Canthon cyanocephalus Harold, 1868: 13 Harold 1869: 990 Bates 1887: 28 A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini) Silva, Fernando A. B. Valois, Marcely Zootaxa 2019 2019-07-08 4629 3 301 341 4W3LF (Harold, 1868) Harold 1868 [151,721,153,180] Insecta Scarabaeidae Scybalocanthon Animalia Coleoptera 18 319 Arthropoda species cyanocephalus  ( Figs. 1G, 4G, 5F, 6B)     Canthon cyanocephalus Harold, 1868: 13, 53 (identification key and description);  Harold 1869: 990(catalog);  Bates 1887: 28(taxonomic remarks); Gillet 1911: 29(catalog); Schmidt 1922: 65, 74 (diagnosis); Balthasar 1939: 191(redescription);  Krajcik 2006: 26(catalog); Krajcik 2012: 63(catalog).   Diagnosis. Specimens of  S. cyanocephalusare readily distinguishable from all other species by the pronotum with a longitudinal dark band on midline ( Fig. 1G); elytra bicolored, with black spot on the anterior portion ( Fig. 1G); the eighth elytral stria lacking carina on anterior portion; aedeagus with parameres elongate, subrectangular, not excavated ventrally ( Fig. 4F); and endophallus lacking bristles or microbristles close to the FLP sclerite ( Fig. 5G).   Description. Body.Oval-elongate, with variable metallic sheen. Surface completely microgranulate. Color. Most of pronotum, disc of elytra, and ventral surface of body yellow or brown. Head, margin of pronotum and elytra, longitudinal dark band on midline of pronotum, semicircular band at the humeral portion of elytra black with green sheen. Length. 5.8–6.9 mm. Thorax. Anterior angles of pronotum acute (approximately 85°). Lateral margin irregularly curved outward, forming an obtuse angle at the middle portion. Elytra. Striae thin and shiny, punctures almost inconspicuous. Eighth stria lacking carina at the anterior portion. Aedeagus.Parameres slightly asymmetrical ( Fig. 4G). Dorsal margin of parameres curved inward from the basal to medial portions and curved outward at the apical portion. Ventral margin of parameres substraight, with a short denticle at the basal portion. SRP circular, with substraight handle-shaped extension ( Fig. 5F). FLP I-shaped ( Fig. 5F). A+SA with two superposed and elongate sclerites ( Fig. 5F). ASI-shaped ( Fig. 5F).   Type material.  Canthon cyanocephalusHarold,   lectotype ♂( here designated; MNHU): “  cyanocephalus/ Harold/ Orinoco” // “  Chaeridium/ nigrocinetum/ Mont./ Orinoco/ Mont.” // “ 26363” // “Type” // “ LECTOTYPE ♂/  Canthon/  cyanocephalus/ Harold/ des. F. Z. Vaz-de-Mello”.  Paralectotype   ♂( MNHU): “ cyanoce/ phalus Har.”// “ 26363” // “Type” // “ PARALECTOTYPE/  Canthon/  cyanocephalus/ Harold/ des. F. Z. Vaz-de-Mello”.  Non-type material [ 22 males, 22 females]: BRAZIL: AMAPÁ: Amapari, Rio Mariru ( 51°57′13″W, 0°46′10″N), N. Degallier— 1♂( CEMT). PARÁ: Monte Dourado, rainforest (52°67’W, 00°85’S), iv.2006, pitfall dung, T.A. Gardner & M.I.M. Hernández— 1♂( CEMT); Monte Dourado, Eucaliptusforest ( 52°51’W, 0°57’S), 19.v. 2008, 180 m, E. Bordoni— 2♂ 2♀( CEMT).  FRENCH GUIANA: Cayenne ( 4°54’3’’N, 52°18’12’’W), x.1918, [anonymous]— 1♂( CMNC).  TRINIDAD AND TOBAGO: COUVA-TABAQUITE-TALPARO, Quesnell Farm, 13 kmS Arima, 2 kmN Talparo, 50 m( 10°31’48’’N, 61°16’26’’W), 12–22.vi.1993, S. & J. Peck, rainforest, flight interception trap (1993-28)— 1♂ 1♀( CMNC); Quesnell Farm, 13 kmS Arima, 2 kmN Talparo, 50 m( 10°31’48’’N, 61°16’26’’W), 22.vi–8.vii.1993, S. & J. Peck, rainforest, flight interception trap (1993-46)— 1♀( CMNC); EAST- ERN TOBAGO, Gilpin Trace, 10 kmNE Roxborough, 400–500 m( 11°17’N, 60°36’W), 26–30.vi.1993, S. & J. Peck, flight interception trap (1993-50)— 1♂ 3♀( CMNC); SAN JUAN-LAVANTILLE, above Loango Village, Maracas Valley, 600 m( 10°44’1’’N, 61°26’10’’W), 9–22.vi.1993, S. & J. Peck, mountain rainforest, flight interception trap (1993-22)— 1♀( CMNC); above Loango Village, Maracas Valley, 600 m( 10°44’1’’N, 61°26’10’’W), 22.vi–6.vii.1993, S. & J. Peck, mountain rainforest, flight interception trap (1993-43)— 1♂( CMNC); Maracas Bay ( 10°45’13’’N, 61°25’58’’W), 23–30.vi.1976, J.D. Glaser— 1♂( CMNC); Simla Research Station, 8 kmN Arima, 240 m( 10°42’20’’N, 61°17’40’’W), 6–10.vi.1993, S. & J. Peck, tropical forest, flight interception trap (1993- 11)— 1♂( CMNC); TUNAPUNA-PIARCO, Andrews Trace, 16 kmN Arima, 620 m( 10°43’45’’N, 61°18’40’’W), 7–24.vi.1993, S. & J. Peck, upper mountain rainforest, flight interception trap (1993-14)— 2♂ 1♀( CMNC); Blan- chisseuse Road, 610 m( 10°43’N, 61°18’W), 14.ii.1987, M.S. Adams— 1♀( CMNC); Lalaja Trace, 19 kmN Arima, 650 m( 10°44’26’’N, 61°18’41’’W), 8–24.vi.1993, S. & J. Peck, mountain rainforest, flight interception trap (1993- 20)— 2♂ 2♀( CMNC); Lalaja Trace, 19 kmN Arima, 650 m( 10°44’26’’N, 61°18’41’’W), 24.vi–7.vii.1993, S. & J. Peck, mountain rainforest, flight interception trap (1993-59)— 1♂( CMNC); Morne Bleu, 823 m( 10°43’N, 61°17’W), 4.viii.1969, H. & A. Howden— 1♀( CMNC); Mount Saint Benedict, Tunapuna, 550 m( 10°40’51’’N, 61°23’36’’W), 5–21.vi.1993, S. & J. Peck, summit rainforest, flight interception trap (1993-06)— 1♂ 2♀( CMNC); Mount Saint Benedict, Tunapuna, 550 m( 10°40’51’’N, 61°23’36’’W), 5.vi.1993, S. Peck, treebase litter and seeds, tree base litter (1993-07)— 1♂( CMNC); Simla Research Station, 8 kmN Arima, 260 m( 10°42’20’’N, 61°17’40’’W), 6–14.vi.1993, S. & J. Peck, tropical forest, flight interception trap (1993-09)— 1♂( CMNC); Simla Research Sta- tion, ArimaValley, North Range, 244 m( 10°41’34’’N, 61°17’22’’W), 24–29.v.2000, A.S. Ramsdale, premontane rainforest, pitfall trap, dung— 1♀( CMNC); Simla, 5 mi.N Arima( 10°43’7’’N, 61°17’58’’W), 18.viii.1969, H. & A. Howden— 1♂ 1♀( CMNC); Simla, 5 mi.N Arima( 10°43’7’’N, 61°17’58’’W), 20.viii.1969, H. & A. Howden— 1♂( CMNC).  VENEZUELA: BOLÍVAR, 26 kmN Guasipati ( 7°41’35’’N, 61°57’19’’W), 24.vi–12.vii.1987, S. & J. Peck, sandy seasonally humid forest, flight interception trap— 1♂ 4♀( CMNC); Anacoco ( 6°43’0’’N, 61°8’00’’ S), 6.viii.2006, Curso NM 2006– 1♀( CEMT); Anacoco ( 6°43’0’’N, 61°8’00’’ S), 6.viii.2006, NM Curso 2007– 1♂( CEMT).   Distribution.Known from Brazil( Amapá, Pará), French Guiana, Trinidad and Tobagoand Venezuela( Fig. 6B). Endemism areas: Brazilian sub-region: Boreal Brazilian dominion: Guianan Lowlands and Roraima provinces; Mesoamerican dominion: Trinidadprovince (see Morrone 2014; fig. 12). 2847261438 MNHU F. Z. Vaz-de-Mello 18 319 2 2 lectotype 2847261354 MNHU F. Z. Vaz-de-Mello 18 319 1 1 paralectotype