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        <dc:title>Übersicht über die orientalischen Arten der Gattung Stenus LATREILLE 1797 (Coleoptera, Staphylinidae) 330. Beitrag zur Kenntnis der Steninen</dc:title>
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        <spm:hasContent> Wie die hohe Zahl der Synonyme ausweist, handelt es sich hier um eine Art, die dem Taxonomen Kopfzerbrechen bereitet hat und weiter bereiten wird. Solange der Aedoeagus der jeweiligen Taxa nicht untersucht worden war, musste man beim Vergleich ektoskelettaler Merkmale zu dem Resultat kommen, verschiedene Arten vor sich zu haben. Diese Auffassung wurde sicher auch dadurch bestärkt, dass man nicht gerade vermuten wollte, ähnliche Stücke aus Myanmar, von den Philippinenund von Java gehörten zu einer einzigen Art. Auch sind alle diese Taxa beschrieben worden, bevor man auf ihren eigenartigen Paraglossentyp geachtet hat. Und es war seinerzeit nicht bekannt, dass die jeweiligen Weibchen alle darin übereinstimmen, dass sie keine klar sklerotisierte Spermatheka besitzen (was ja im Vergleich zu den meisten gestaltlich ähnlichen Stenen die Ausnahme bildet). Die Paraglossen der Arten dieser Gruppe ähneln einem Springbrunnen (ein mittlerer langer Wasserstrahl wird von kurzen Wasserstrahlen umgeben), weshalb ich für diese Paraglossenform den Begriff " sileniform" wähle. Ob es sich möglicherweise nur um eine Modifikation des ovalen Paraglossentyps handelt, kann erst entschieden werden, wenn REM-Aufnahmen vorliegen. Dazu war aber bisher kein Material zu erhalten. Der Aedoeagus aller der genannten Taxa stimmt im Umriss des Medianlobus, in dessen Innnenbau und oft auch im Bau der Parameren so stark überein (Vergleich dazu: die nah verwandten Arten  S. emancipatus nov.sp.,  S. svenhedini nov.sp.,  S. nepalensisPUTHZ,  S. indecoratusPUTHZ,  S. calamei nov.sp.), dass er nicht als arttrennendes Argument herangezogen werden kann. Der Aedoeagus besitzt einen Medianlobus, der sich zur Spitze dreieckig (mehr oder weniger spitzwinklig) verengt, die Spitze ist schmal abgerundet [bei manchen Stücken kann die Apikalpartie aber auch vorne breiter gerundet und insgesamt schmäler erscheinen: Abb. 68, PUTHZ 1969b, Abb. 383]. Die Apikalpartie ist vorn ventral dicht beborstet. Im Innern befinden sich median verbundene Ausstülphaken (diese deutlich anders geformt als bei  S. emancipatus nov.sp.und  S. svenhedini nov.sp.), Ausstülpbänder und ein Komplex aus eng stehenden, länglichen, am Ende zugespitzten chikoreeähnlichen Schuppen sowie der länglich-zangenförmige Innensack ( Abb. 370-374). Die Parameren sind meist deutlich länger als der Medianlobus, an ihren Spitzen meist stark dreieckig verbreitert, apikal eingebuchtet; ihre konstante Beborstung besteht aus drei Gruppen von je zwei langen Borsten: zwei dorsal inserierenden seitlichen Borsten, zwei auf der ventralen Innenfläche und zwei am äusseren Spitzenende. Die ventrale Innenfläche weist einige Längsriefen auf ( Abb. 379). Wenn man nun versucht, ektoskelettale Unterschiede bei den männlichen Sexualcharakteren zu finden, die hier weiterhelfen könnten, wird man ebenfalls enttäuscht: die Metaventrite verschiedener Populationen zeigen zwar auffällige (nicht geschlechtsspezifische!) Unterschiede (gekielt/ungekielt (z. B. Abb. 336, 337, 346, 347/338, 339, 340- 345), dicht punktiert/wenig dicht punktiert (z.B. Abb. 339, 348/344, 346), genetzt/ungenetzt (z.B. Abb. 343, 344/345, 349) und auch die Ventralseite des Abdomens ist deutlich verschieden bei den Stücken unterschiedlicher Herkunft (Sternite 3-6 mit deutlichen Mitteleindrücken/Sternite 3-6 ohne Eindrücke: Abb. 352-357). Die Schenkel der Männchen sind – im Unterschied zu anderen nah verwandten Arten – deutlich gekeult, die Hinterschienen zur Spitze etwas nach innen gebogen und daselbst mit einem Präapikaldorn versehen ( Abb. 369). Prüft man dann ektoskelettale Merkmale der Oberseite, so stellt man die in den Beschreibungen der oben genannten Taxa beobachteten deutlichen Unterschiede fest: Grösse der Elytren, Grobheits- und Dichtegrad der Punktierung sind auffällig. Nebeneinander gestellt, wird man es nicht für möglich halten, dass Stücke aus Burmamit grossen, langen, etwa quadratischen Elytren und dichter Punktierung ( Abb. 316) konspezifisch sein sollen mit solchen von Java mit kurzen, trapezoiden, grob punktierten Elytren und mässig fein und weitläufig punktiertem Abdomen ( Abb. 325-327), aber auch Stücke von verschiedenen Orten Javas ( Abb. 326und Abb. 327). Auffällig sind die ektoskelettalen Unterschiede verschiedener Populationen von Java ( Abb. 319, 321, 323, 325-327). Ob hier nun eine polytypische Art vorliegt oder ob wir es mit klinalen Unterschieden zu tun haben, möchte ich zum gegenwärtigen Zeitpunkt nicht entscheiden. Ich verzeichne im Folgenden alle mir bekannten Funde.</spm:hasContent>
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        <spm:hasContent> Material:  Nepal: 1: Prov. Bagmati, Burlung Bhanjyang,  2600m,  5.IV.1981, Löbl&amp; Smetana( MHNG);  1: Prov. Kosi, Chichila s/Ahale,  2200m,  24.IV.1984, tamisage des bois pourrie avec champignons,  24.IV.1984, Löbl&amp; Smetana( MHNG);  1: Dolakha Distr., S slope of Khare Khola,  2100m,  2.VI.2000, Schawaller( SMNS);  2, 3: SW Dhaulagiri Himal, Dhara Khola Valley, 28 o30‘36‘‘N, 83 o18‘16‘‘E,  1900m, 21./  22.V.2012, Schmidt( NME, cP).  Indien: 5, 9: Meghalaya. East Khasi Hillsdistrict, Shillong Peak,  1800m, 25 o32‘30‘‘N, 91 o50‘55‘‘E,  21.X.2004, Cuccodoroet al. #9a( MHNG, cP);  1ibidem  1900m, 35 o32‘20‘‘N, 91 o51‘03‘‘E,  21.X.2004, idem ( cP);  1: Assam: North Cachar Hillsdistr., Mt. Borail, Borail Peak,  1700m, 25 o06.43’N, 93 o03.1’E,  19.X.12005, sifting bamboo litter in mountainous rain forest, Cuccodcoro&amp; Martella( MHNG).  Myanmar: - Lektotypus und 1- Paralektotypusvon  S. pilicornis: CarinCheba,  900-1100m, V-VII.1888, L. Fea( IRSNB, FMCh);  2- Paralektotypen: Carin Asciuii Ghécu,  1400-1500m, III-IV.1888, L. Fea( IRSNB, MCSNG);  2, 3: Shan prov., Namhsan,  1600m,  18.-20.II.1996, S. Kurbatov( MHNG, cP);  2: Kalaw,  19.III.1982, Rougemont( cR);  1, 1( Typenvon  S. meracus): Kambaiti,  2000m, Malaise( NHRM, FMCh).  Thailand: 1: C. Rai, Mae Lao,  III.1987, Rougemont( cR);  1: Ban Hua Mo,  12.III.1992, Rougemont( cR);  1: Doi Suthep,  6.III.1979, Rougemont( cR);  1: ibidem,  1100m, Mahidolwaterfall,,  20.II.1987, Schwendinger( MHNG);  1: ibidem  1450m,  4.XI.1985, tamisage de débris végétaux dans un ravin assez humide,  4.XI.1985, Burckhardt&amp; Löbl( MHNG);  1: Doi Angkhang Fang Distr., Ban Kum,  1800m,  28.VIII.1990, Schwendinger( MHNG);  2: Huay Nam Dang, Mae Taeng Distr.,  1400m,  17.XII.1990, Schwendinger( MHNG);  1: Taksin Maharat N. P.,  1000m,  9.II.1993, Schwendinger( cP);  1: Huay Nam Dang, Mae Tang Distr.,  1400m,  17.XII.1990, Schwendinger( NHMW);  1, 2: Chiang Mai,  10 kmW Wlang Pa Pao Ban HuySal  750m,  28.I.1988, Schwendinger( MHNG);  1: Doi Inthanon  1080m,  24.II.1987, Schwendinger( MHNG);  1: ibidem  1720m, ravin en forêt,  7.XI.1985, Burckhardt&amp; Löbl( MHNG);  2: Doi Saket  1130m,  4.XII.1987, Schwendinger( MHNG);  1: Nan Prov., Doi Phuka N. P.,  28.IV.-12.V.2002, PrĤdek&amp; OboĜil( cKecval).  Laos: 1: Vientiane Prov., Ban Van Eue,  30.VIII.1967, Native Collector( BPBM);  1: Sedone Prov., Paksong,  18.V.1965, P.D.Ashlock( BPBM).  China: Yunnan: 1: Huanxipo,  1750m, Tengchong X.,  14.X.1996, S. Nomura( cWatanabe);  3, 3: Dianshita Nabanhe,  1900m,  30.VII.2005, Li&amp; Li( SHNU);  4, 8: Bangganglahu, Nabanhe, 14./  15.I.2004, Li&amp; Tang( SHNU);  1: Bengganghani,  2000m,  29.IV.2009, Zi( SHNU);  3, 5: ibidem,  1800m, 22 o10.382’N, 100 o35.408’E,  18.XI.2008, Hu&amp; Tang( SHNU);  2: Bengganghani Nanmugahe,  1650m,  30.IV.2009, Zi( SHNU);  2: ibidem  1750m, 22 o08.383’N, 100 o34.126’E,  12.XI.2008, Hu&amp; Tang( SHNU);  1, 2: Nabanhe, Bengganhani Shanshenmiao,  1750m, 22 o08.450’N, 100 o35.289’E,  12.XI,.2008, Hu&amp; Tang( SHNU);  3: Benggangxinzhai, ca.  1750m,  3.V.2009, Jin&amp; Zi( SHNU, cP);  6, 1: ibidem  1850m,  4.V.2009, Zi( SHNU);  1: Xiaonuoyoushangzai,  1550m, 22.13.489 N, 100.35.342 E, Hu&amp; Tang( SHNU);  2: Nabanhe,  1900m,  30.VII.2005, Li&amp; Li( SHNU, cP);  2: Nabanheconv.,  15.I.2004, Li&amp; Tang( SHNU); : Baoshanpref., mountain range  25 kmS Tengchong,  1900m, 24 o48‘21‘‘N, 98 o32‘05‘‘E, cleft with devasted primary forest, litter &amp; mushrooms,  30.VIII.2009, Schülke( cSch);  1, 1: Baoshanpref., mountain range  14 kmE Tengchong,  1850m, 25 o00.28’N, 98 o38.07’E, secondary mixed forest,  1.VI.2007, Schülke, Pütz( cSch, cPütz);  1: Tengchong Co., Nakany,  4.V.2005, Huang( SHNU).  Sichuan: 1: Ganzi Tibet Aut. Pref., Kangding Co., Daxue Shan, brook valley  3 kmE Kangding,  2500-2600m, 30 o03’N, 102 o00’E,  24.VI.1999, Wrase( cSch).  Fujian: 1: Shaowu, Tachuland,  6.IV.1942, T.C.Maa( BPBM);  Taiwan: 1: Kaohsiung Hsien, road from Tengchihto Chuyunshan,  1400m,  25.IV.1990, A. Smetana(T 23) ( cS);  2: Nantou Hsien, near Lushan,  1200m, 27.,  29.VII.1977, Shibata( cShibata);  1: Lushan-Spa,  31.VII.1971, Shibata( cShibata);  1, 1: Chiai Hsien: Fenchihu,  8.VIII.1971, Shibata,  3.IX.1971, M. Nishikawa( cP, cShibata).  Malaysia: 1: Pahang,  15 miNKuala Lumpur,  29.IV.1977, berlese leaf litter &amp; rotten figs, L. E. Watrous( FMCH).  Indonesien: Sumatra: - HT von S. jacobsoni: Fort de Kock( NHML);  5, 3: Brastagi,  6.VI.1984, Rougemont( cR);  1: Propat,  3.VI.1984, Rougemont( cR);  1, 2: Brastagi, G. Sibayak,  1450-1900m,  19.-23.II.1991, Bocák&amp; Bocáková( NHMB, cP);  2, 2: 5 km W Brastagi, Tongkoh,  1450m, mixed pine forest with many epiphytes, sifting of vegetational debris,  3.XII.1989, Löbl&amp; al. ( MHNG, cP);  4: Jambi: Mt. Kerinci,  1750-1850m, montane Lithocarpus-Castanopsisforest, sifting of vegetational debris,  11.-12.XI.1989, idem (ibidem);  2, 5: Mt. Kerinci,  1750-1850m,  11.-12.XI.1989, Riedel( cRiedel, cP);  1: ibidem  3300m,  12.XI.1989, idem (ibidem);  1: Jambi, km 8 Sungaipenu to Tapan,  1200m,  9.XI.1989( cRiedel);  1: Jambi, 12 km W Sungaipenuh,  1250m,  4.III.1991, Bocák&amp; Bocáková( NHMB);  1: W Mt. Tujuh Lake,  1400m,  14.XI.1989, Riedel( cRiedel);  3, 3: Bukittinggi, Gn. Merapi,  1900-2100m,  10.X.1990, Riedel( ZMB, cP);  1: ibidem  1800m,  17.II.1991, Bocák&amp; Bocáková( NHMB);  1: Gn Merapi, 5 km E of Kotabaru,  1600m,  18.- 25.V.2001, Bolm( NHMB);  1: Batang Palupuh Nat. Res.,  850m,  15.II.1991, Bocák&amp; Bocáková( NNHMB).  Java: 2, 4: Pangrango,  3000m,  VIII.1921,  V.1922, Dammerman( Typenvon  S. pangrangoensis; Museum Leiden, DEI, cP);  1: Mt. Gede, 50 km SE Bogor,  2000-2200m, upper montane forest with a lot of moss, sifting vegetational debris,  5.XI.1989, Burckhardt, Löbl&amp; Agosti( MHNG, cP);  5, 1: ibidem  2600m, Ericaceaeforest, sifting vegetational debris,  5.XI.1989, idem (ibidem);  17, 9, ibidem  2800-2950m, low Ericaceaescrub, sifting vegetational debris,  5.XI.1989, idem (ibidem);  2: Mt. Gede,  1400-1500m,  24.-28.V.1997, Kurbatov( MHNG);  1: Cibodas Botanical Garden, 50 km E Bogor,  1400m, montane Lithocarpus-Castanopsisforest, sifting vegetational debris,  3.-6.XI.1989, idem (ibidem);  1, 4: Ijen National Park,  1800m, Sosong,  26.-27.II.1994, Bolm( SMNS, cP);  1, 1: 50 km S Surabaya,  800m, Tretes, Kekes BodoWF,  20.IX.1995, Koubek( NHMW);  1: G. Tangkoeban Prahoe,  4000-5000 feet, Preanger,  12.-18.I.1933( Typusvon  S. limbatus; FMCh);  1: G. Kawi,  IV.1919, Drescher( Museum Amsterdam);  1: G. Papandajan,  III.1915, Drescher( Mus. Amsterdam);  1: Pengalengan,  4000 feet, 1893, H. Fruhstorfer(HT von  S. makak; IRSCNB);  7, 4: Dieng,  23.VI.1984, Rougemont( cR, cP);  1: ibidem,  2100m,  12.XI.1978, Huber( MHNG);  4, 6: Gn Lawu 8 km W Sarangan,  1850m,  10.-11.V.2001, Bolm( NHMB, cP);  8, 5: SE Probolingo Gn. Argopuro, Bremi,  1600m,  18.VII.2003, Riedel( cRiedel, cP).  Bali: 1, 1: Penelokan,  25.XI.1978, Huber( MHNG);  1: L. Bunyan, 27,  VII.1981, Rougemont( cR);  2: ibidem,  1200m, forest,  8.-9.XI.1991, Löbl( MHNG);  17, 9: Danau Buyan  1300m,  19.-21.II.1994, Bolm( SMNS, cP);  6, 1: Kebun Raya, ca.  1600m,  4.II.1994, Pedersen( ZMK, cP);  1, 1: Kintamani, Gn Penulisan,  1485m,  9.XI.2007, Riedel( cRiedel);  1, Gn Abang,  1605m,  9.XI.2007, idem (ibidem);  1: Bedugul, Gn Catur,  1690m,  7.XI.2007, idem (ibidem),  3, 3: Gn Pohan,  1405m,  1.XI.2007, idem (ibidem, cP);  1, 1: Gn Catur, env. Bratansee,  1500m, Bergwald,  26.IX.1999, E. Heiss( cP);  1: Mt. Ahung, above Besahik Temple,  31.X.- 1.XI.1991, Löbl( cP);  Lombok: 6, 3: Sembalun Lawang, Mt. Rinjan,  1700m,  6.-8.II.1994, Bolm( SMNS, cP);  Sulawesi: 5, 5: Tanah Toraja, Pulu Pulu  1700m,  13.-16.VIII.1990, Riedel( ZMB, cP).  Philippinen: Luzon: 1: Trinidad(HT von  S. seriatipennis; FMCh);  1, 2: Subuagrn( Typenvon  S. plumbellus; FMCh, NHMW);  9, 12: Mountain Province, Sagada,  1500m, mousses, lavage litière, lavage humus,  19.XII.1979, Orousset( cOrousset, cP);  1: Ambasing  1440m,  16.XII.1979, idem (ibidem);  2, 2: Mt. Data,  2300m, mousses,  22.XII.1979, idem (ibidem);  1: Baguio,  1500m,  2.I.1980, litières, idem (ibidem);  1: Balbalason( cP).</spm:hasContent>
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