Damenglong,  650 m/ 1958.VI.9, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengsong,  1600 m/ 1958.IV.23, Yiran Zhangcoll.;  1♀, Xishuangbanna, Mengsong,  1600 m/ 1958.IV.28, Fuji Pucoll.;  1♀, Xishuangbanna, Kongming Shan,  2600 m/ 1957.IX.23, Lingchao Zangcoll.;  2♀, Lancang,  1000 m/ 1957.VII.26, Shuyong Wangcoll.;  1♀, Lancang,  1100 m/ 1957.VIII.8, Lingchao Zangcoll.;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.VI.1, Fuji Pucoll.;  1♀, Lancang,  1100 m/ 1957.VIII.2, Lingchao Zangcoll.;  1♀, Shuangjiang, 1953.VI;  1♀, 22 kmnortheast of Jingdong, 1957.V.9, Menchalsky[in Chinese]coll.;  1♀, Xishuangbanna, Menghun,  650 m/ 1958.VI.15, Xuwu Mengcoll.;  1♀, Xishuangbanna, Menga,  1000 m/ 1958.V.16, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Fuji Pucoll.;  1♀, Tengchong, Qingshui, Rehai, on vegetation, 24.94861°N, 98.45181°E/  1470 m, 2006.6.1, Liang HBand Hu Pcoll.,  California Academyand IOZ, Chinese Acad. Sci.;  1♂, Xishuangbanna, Mengzhe,  870 m/ 1958.IX.2, Shuyong Wangcoll.;  1♂, Xishuangbanna, Mengzhe,  870 m/ 1958.IX.2, Fuji Pucoll.;  1♂, Ruili, Ruili Botanicalgarden, 24.07230°N, 97.81944°E/ 2012.X.25,  1152 m, Xinlei Huangcoll.;  1♂, Ruili, Nongdao, Wudian, 23.98207°N, 97.61129°E/ 2012.XI,  990 m, Xinlei Huangcoll./ Lilioceris latissima, det. Liang H.B. 2022;  1♂, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.16, Shuyong Wangcoll.;  1♂, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.10, Shuyong Wangcoll.;  1♂, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.16, Fuji Pucoll.;  1♂, Xishuangbanna, Menghun,  1200 m/ 1958.VI.28, Shuyong Wangcoll.;  1♂, Xishuangbanna, Mengzhe,  870 m/ 1958.IX.2, Fuji Pucoll.;  1♂, Xishuangbanna, Mengzhe,  870 m/ 1958.IX.9, Fuji Pucoll.;  1♂, Xishuangbanna, Mengzhe,  870 m/ 1958.VII.4, Shuyong Wangcoll.;  1♂, Xishuangbanna, Mengzhe,  1200 m/ 1958.IX.14, Shuyong Wangcoll.;  1♂, Xiaomengyang,  850 m;  1♂, Xishuangbanna, Menghun,  1200 m/ 1958.V.23, Xuwu Mengcoll.;  3♂, Xishuangbanna, Mengzhe,  1200 m/ 1958.VI.14, Shuyong Wangcoll.;  2♂, Xishuangbanna, Mengla,  620-650 m/ 1959.VII.9, Fuji Pucoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Suofu Licoll.;  1♂, Xishuangbanna, Mengsong,  1600 m/ 1958.IV.24, Fuji Pucoll.;  1♂, Xishuangbanna, Menga,  800 m/ 1958.V.30, Fuji Pucoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.11, Fuji Pucoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.16, Fuji Pucoll.;  1♂, Simaoto Puwen,  950-1200 m/ 1957.V.11, Guangji Hongand Zhiran Mengcoll.;  1♂, Xishuangbanna, Mengzhe,  620-650 m/ 1959.VII.9, Facai Zhangcoll.;  1♂, Shuangjiang, 1953.VI;  1♂, 22 kmnortheast of Jingdong, 1957.V.9, Mengqiacijicoll.;  1♂, Xishuangbanna, Menghun,  1200 m/ 1958.VI.28, Shuyong Wangcoll.;  1♂, Xishuangbanna, Menghun,  750 m/ 1958.VI.2;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.11, Facai Zhangcoll.;  1♂, Xiaomengyang,  850 m/ 1957.IX.13, Shuyong Wangcoll.;  1♂, Xishuangbanna, Damenglong,  650 m/ 1958.IV.7;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.16, Fuji Pucoll.;  1♂, Xiaomengyang,  850 m/ 1957.VI.18, Shuyong Wangcoll.;  1♂, Xiaomengyang,  620-650 m/ 1959.V.5, Yiran Zhangcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.7, Facai Zhangcoll.;  2♂, Xiaomengyang,  850 m/ 1957.VII.11-12, Lingchao Zangcoll.;  1♂, Xishuangbanna, Damenglong,  650 m/ 1958.IV.6, Yiran Zhangcoll.;  2♂, Xiaomengyang,  850 m/ 1957.V.7, Bonfirov[in Chinese] coll.;  1♂, Cheli[= Jinghong], Shihuiyao,  750 m, 1957.IV.27, Bonfirov[in Chinese] coll.;  1♂, Gaoligong Shan,  1200 m/ 1958.VIII.22, Fulong Licoll.;  1♂, Xishuangbanna, Menglun,  650 m/ 1964.V.10, Baolin Zhangcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.VI.6, Fuji Pucoll.;  1♂, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.15, Fuji Pucoll.;  1♂, Puwen, 1957.III. 27, Mengqiacijicoll.;  1♂, Xishuangbanna, Menga,  1050-1080 m/ 1958.V.23, Shuyong Wangcoll.;  1♂, Xishuangbanna, Menglun,  650 m/ 1959.VII.23, Fuji Pucoll.;  2♂, Menga, 1958.V.30/  800 m;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Fuji Pucoll.;  1♂, Xishuangbanna, Damenglong,  650 m/ 1958.IV.18, Chunpei Hongcoll.;  1♂, Xishuangbanna, Mengla,  1050-1080 m/ 1958.V.20, Fuji Pucoll.;  1♂, Xishuangbanna, Mengla,  1050-1080 m/ 1958.VIII.18, Fuji Pucoll.;  1♂, Xishuangbanna, Menglun,  650 m/ 1959.VIII.30, Yiran Zhangcoll.;  1♂, Xiaomengyang,  850 m/ 1957.IX.12, Shuyong Wangcoll.;  1♂, Mengla/ 1982.IV.20, Shengqiao Jiangcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.8, Fuji Pucoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.VII.9, Yiran Zhangcoll.;  2♂, Xishuangbanna, Mengla,  620-650 m/ 1959.VI.6-9, Fuji Pucoll.;  1♂, Xishuangbanna, Menghun,  1200-1400 m/ 1958.VI.3, Shuyong Wangcoll.;  1♂, Xishuangbanna, Menghun,  1300 m/ 1958.VI.7, Leyi Zhengcoll.;  1♂, Xishuangbanna, Menghun,  750 m/ 1958.VI.7;  1♂, Xishuangbanna, Menghun,  1200 m/ 1958.V.23, Xuwu Mengcoll.;  1♂, Xishuangbanna, Menghun,  750 m/ 1958.VI.2, Xuwu Mengcoll.;  1♂, Xishuangbanna, Menghun,  1200 m/ 1958.V.22, Xuwu Mengcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Facai Zhangcoll.;  1♂, Xishuangbanna, Menghun,  1200 m/ 1958.VI.3;  1♂, Xishuangbanna, Menghun,  750 m/ 1958.VI.1;  1♂, Xishuangbanna, Mengzhe,  870 m/ 1958.VII.10, Shuyong Wangcoll.;  1♂, Xishuangbanna, Menga,  1800 m/ 1958.VI.2, Fuji Pucoll.;  2♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Yiran Zhangcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.9, Facai Zhangcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.VI.6, Fuji Pucoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.4, Yiran Zhangcoll.;  1♂, Xishuangbanna, Menglun,  650 m/ 1959.VIII.25, Facai Zhangcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.4, Facai Zhangcoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.19, Facai Zhangcoll.;  1♂, Xishuangbanna, Menglun,  650 m/ 1964.IV.29, Baolin Zhangcoll.;  1♂, Cheli[= Jinghong],  620 m, 1957.IV.17, Lingchao Zangcoll.;  1♂, Baoshanto Yongping, 1955.V.28, Tianrong Huangcoll.;  1♂, Ruili, Dengga/ 1992.VI.8, Decheng Yuancoll.;  1♂, Lincang,  1000 m/ 1957.VII.26, Shuyong Wangcoll.;  1♂, Lincang,  1100 m/ 1957.VIII.8, Lingchao Zangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1958.V.30, Facai Zhangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1958.V.23, Facai Zhangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1958.V.22, Fuji Pucoll.;  1♀, Mengla/ 1982.IV.20, Shengqiao Jiangcoll.; 1♀, 1984.V.9, Menglun / Lilioceris impressa(Fabricius);  1♀, Xishuangbanna, Mengla,  620-650 m/ 1958.VI.6, Fuji Pucoll.;  1♀, Xishuangbanna, Mengzhe,  890 m/ 1958.VII.3, Fuji Pucoll.;  1♀, Xishuangbanna, Mengzhe,  890 m/ 1958.IX.5, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Fuji Pucoll.;  1♀, Xishuangbanna, Mengzhe,  890 m/ 1958.VII.8, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.VI.2, Fuji Pucoll./ Lilioceris semipunctata, det. Yuan Xu, 2021;  1♀, Xishuangbanna, Mengsong,  1600 m/ 1958.IV.23, Yiran Zhangcoll.;  1♀, Jinghong, 1979.X.30, Damenglong, Zuopei Lincoll./ Lilioceris latissima, det. Liang H.B. 2020;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.VI.5, Yiran Zhangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.20, Fuji Pucoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Fuji Pucoll.;  1♂, Xishuangbanna, Mengla,  620-650 m/ 1959.V.23, Fuji Pucoll.;  1♂, Xishuangbanna, Xiaomengyang, 1957.VI.15, Shuyong Wangcoll.;  1♂, Xishuangbanna, Mengzhe,  1200 m/ 1958.VI.14, Shuyong Wangcoll.;  1♂, Xishuangbanna, 1958.VII.8/ Mengzhe, Mengman,  870 m, Fuji Pucoll.;  1♂, Xishuangbanna, Mengzhe,  870 m/ 1958.IX.7, Shuyong Wangcoll.;  1♂, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.4, Fuji Pucoll.;  1♂, Xishuangbanna, Xiaomengyang,  850 m, 1957.VI.13, Shuyong Wangcoll.;  1♀, Xishuangbanna, Mengla,  620-650 m/ 1959.V.16, Fuji Pucoll.;  1♀, Lancang,  1000 m/ 1957.VII.26, Lingchao Zangcoll.;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.VIII.16, Shuyong Wangcoll.;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.V.10;  1♀, Xishuangbanna, Menglun,  650 m/ 1964.IV.14, Baolin Zhangcoll.;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.V.18, Fuji Pucoll.;  1♀, Xishuangbanna, Menga,  1050-1080 m/ 1958.V.18, Fuji Pucoll.;  1♀, Xishuangbanna, Xiaomengyang,  1100 m, 1957.V.6, Dahua Liucoll.;  1♂, Xishuangbanna, Mengzhe,  800 m/ 1958.VII.3, Fuji Pucoll.;  1♂, Xishuangbanna, Menghun,  750 m/ 1958.VI.2;  1♂, Ruili, Ruili Botanicalgarden, 24.07230°N, 97.81944°E/ 2012.X.27,  1152 m, Xinlei Huangcoll.;  1♀ 1♂, Xishuangbanna, 2020.VI.30, Ying Yancoll.;  1♀, Xishuangbanna, Guomen Shan, 2019.V.11, Kaiqin Licoll.;  1♀ 2♂, Xishuangbanna, Menghai, Menghun, 2011.IV.26, Hongbin Liangand Kaiqin Licoll.;  8♀ 5♂, Xishuangbanna, Menghai, Mannong Xinzhai, 21.78233°N, 100.50706°E,  1582 m, 2021.IV.3-8, Hongbin Liang, Yuan Xuand Neng Zhangcoll.;  1♂, Xishuangbanna, Guanlei, Nanlahe, 21.5796°N, 101.1879°E,  465 m, 2021.XI.30, Yong Wangcoll.;  2♀ 4♂, Xishuangbanna, Mengla, Longlin New village, 2020.VI.5-6, 21.52914°N, 101.49415°E,  1066 m, Hongbin Liangand Yuan Xucoll.;  3♀ 3♂, Xishuangbanna, Xiaomengyang, Baihua Shan, 22.17720°N, 100.92428°E,  876 m, 2021.IV.3, Hongbin Liangand Yuan Xucoll.;  3♀ 5♂, Xishuangbanna, Menghai, Nanlanghe, 22.17720°N, 100.92428°E,  876 m, 2020.IV.9, Hongbin Liangand Yuan Xucoll.;  1♀ 1♂, Baoshan, Baihualing, N 25.30499, 98.80008,  1622 m, 2020.IX.29, Yuan Xucoll.;  1♀, Baoshan, Baihualing, N 25.30499, 98.80008,  1622 m, 2019.X.2, Hongbin Liangand Yuan Xucoll.;  1♀, Cangyuan, Banhong, 2011.V.3, Kaiqin Licoll.;  1♀, Eshan, Ayi Power Station, 24.09573°N, 102.54589°E,  1444 m, 2022.VII.2, Neng Zhangcoll.;  1♀, Hekou, Huayudong, 22.67222°N, 103.93769°E,  97 m, 2021.IV.22, Hongbin Liangand Yuan Xucoll.;  3♀, Jianshui, Yunlongshan, 23.77582°N, 102.82258°E,  1645 m, 2021.IV.23, Hongbin Liangand Yuan Xucoll.;  1♀, Lvchun, Huanglian Shan, 2018.5.23, Kaiqin Licoll.;  3♀ 3♂, Xishuangbanna, Menghai, Nanlanghe, 22.17720°N, 100.92428°E,  876 m, 2020.VI.1, Hongbin Liangand Yuan Xucoll.;  16♀ 8♂, Xishuangbanna, Mengla, Longlin New village, 2020.III.30, 21.52914°N, 101.49415°E,  1066 m, Hongbin Liangand Yuan Xucoll.;  1♂, Xishuangbanna, Menghai, Mengsong, Sanmai, 21.99943°N, 100.61022°E,  1028 m, 2020.VI.2, Hongbin Liangand Yuan Xucoll.;  2♀, Ruili, Nongdao, 2019.X.10, Yuan Xucoll.;  1♂, Yingjiang, Nabang, 2012.IX.29, Hongbin Liangcoll.;  6♀ 3♂, Eshan, Ayi Power Station, 24.09573°N, 102.54589°E,  1444 m, 2021.IV.23, Hongbin Liangand Neng Zhangcoll.;  1♀, Zhenyuan, Enle, Najiu, 23.977826°N, 101.031797°E,  1359 m, 2022.VII.8, Yuan Xuand Neng Zhangcoll.;  2♀, Zhenyuan, Sanzhangtian, 2022.V.28, Neng Zhangcoll.;   Guangxi: 1♂, Yangshuo, Baisha/ 1963.VII.23, Shuyong Wangcoll.;  2♀, Baishou, 1952.6.23;  1♀, Baishou, 1952.7.2;  1♀, Fangcheng, Fulong,  240 m, 1998.IV.20, Gexia Qiaocoll.;  1♀, Longlin/ 1984.V.21, Shufang Wangcoll.;  1♀, Yangshuo, Baisha/ 1963.VII.23, Shuyong Wangcoll.;  1♀, Longzhou, Daqing Shan,  360 m, 1963.VII.23, Shuyong Wangcoll.;  1♀, Longzhou, Daqing Shan,  360 m, 1963.IV.18, Shuyong Wangcoll.;  1♀, Longzhou, Daqing Shan,  360 m, 1963.IV.27, Shuyong Wangcoll.;  1♀, Longzhou, Daqing Shan,  360 m, 1963.IV.27, Shuyong Wangcoll.;  1♀, Wuming, Daming Shan, 1963.V.21, Sikong Liucoll.;  1♂, Longzhou, Daqing Shan,  360 m, 1963.IV.25, Shuyong Wangcoll.;  1♂, Wuming, Daming Shan, 1963.V.24, Jikun Yangcoll.;  1♂, Longlin/ 1980.IX.9, Lin Dongcoll.;   Tibet: 1♀,  Medog, Bangxin, 29.48488°N, 95.43508°E,  1418 m, 2020.IX.13,  1♀,  Medog, Baibung Tea Farm, 29.26310°N, 95.20983°E,  1047 m, 2020.IX.7, Hongbin Liangcoll.;  1♀,  Medog, Baibung Tea Farm, 29.26310°N, 95.20983°E,  1047 m, 2021.VI.16, Hongbin Liangand Neng Zhangcoll.;  9♀ 7♂,  Medog, Baibung, Gelin, 29.23370°N, 95.17707°E,  1408 m, 2020.VI.11-15, Hongbin Liang, Yuan Xuand Neng Zhangcoll.;  1♀,  Medog, Dexing, Wenlang, 29.36709°N, 95.34012°E,  1251 m, 2020.IX.4, Hongbin Liangcoll.;  5♀ 5♂,  Medog, Baibung, Gelin, 29.23370°N, 95.17707°E,  1408 m, 2022.VII.17-20, Hongbin Liang, Yuan Xuand Neng Zhangcoll.;  1♀,  Medog, Lagong Tea Farm, 29.31879°N, 95.31570°E,  1294 m, 2021.VI.8, Yuan Xuand Neng Zhangcoll.;   Guizhou: 2♀ 1♂, 1912;  1♀, Xishui, Linjiang,  300 m/ IOZ and Guizhou Univ. Joint Expedition, 2000.VI.1, Song QZcoll.;  1♀, Libo, Maolan,  450 m, 1998.X.27, Xinke Yangcoll.;  1♂, Libo, Maolan,  450 m, 1998.X.27, Xingke Yangcoll.;  3♂, Maolan, 1998.V, Qiongzhang Songcoll./ Lilioceris semipunctata;  1♂, Libo, Maolan,  450 m, 1998.X.26, Wenzhu Licoll.;  1♂, Wangmo, 1982.VI;  1♂, Fanjing Shan, Huguotemple,  1350 m, 2001.VIII.03, Qiongzhang Songcoll.;  1♀, Libo,  500-700 m/ 1998.V.24-30/ Lilioceris semipunctata, Det. Peiyu Yu;  1♂, Libo, Xiaoqikong,  700 m, 1998.V.30, Runzhi Zhangcoll.;  1♂, Libo, Yonggui,  700 m, 1998.V.29, Runzhi Zhangcoll.;   Hainan: 1♀, Baisha, Yinggeling, Nankai, Daoyinto Zafu village, 2009.XI.22, Meiying Lincoll.;  1♀, Changjiang, Bawangstreet, light trap, 19.11104, 109.08168/  145 m, 2007.5.12, Hongbin Liangand Fuqiang Chencoll.;  1♀, Xinglong/ 1963.III.12, Baolin Zhangcoll.;  1♀, Tongzha,  340 m/ 1960.VII.31, Changqing Licoll.;  1♀, Tongzha,  340 m/ 1960.VII.31, Xuezhong Zhangcoll.;  2♀, Tongzha,  340 m/ 1960.VII.4, Suofu Licoll.; 1♀, 1934.10.27;  2♂, Bawangling, Dongliu, 19.054112, 109.19445/  635 m, 2007.5.11, Hongbin Liangcoll.;  1♂, Nada/ 1954.IV.27, Keren Huangcoll.;  1♂, Tongzha,  340 m/ 1960.VI.29, Xuezhong Zhangcoll.;  2♂, Tongzha,  340 m/ 1960.VIII.4-5, Suofu Licoll.;  1♂, Tongzha,  340 m/ 1960.VII.31, Suofu Licoll.;  2♂, Bawangling, 2019.IV.19, Run Zhoucoll.;  1♀ 1♂, Baisha, Gaofeng village, 19.04059°N, 109.31583°E,  886 m, 2020.VII.28, Yuan Xucoll.;  2♀, Baisha, Fangza village, 19.07599°N, 109.52348°E,  438 m, 2020.VII.29, Yuan Xucoll.;  1♀ 1♂, Nanfaling, 18.87220°N, 109.29446°E,  537 m, 2020.VII.26, Yuan Xucoll.;   Fujian: 1♀, Tongmuguan/ 1979.VIII.4, Fusheng Huangcoll./ Lilioceris semipunctata, det. Peiyu Yu/ Lilioceris rufimembris, det. Liang H.B.;  Vietnam:  1♀, Tonkin, Hoa Binh, leg. A. de Cooman, 1939.VII;  2♀, Tonkin, Hoa Binh, leg. A. de Cooman, 1940.VIII;  1♀, Tonkin, Hoa Binh, L. Ouport;  1♂, Tonkin, Hoa Binh, leg. A. de Cooman, 1940.VIII;  2♂, Tonkin, Hoa Binh, leg. A. de Cooman/ Lilioceris rufimembris, det. Liang H.B; 2♂, Tonkin.  Diagnosis. Anterior angles of pronotum protruding, pronotal disc with 2-5 irregular rows of fine punctures. Humeral groove of elytra distinct, punctures of elytra sparse and diminishing posteriorly, absent on apical 1/3. Femora of legs without tooth. Lateral metasternum with a long strip of pubescence.  Redescription. BL = 7.5-9.0 mm, BW = 3.0-3.8 mm. Bodybrownish red.  Head(Figs 21, 45A). Vertex slightly convex, with a shallow groove in the middle, sparse punctate and pubescent laterally; frontoclypeal area triangular, lateral disc with sparse punctures and pubescent; labrum transverse, with sparse pubescent; antennae nearly 1/2 of body length, antennomeres 1-4 nearly globular, antennomere 2 shortest, antennomeres 5-10 nearly 1.1 xas long as wide, antennomere 11 slender.  Pronotum(Figs 21, 45B). Anterior angles protruding, posterior angles not protruding; sides slightly constricted in the middle; disc slightly raised; middle of disc with 2-5 rows of fine punctures. Scutellum triangular and pubescent.  Elytra(Figs 21, 35B). Humeri protruding, humeral groove and basal impression distinct; elytra without completely punctate striae, punctures sparse and large in the base, diminishing posteriorly, absent on apical 1/3; intervals smooth; epipleura raised, with a sparse row of fine punctures.   Mesosternum pubescent.Lateral metasternum with a long and straight strip of pubescence, metepisternum densely pubescent (Fig. 45C).  Abdominal sternite(Fig. 35A). Lateral transverse impressions small and indistinct on sternites 1-4, other areas of sternites 1-4 and all of sternite 5 densely pubescent.  Leg(Fig. 35C). Femora with dense pubescence on the dorsal surface, with sparse pubescence on the ventral surface, without tooth.  Male genitalia(Fig. 50A-D). Ostium occupying 1/4 length of median lobe (Fig. 50A); apex hooked (Fig. 50B); tegmen Y-shaped, basal piece of tegmen triangular and broad, lateral lobes slightly sclerotized and combined with second connecting membrane; internal sac membranous, with distinct dorsal, median and ventral sclerites, posterior part of dorsal sclerite paralleled in dorsal view, ventral sclerite short and flaky, median sclerite distinctly sclerotized (Fig. 50C, D).  Female reproductive organs(Fig. 62A-C). Tergites 8 and 9, sternites 8 and 9 sclerotized, posterior areas of tergite 8, sternite 8, and apodemes with pubescence, spiculum gastrale Y-shaped and long; ovipositor with dense pubescence, distal part of ovipositor cylindrical, long and with a protuberance; spermatheca simple and hooked.  Distribution  (Fig. 67).China (Fujian, Guangxi, Guizhou, Hainan Tibet, Yunnan); Indonesia (Java, Sumatra); Nepal; India ( Kimoto and Takizawa 1973;  Bezdekand Schmitt 2017).  Host plant and habitat  (Fig. 73A, B).Host plant of this species is  Smilax bracteataPresl. and  Lilioceris semipunctatashares the same habitat as  L. latissimaaccording to our observations in Baihua Shan of Xishuangbanna, Yunnan.  Remark.  Fabricius (1801: 472) described  Lilioceris semipunctatabased on specimens from Sumatra. We examined three syntype photographs of  L. semipunctata. They obviously belong to two different species: the specimen with the identification label and locality is here designated as lectotype (Figs 21, 22). The other two specimens (TYPE / zmuc 00031032; TYPE / zmuc 00031031) with no label and no locality are here excluded from the type series. These two specimens with a glabrous metasternum are clearly different from the lectotype, but identical to  Lilioceris latissima.  Lacordaire (1845: 559) redescribed  Lema semipunctatabased the specimens from Java, and indicated that  Lema dehaanii( Guerin-Meneville, 1844) was completely similar to  L. semipunctata, and moved it to the genus  Crioceris. Then Kimoto and Gressitt (1979: 230) synonymized  L. dehaaniiwith  L. semipunctata, and subsequent researchers followed this treatment. We also examined a syntype of  Lilioceris rufimembris(Pic, 1921) in NHHN, and no significant morphological differences were found from the lectotype of  L. semipunctata, except for two rows fine punctures on pronotum (Fig. 23), which are absent in the lectotype of  L. semipunctata(Fig. 21). In IZCAS, there are more than 150 specimens from Yunnan that perfectly match these two types of punctures on the pronotum. We compared all these specimens and found that there were no significant morphological differences including male genitalia. Therefore, we conclude that  L. semipunctataand  L. rufimembrisare conspecific, and that the pronotal punctures in  L. semipunctatacan be variable.   Figures 23-26.Habitus of  Liliocerisspp. 23, 24  L. rufimembris, syntype, China (Yunnan), photographed by Christophe Rivier (MNHN) 25, 26  L. yuae, holotype, China (Hainan), photographed by Yuan Xu. Scale bars: 5.0 mm. 2024-01-01 2024-12-31 2024-01-01 China Yunnan 451 Yunnan 1957-05-06 Dahua Liu China 1000 Xiaomengyang 2 2 Yunnan 1959-07-09 Yiran Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1964-04-14 Baolin Zhang China 650 Yiwu 1 1 Yunnan 1958-07-25 Shuyong Wang China 1400 Menghai Xishuangbanna 1 1 Yunnan 1958-04-29 Yiran Zhang China 710 Jinghong Xishuangbanna 1 1 Yunnan 1958-07-11 China 650 Damenglong Xishuangbanna 1 1 Yunnan 1958-08-19 Fuji Pu China 1065 Menga Xishuangbanna 1 1 Yunnan 1958-05-31 Chunpei Hong China 750 Xishuangbanna 2 2 Yunnan 1958-04-30 Shuyong Wang China 1065 Menga Xishuangbanna 2 2 Yunnan 1958-04-30 Fuji Pu China 1065 Menga Xishuangbanna 1 1 Yunnan 1958-08-31 China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1959-05-08 Fuji Pu China 635 Mengla Xishuangbanna 1 1 Yunnan 2024-01-01 2024-12-31 2024-01-01 China 650 Damenglong Xishuangbanna 1 1 Yunnan 1958-08-10 Shuyong Wang China 1065 Menga Xishuangbanna 1 1 Yunnan 2024-01-01 2024-12-31 2024-01-01 China 650 Damenglong Xishuangbanna 1 1 Yunnan 1958-05-06 Fuji Pu China 1065 Menga Xishuangbanna 1 1 Yunnan 1958-07-07 Fuji Pu China 1890 Mengzhe Xishuangbanna 1 1 Yunnan 1957-06-21 Shuyong Wang China 850 Xiaomengyang 1 1 Yunnan 1959-07-09 Yiran Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1958-07-20 Yiran Zhang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1959-07-10 Yiran Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1959-06-06 Fuji Pu China 635 Mengla Xishuangbanna 1 1 Yunnan 1959-05-19 Facai Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1959-05-08 Fuji Pu China 635 Mengzhe Xishuangbanna 1 1 Yunnan 1958-07-03 Fuji Pu China 850 Mengzhe Xishuangbanna 1 1 Yunnan 1958-07-04 Shuyong Wang China 850 Mengzhe Xishuangbanna 1 1 Yunnan 1958-06-14 Shuyong Wang China 1200 Mengzhe Xishuangbanna 2 2 Yunnan 1958-07-04 Zhizi Chen China 875 Mengzhe Xishuangbanna 1 1 Yunnan 1959-05-16 Facai Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1957-06-09 Shuyong Wang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1957-06-25 Shuyong Wang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1958-09-03 Shuyong Wang China 870 Mengzhe Xishuangbanna 2 2 Yunnan 1958-06-14 Shuyong Wang China 1200 Mengzhe Xishuangbanna 1 1 Yunnan 1958-07-01 Fuji Pu China 875 Mengzhe Xishuangbanna 1 1 Yunnan 1959-05-12 Facai Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1959-05-22 Fuji Pu China 635 Mengla Xishuangbanna 1 1 Yunnan 1959-06-29 Facai Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1958-07-25 Leyi Zheng China 1600 Mengsong Xishuangbanna 1 1 Yunnan 1958-09-24 Leyi Zheng China 1600 Mengsong Xishuangbanna 1 1 Yunnan 1958-07-11 Fuji Pu China 870 Mengzhe Xishuangbanna 1 1 Yunnan 1980-05-01 Zuyun Guo China 1060 Menghai 1 1 Yunnan 1958-05-30 China 750 Menghun Xishuangbanna 1 1 Yunnan 1958-06-03 China 850 Menghun Xishuangbanna 1 1 Yunnan 1958-06-01 Shuyong Wang China 800 Menga Xishuangbanna 1 1 Yunnan 1957-09-03 Shuyong Wang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1958-04-21 Xuwu Meng China 650 Damenglong Xishuangbanna 1 1 Yunnan 2024-01-01 2024-12-31 2024-01-01 Xinlei Huang China Ruili 4990 23.98207 Wudian 1 97.61129 Nongdao 1 1 Yunnan 2008-08-06 Jingxin Liu China 22.16713 Nabanhe Reserve 1 100.65887 1 1 Yunnan 2005-08-22 Liang HB, Zhang JF China 1177 26.86203 Shangpa Town 1 98.87142 Fugong 1 1 Yunnan 1959-05-12 Facai Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1958-08-15 Fuji Pu China 1065 Mengla Xishuangbanna 1 1 Yunnan 1958-08-20 Fuji Pu China 1065 Mengla Xishuangbanna 1 1 Yunnan 1958-08-16 Shuyong Wang China 1065 Mengla Xishuangbanna 1 1 Yunnan 1957-06-18 Shuyong Wang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1957-07-17 Shuyong Wang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1957-07-12 Shuyong Wang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1957-07-09 Shuyong Wang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1957-06-27 Lingchao Zang China 850 Xiaomengyang Xishuangbanna 1 1 Yunnan 1964-04-14 Baolin Zhang China 650 Menglun Xishuangbanna 1 1 Yunnan 1957-04-17 Fuji Pu China 540 Ganlanba Xishuangbanna 1 1 Yunnan 1955-05-28 Tianrong Huang China Yongping Baoshan 1 1 Yunnan 1958-08-02 China Wenshan 1 1 Yunnan 1958-10-28 Shuyong Wang China 1300 Menghun Xishuangbanna 1 1 Yunnan 1958-06-01 Shuyong Wang China 750 Menghun Xishuangbanna 2 2 Yunnan 1958-06-05 Chunpei Hong China 750 Menghun Xishuangbanna 3 3 Yunnan 1958-09-03 Shuyong Wang China 800 Menga Xishuangbanna 1 1 Yunnan 1958-08-16 Shuyong Wang China 1065 Menga Xishuangbanna 1 1 Yunnan 1958-07-09 Fuji Pu China 635 Mengla Xishuangbanna 1 1 Yunnan 1958-05-30 Facai Zhang China 635 Mengla Xishuangbanna 1 1 Yunnan 1958-04-07 China 650 Damenglong Xishuangbanna 1 1 Yunnan 1959-05-31 Suofu Li China 635 Mengla Xishuangbanna 1 1 Yunnan 1959-05-08 Fuji Pu China 635 Mengla Xishuangbanna 1 1 Yunnan 1979-09-02 Mengqiao, Leyi Zheng China Ruili 1 1 Yunnan 1929-04-02 Xiaohong Ou China Tengchong 1530 Wujia Jidi Jietou 1 1 Yunnan 1958-06-09 Shuyong Wang China 650 Damenglong Xishuangbanna 1 1 Yunnan 1958-04-23 Yiran Zhang China 1600 Mengsong Xishuangbanna 1 1 Yunnan 1958-04-28 Fuji Pu China 1600 Mengsong Xishuangbanna 1 1 Yunnan 1957-09-23 Lingchao Zang China 2600 Kongming Shan Xishuangbanna 1 1 Yunnan 1957-07-26 Shuyong Wang China 1000 Lancang 2 2 Yunnan 1957-08-08 Lingchao Zang China 1100 Lancang 1 1 Yunnan 1958-06-01 Fuji Pu China 1065 Menga Xishuangbanna 1 1 Yunnan 1957-08-02 Lingchao Zang China 1100 Lancang 1 1 Yunnan 2024-01-01 2024-12-31 2024-01-01 China Shuangjiang 1 1 Yunnan 1957-05-09 Menchalsky, Chinese China Jingdong 1 1 Yunnan 1958-06-15 Xuwu Meng China 650 Menghun Xishuangbanna 1 1 Yunnan 1958-05-16 Fuji Pu China 1000 Menga Xishuangbanna 1 1 Yunnan 1959-05-23 Fuji Pu China 635 Mengla Xishuangbanna 1 1 Yunnan 2006-06-01 Liang HB, Hu P China Tengchong 1470 24.94861 Rehai 1 98.45181 Qingshui 1 1 Yunnan 2024-01-01 2024-12-31 2024-01-01 Chinese Acad. 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