Ebulea obliquata Leucocraspeda nissoralis A revision of the genus Ecpyrrhorrhoe Huebner, 1825 from China based on morphology and molecular data, with descriptions of five new species (Lepidoptera, Crambidae, Pyraustinae) Xiang, Lanbin Chen, Kai Chen, Xiaohua Duan, Yongjiang Zhang, Dandan ZooKeys 2022 2022-03-23 1090 1 44 974A77DA-373D-57BB-A33D-D9E2AE5E3FBA (Moore, 1888) Xiang & Chen & Chen & Duan & Zhang 2022 Moore 1888 Insecta Crambidae Ecpyrrhorrhoe CoL Animalia Ecpyrrhorrhoe obliquata Lepidoptera 0 1 Arthropoda species obliquata comb. nov.   Figs 8 , 25 , 40   Ebulea obliquataMoore, 1888: 224.  Leucocraspeda nissoralisSwinhoe, 1894: 145.  Diagnosis. In appearance,  Ecpyrrhorrhoe obliquatais similar to  E. exigistriain the brown subterminal area of the wings, as well as in the oblique, dark brown streak on forewing which distinguish them from all other species of the genus. Genitalia characters readily distinguish  E. obliquatafrom other members of the genus in the semi-circular sella with basal half of ventral margin bearing curved spines. It can be best distinguished from  E. exigistriaby the larger forewing length (forewing length: 10.0-12.0 mm), the more distinct and longer streak of the forewing, in the male genitalia (Fig. 25) by the shape of sella, and the weakly sclerotized arms of the bifid juxta, which is without teeth, and in the female genitalia (Fig. 40) by the shape of antrum.  Redescription (Fig. 8). Head.Frons and vertex yellow, frons with lateral white bands. Labial palpus yellowish brown, contrastingly white at base ventrally. Maxillary palpus yellowish brown, white terminally. Thorax.Dorsal side yellow and ventral side white; tegula yellow with brown base. Wings.Forewing length: 10.0-12.0 mm. Forewing pale yellow, basal half of costal band and posterior half of subterminal area brown; stigmata and lines dark brown; antemedial line strongly oblique from 1/5 of costa to 1/2 of 1A, then dentated to 2/5 of posterior margin; orbicular stigma dot-shaped, sometimes indistinct; reniform stigma comma-shaped, thick and straight; an oblique, dark brown streak from posterior end of reniform stigma weakly curved and extended to tornus; postmedial line obliquely outwards from 3/5 of costa to 1/2 of M1, then dentated inwards to 2/3 of posterior margin; terminal line dark brown and intermittent; fringe with basal half yellow and blackish brown, distal half with anterior half yellow and posterior half dark brown. Hindwing pale yellow, subterminal area from dark brown to brown, gradually paler to tornus; postmedial line dark brown, slightly dentate from 2/3 of M1 to 1/2 of CuA1, then bending inwards to base of CuA1, finally undulated to 2/3 of posterior margin; terminal line and fringe as in forewing. Abdomen.Pale yellow dorsally, each segment with posterior margin whitish, black on distal end; white ventrally.  Male genitalia  (Fig. 25).Uncus with basal 1/3 nearly triangular and naked. Valva with costa nearly straight or concave and ventral margin curved, basal part weakly narrowed, remainder of even width, apex obtusely rounded; sella nearly semi-circular, bearing short and curved spines on distal half of ventral margins; sacculus with middle part inflated into a thumb-shaped protrusion. Juxta with base wide, distal 3/4 bifid into pointed arms. Saccus narrowly triangular. Phallus long and straight, with a bundle of short spines assembling into cone-shape at distal end.  Female genitalia  (Fig. 40).Anterior apophysis ~ 2 xlength of posterior apophysis. Antrum cup-shaped, with anterior 1/3 strongly sclerotized and covered with spinules on inner wall, posterior 1/3 abruptly broad and partly wrinkled; colliculum long; ductus seminalis connecting to anterior end of colliculum and with a short sclerite located opposite to colliculum; ductus bursae ~ 2.5-3 xas long as diameter of corpus bursae, basal 1/3 bearing a slim sclerite. Corpus bursae globular; rhomboid signum with maximal length approximately 1/2 as long as diameter of corpus bursae; the second (posterior) signum nearly thumb-shaped bearing a wide base, sparsely covered with long spines.  Material examined.  Type material.  Leucocraspeda nissoralis:  Lectotype(designated here) ♂, Kahsia Hs [Hills]. 94-66, PyralidaeBrit. Mus. Slide No. 10897 (NHMUK);    Paralectotype(designated here): 1♂, [  India]: Cherre Punji(NHMUK).  Other material examined.    China: Zhejiang: 1♂, Mt. Jiulongshan, 5.VIII.2011, Fu Xiaobingleg.;   Jiangxi: 2♂, Mt. Jiulianshan, Longnan, 24.58°N, 114.43°E, alt.  620 m, 26.IX.2016, 24.IX.2017, Chen Kai, Duan Yongjiangleg.;   Hunan: 1♂, Xijialing, Mt. Shunhuangshan, Xinning, Shaoyang, 26.43°N, 111.01°E, alt.  1000 m, 6.IX.2020, Jin Mengjie, Xiang Lanbinleg.;  6♂ 2♀, Mt. Shunhuangshan, Xinning, Shaoyang, 26.40°N, 111°E, alt.  810 m, 4-6.IX.2020, Jin Mengjie, Xiang Lanbinleg.;  1♂, Dupangling National Natural Reserve, Dao County, Yongzhou, 25.48°N, 111.37°E, alt.  430 m, 29-30.VIII.2020, Jin Mengjie, Xiang Lanbinleg.;  1♀, Dupangling National Natural Reserve, Dao County, Yongzhou, 25.49°N, 111.39°E, alt.  350 m, 28-31.VIII.2020, Jin Mengjie, Xiang Lanbinleg.;  1♂, Qiaotoupu, Chengbu, Shaoyang, 26.25°N, 110.38°E, alt.  640 m, 8.IX.2020, Jin Mengjie, Xiang Lanbinleg.;   Guangdong: 2♂ 1♀, Heishiding, Fengkai, 15.VI.2009, 9.X.2010, 2.VIII.2011, 9.VII.2017, 23.47°N, 111.90°E, alt. 214 m, Zhang Dandan et al. leg., genitalia slide No. LJW121067 ( ♂), SYSU0239 ( ♂), SYSU1236 ( ♀, molecular voucher no. LEP0297);  1♀, Yanshuitian, Fengkai, 6.IX.2011, Yang Lijun, Liao Junleileg., genitalia slide No. SYSU0269;  2♂, Mt. Danxiashan, Shaoguan, 25.04°N, 113.64°E, alt.  96 m, 6-7.VI.2012, Li Jinweileg., genitalia slide No. LJW12075, CXH12186;  1♀, Mt. Nankunshan, Huizhou, 16.VII.2003, Zhang Dandan, Li Zhiqiangleg., genitalia slide No. ZDD03057;  3♂ 2♀, Hongri Village, Mt. Nankunshan, Huizhou, 6-9.XI.2020, Jin Mengjieleg.;   Hainan: 2♂, Bawangling Natural Reserve, 8.V.2011, Yang Lijunleg., genitalia slide No. LJW12101;  2♂, Yinggeling Natural Reserve, 19.05°N, 109.50°E, alt.  954 m, 4.IX.2013, Chen Xiaohualeg., genitalia slide No. CXH12212, SYSU0228 (molecular voucher No.LEP0034);  2♂, Hongkan, Yinggeling Natural Reserve, 19.08°N, 109.50°E, alt.  508 m, 15-16.VI.2015, Cong Peixin, Guan Wei, Hu Shaleg. (NKU);  1♂, Limushan ForestPark, 19.17°N, 109.73°E, alt.  607 m, 25.VII.2014, Cong Peixin, Liu Linjie, Hushaleg. (NKU);  1♂, Wuzhishan ForestPark, 18.88°N, 109.67°E, alt.  766 m, 9.I.2016, Teng Kaijian, Bai Xia, Chen Mengtingleg. (NKU);   Guangxi: 1♂, Nonggang, Longzhou, 22.47°N, 106.96°E, alt.  271 m, 19.IV.2012, Li Jinweileg., genitalia slide No. CXH12185;  1♂, Hekou, Dayaoshan Natural Reserve, Jinxiu, 24.14°N, 110.09°E, alt.  823 m, 20.VII.2015, Qi Mujie, Zhao Shengnanleg. (NKU);  2♂, Mt. Jiuwanshan, Hechi, alt.  1600 m, 23.VII.2015, Wang Jipingleg.;  1♂, Technology Building, Huaping Natural Reserve, Guilin, 25.63°N, 109.91°E, alt.  760 m, 10-12.IX.2020, Jin Mengjie, Xiang Lanbinleg.;   Yunnan: 1♂, Taiyanghe Natural Reserve, alt.  1450 m, 23.VIII.2014, Zhang Zhenguoleg. (NKU);  1♂, Liaowangtai, Taiyanghe ForestPark,  Pu'er, 22.60°N, 101.11°E, alt.  1626 m, 8.VII.2013, LiuShurong, Wang Yuqi, Teng Kaijianleg. (NKU);   Sichuan: 1♂, Nuoshuihe Natural Reserve, Tongjiang, alt.  700 m, 5.VII.2013, He Guiqing, Xu Lijunleg.;   Chongqing: 1♂, Daheba, Mt. Jinfoshan, alt.  800-850 m, 15.VII.2010, Du Xicui, Song Lifangleg.;  1♂, Tudiyan, Mt. Simianshan, alt.  1200 m, 9.VIII.2011, He Guiqing, Song Lifangleg.;   Tibet: 1♂, Air-raidshelter, Beibeng, Medog, 29.24°N, 95.17°E, alt.  750 m, 31.VII.2018, Qi Mujieleg. (NKU);  1♂, Gelin, Beibeng, Medog, 29.25°N, 95.19°E, alt.  1063 m, 29.VII.2018, Qi Mujieleg. (NKU);  1♂, Yadong, Medog, 29.33°N, 95.34°E, alt.  833 m, 2.VIII.2018, Qi Mujieleg.(NKU).  Distribution. China (Zhejiang, Jiangxi, Hunan, Guangdong, Hainan, Guangxi, Yunnan, Sichuan, Chongqing, Tibet), Burma, India, Sri Lanka. 2022-01-01 2022-12-31 2022-01-01 India Cherre Punji 1 1 Paralectotype 2011-08-05 Fu Xiaobing China Mt. Jiulongshan 1 1 Zhejiang 2022-01-01 2022-12-31 2022-01-01 Chen Kai, Duan Yongjiang China 620 24.58 Longnan 750 114.43 Mt. Jiulianshan 2 2 Jiangxi 2020-09-06 Jin Mengjie, Xiang Lanbin China Xijialing 1000 26.43 Xinning 745 111.01 Mt. Shunhuangshan 1 1 Hunan 2020-09-04 2020-09-06 2020-09-04 Jin Mengjie, Xiang Lanbin China Mt. Shunhuangshan 810 26.4 Shaoyang 0 111.0 Xinning 8 2 6 Hunan 2020-08-29 2020-08-30 2020-08-29 Jin Mengjie, Xiang Lanbin China Dao County 430 25.48 Yongzhou 748 111.37 Dupangling National Natural Reserve 1 1 Hunan 2020-08-28 2020-08-31 2020-08-28 Jin Mengjie, Xiang Lanbin China Dao County 350 25.49 Yongzhou 748 111.39 Dupangling National Natural Reserve 1 1 Hunan 2020-09-08 Jin Mengjie, Xiang Lanbin China Qiaotoupu 640 26.25 Shaoyang 746 110.38 Chengbu 1 1 Hunan 2022-01-01 2022-12-31 2022-01-01 China Fengkai Heishiding 3 1 2 Guangdong 2011-09-06 Yang Lijun, Liao Junlei China Fengkai Yanshuitian 1 1 Guangdong 2012-06-06 2012-06-07 2012-06-06 Li Jinwei China 96 25.04 Shaoguan 749 113.64 Mt. Danxiashan 2 2 Guangdong LJW12075, CXH12186 2022-01-01 2022-12-31 2022-01-01 China No. 1 Guangdong 2003-07-16 Zhang Dandan, Li Zhiqiang China Huizhou Mt. Nankunshan 1 1 Guangdong ZDD03057 2022-01-01 2022-12-31 2022-01-01 China 1 Guangdong 2020-11-06 2020-11-09 2020-11-06 Jin Mengjie China Hongri Village Huizhou Mt. Nankunshan 5 2 3 Guangdong 2011-05-08 Yang Lijun China Bawangling Natural Reserve 2 2 Hainan LJW12101 2022-01-01 2022-12-31 2022-01-01 China 1 Hainan 2013-09-04 Chen Xiaohua China 954 19.05 Yinggeling Natural Reserve 764 109.5 2 2 Hainan CXH12212 2022-01-01 2022-12-31 2022-01-01 No. China No. 1 Hainan 2015-06-15 2015-06-16 2015-06-15 Cong Peixin, Guan Wei, Hu Sha China 508 19.08 Yinggeling Natural Reserve 764 109.5 Hongkan 2 2 Hainan 2014-07-25 Cong Peixin, Liu Linjie, Husha China 607 19.17 Limushan Forest 763 109.73 1 1 Hainan 2016-01-09 Teng Kaijian, Bai Xia, Chen Mengting China 766 18.88 Wuzhishan Forest 764 109.67 1 1 Hainan 2012-04-19 Li Jinwei China 271 22.47 Longzhou 756 106.96 Nonggang 1 1 Guangxi CXH12185 2022-01-01 2022-12-31 2022-01-01 China 1 Guangxi 2015-07-20 Qi Mujie, Zhao Shengnan China Hekou 823 24.14 Jinxiu 751 110.09 Dayaoshan Natural Reserve 1 1 Guangxi 2015-07-23 Wang Jiping China 1600 Hechi Mt. Jiuwanshan 2 2 Guangxi 2020-09-10 2020-09-12 2020-09-10 Jin Mengjie, Xiang Lanbin China Technology Building 760 25.63 Guilin 747 109.91 Huaping Natural Reserve 1 1 Guangxi 2014-08-23 Zhang Zhenguo China 1450 Taiyanghe Natural Reserve 1 1 Yunnan 2013-07-08 LiuShurong, Wang Yuqi, Teng Kaijian China Liaowangtai 1626 22.6 Pu'er 755 101.11 Taiyanghe Forest 1 1 Yunnan 2013-07-05 He Guiqing, Xu Lijun China 700 Tongjiang Nuoshuihe Natural Reserve 1 1 Sichuan 2010-07-15 Du Xicui, Song Lifang China 825 Mt. Jinfoshan Daheba 1 1 Chongqing 2011-08-09 He Guiqing, Song Lifang China 1200 Mt. Simianshan Tudiyan 1 1 Chongqing 2018-07-31 Qi Mujie China Air-raid 750 29.24 Medog 736 95.17 Beibeng 1 1 Tibet 2018-07-29 Qi Mujie China Gelin 1063 29.25 Medog 736 95.19 Beibeng 1 1 Xizang 2018-08-02 Qi Mujie China 833 29.33 Medog 736 95.34 Yadong 1 1 Xizang