Coiloprosopa nitida Macquart, 1850: 449 Revision of the Psilota Meigen, 1822 flower flies (Diptera: Syrphidae) of Australia Young, Andrew D. Skevington, Jeffrey H. Steenis, Wouter Van Zootaxa 2020 2020-02-17 4737 1 1 127 4PJQ6 (Macquart, 1850) Macquart 1850 [151,537,1377,1404] Insecta Syrphidae Psilota Animalia Diptera 62 63 Arthropoda species nitida      Coiloprosopa nitida Macquart, 1850: 449. Typelocality: Tasmania, Australia   Diagnosis:Face black, densely white pollinose, with white pile. Frons and vertex black pilose. Thorax black. Hind femur with a low triangular apicoventral setulose carina. Hind tibia with basoventral spur. Calypter with dark brown border. Halter pale. Abdomen black. Lateral edges of tergite 3 with many pale pile. Male genitalia with inner lobe narrow, nearly parallel-sided, broadening slightly and regularly over entire length. Body length: 11.6–15.5 mm.   Description: MALE: Head:Head black, face yellowish-brown in some specimens, face pale pollinose. Face pale pilose. Scape and pedicel orange-brown, postpedicel brown. Postpedicel triangular, approximately 1.5 times as long as widest point. Eye densely brown pilose.  Thorax:Thorax black. Legs mostly orange, with apical 1/3 rdof femora, base and apex of hind tibia, and apical tarsomeres brown. Hind femur with a low, triangular apicoventral setulose carina. Hind tibia with a basoventral spur. Pleuron densely grey pollinose. Pile on anepisternum and anepimeron mixed black and white, pile on remainder of pleuron white. Pile on scutum and scutellum mixed black and whitish-yellow. Wing faintly brown tinted on apical half. Wing bare on anterior three-quarters of cell bm and one-quarter of cell cua, cell c bare on basal one-third. Alula completely microtrichose. Halter pale, sometimes with knob slightly pigmented. Calypter with border dark brown, upper calypter hairs brown, lower calypter hairs yellow.   FIGURE 27.  P. nitida(Macquart). A) male dorsal habitus, B) male anterolateral face, C) male lateral habitus. Scale bar =1 mm.   Abdomen:Abdomen black. Tergite 2 mostly yellow pilose, with some central black pile at posterior border. Tergite 3 yellow pilose laterally, black pilose medially. Tergite 4 with mixed black and yellow pile. Sternites black, with white pilosity.  Genitalia:Epandrium compact, slightly broader than long. Cercus approximately triangular. Outer lobe of surstylus inserted near top of inner lobe, curved smoothly downwards at midlength, and then swollen at apex. Inner lobe of surstylus narrow, nearly parallel-sided, broadening slightly and regularly over its entire length, with the apex rounded. Phallapodeme with large ventral projection. Gonostylus thick, not darkened.  FEMALE:Identical except for normal sexual dimorphism.  Specimens examined:  LECTOTYPE: AUSTRALIA:  Tasmania:“4/46” [circular label], “ Holotype,” “MNHN, Paris/ ED9344” and “ Coiloprosopa/ nitida/ ♀/ Macq. n. g., n. sp./ Tasm.” ( 1♀, OUMNH, photograph);  Othermate- rial:  Australian Capital Territory: Namadgi National Park,  28.3kmWSW Canberranear Lees Creek, -35.366111 148.835558,  900m,  18.ii–2.iii.2002, C.L. Lambkin, N.T. Starick, ANIC29035761( 1♀, ANIC);   New South Wales:  3 kmNortheast, Bilpin, Blue Mountains; -33.484094 150.552878; N.W. Rodd;  3.XII.1986; AustralianMuseumK410125 ( 1♂, AM);  Coocumbac Island Nature Reserve, Taree, -31.920678 152.454450, floodplain rainfor- est landing near small pool of water on fallen log,  18.i.1994, G. & T. Williams, AustralianMuseumK410624 ( 1♀, AM);  Bawley Point, -35.500000 150.400000,  9–13.v.2001, D.C.F. Rentz, S.&O. Schmidt, ANIC29035760( 1♂, ANIC);  Kangaroo Park, Manly, -33.803722 151.294769, on flowering Cotoneaster,  23.xi.1986, D.S. Horning, Jr., ANIC29035752( 1♀, ANIC);   3 kmNorth of Lansdowne, -31.758753 152.534258, on  Cuttsia viburnea,  16.xii.1991, G. Williams, AustralianMuseumK408020, AustralianMuseumK410243 ( 1♂, 1♀ AM);    4 m.N. of Bateman’s Bay, - 35.658458 150.177744,  14.x.1952, S.J. Paramonov, CNC_ Diptera253693; USNM_ENT_00249828, ANIC29034901( 2♂♂, 1♀, ANIC USNM);   20.x.1953, S.J. Paramonov, ANIC29034890( 1♂, ANIC);   29.ix.1959, S.J. Paramonov, ANIC29034900( 1♂, ANIC);   22.x.1952, S.J. Paramonov, CNC_ Diptera253694 ( 1♀, ANIC);   20.x.1952, S.J. Para- monov, CNC_ Diptera255080 ( 1♂, ANIC);   22.x.1952, S.J. Paramonov, CNC_ Diptera255079 ( 1♂, ANIC);  Bald Rock N.P., -28.860783 152.055686, on  Leptospermum minutifoliumblossom,  6–7.xi.1984, D.K. Yeates, UQIC_ Reg94642 ( 1♂, QM);  Bellmore Falls, -34.633958 150.555914, 1948, S.J. Paramonov, [Date is ambiguous reads ”12.11.48”], ANIC29034897( 1♂, ANIC);  Bilpin Blue Mountains, -33.500261 150.532989, 1978, N.W. Rodd, Aus- tralianMuseumK408019 ( 1♂, AM);  Burralow Swamp, -33.548342 150.592047,  24.ii.1978, N.W. Rodd, Austra- lianMuseumK410170 ( 1♂, AM);  Durras Bay, -35.645003 150.291439,  12.xi.1953, F.M. Hull, UQIC_ Reg94722; CNC_ Diptera254978–254979; CNC_ Diptera254974–254975 ( 3♂, 2♀, QM; CNC);  Mannus, near Tumbarumba, -35.808336 147.935786,  1.xii.1950, T.G. Campbell, ANIC29034891( 1♀, ANIC);  Mooney Mooney Cknr. Bosford, -33.437894 151.246467,  25.xi.1975, G. Daniels, G. Daniels Collection, GDCBReg_19721 ( 1♀, GDCB);  Mount Tomah, Blue Mountains, -33.549614 150.416033,  28.xi.1977, 26–27.x.1978, 25.i.1979, 17.xii.1982, 13.ii.1984, 20.i.1985, 21.i.1985, 28–29.xii.1985, 7.ii.1987, 3.iv.1987, N.W. Rodd, AustralianMuseumK408011; AustralianMuseumK408014, AustralianMuseumK410123, AustralianMuseumK410129, AustralianMuseumK410136–410138, AustralianMuseumK410142, AustralianMuseumK410161, AustralianMuseumK410171–410172, AustralianMuseumK410276, AustralianMuseumK410166, AustralianMuseumK410280, AustralianMuseumK410104 ( 5♂♂, 10♀♀, AM);  Tooloom, -28.592394 152.453372,  26.i.1926, UQIC_ Reg94665 ( 1♀, QM);   3 kmNorth of Lansdowne, -31.758753 152.534258,  Waterhousia floribunda,  29.xi.1991, G. Williams, AustralianMuseumK409934 ( 1♀, AM);  Willoughbynear Sydney, -33.926375 151.125936,  16.ix.2001, D.K. McAlpine, AustralianMuseumK410128 ( 1♀, AM);   Queensland: Blue Mountain,  0.6 kmSoutheast, -21.6 148.96,  800 to 1058m,  3–4.x.1999, G. Monteith, C. Burwell, Cook, S. Evans, QM_REG._NO._T223197–223199 ( 3♀, QM);  Clarke Range, North Road-Pine Road, Cathu State Forest, site 2, -20.805278 148.483333,  750m,  24.i.1996, G. & A. Daniels, AustralianMuseumK410376 ( 1♀, AM);  6kmW by N of Paluma, -19.000000 146.090000,  25.ix.1980, D.H. Colless, ANIC29034484( 1♀, ANIC); Blue Mountain,  0.6 kmSoutheast, -21.6 148.96,  800 to 1058m,  3–4.x.1999, G. Monteith, C. Burwell, Cook, S. Evans, QM_REG._NO._T223200–T223207 ( 8♀♀, QM);  Gatton, -27.563583 152.279983,  22.iii.1960, J. Mar- tin, UQIC_ Reg94595 ( 1♂, QM);  Scrub Road,  Brisbane ForestPark, -27.416667 152.833333, [Latitude] 27°25’S[Longitude] 152°50’E  3–10.x.1997, S. Winterton, N. Power, D. White, Malaise trap, UQIC_ Reg94798 ( 1♂, QM);  Gambubal State Forest, via Emu Vale, -28.235000 152.376111,  1090m,  17.iii.2009, G. Daniels, AustralianMuseumK408007 ( 1♂, AM);  Eungella National Park, Chelmer’s Road, -21.132822 148.492683,  19.xii.2014, J.H., A.M. and A.W. Skevington; CNC451698( 1♀, CNC);   Tasmania: Geeveston, -43.166500 146.933397,  28.xii.1914, G.H. Hardy, ANIC29034896( 1♀, ANIC);  Hobart, -42.882139 147.327194,  i.1924, G.H. Hardy, ANIC29034898( 1♂, ANIC);  Huon-Pictonjunction, -43.139689 146.714758,  17.ii.1967, E.F. Riek, ANIC29034893( 1♀, ANIC);  South Bruni [Bruny] Island, -43.488236 147.295425,  i.1916, Clarke, ANIC29034899( 1♂, ANIC);  Warra Long TermEco- logical Research Site170, -43.100000 146.683333,  14.i–12.ii.1998, Malaise trap, UQIC_ Reg94611 ( 1♀, QM);  A. Simson, 3266, SAMADatabaseNo.29-003356 ( 1♂, SAMA);  Victoria: Gunyah, -38.525603 146.255119,  iii.1955, Bornemissza, ANIC29034892( 1♀, ANIC);  Springvale, -37.947175 145.153067, L.B. Thorn, ANIC29034489( 1♀, ANIC);  Tarra Bulga National Park, Tarra Bulga Visitors Centre, -38.429578 146.566172,  27.i.2006, W. van Steenis, Jeff_ Skevington_ Specimen43828 ( 1♀, QM);  Barrington Area, Mt. William, -40.934189 148.235269,  152m,  6.iii.1951, B. McMillan, UQIC_ Reg94667 ( 1♀, GDCB); F.M. Hull, CNC_ Diptera254976 ( 1♂, CNC).   Discussion: Lectotypefemale in MNHN and labeled “4/46” [circular label], “ Holotype,” “MNHN, Paris/ ED9344” and “ Coiloprosopa/ nitida/ ♀/ Macq. n. g., n. sp./ Tasm.” [in Macquart’s handwriting]. This specimen is herewith designated lectotypeand has so been labeled.   Psilota nitidadisplays more divergence on COI than other  Psilotaand may represent a species complex. However, no morphological characters could be found to distinguish specimens that were more similar on COI than others, and it was therefore treated as a single species. This species has been collected on  Cuttsia viburnea(Rousseaceae)and on  Leptospermum minutifolium(Myrtaceae)blossoms. 2571544699 OUMNH Australia 64 65 1 1 Tasmania lectotype 2571544753 2002-02-18 ANIC C.L. Lambkin & N.T. Starick Australia -35.36611 148.83556 64 65 ANIC29035761 1 1 Australian Capital Territory 2571544734 1986-12-03 AM N.W. Rodd Australia -33.484093 150.55287 64 65 1 1 New South Wales 2571544702 1994-01-18 AM Taree & G. & T. Williams Australia -31.920677 152.45445 64 65 1 1 New South Wales 2571544750 2001-05-09 ANIC Bawley Point & D.C.F. Rentz & S.&O. Schmidt Australia -35.5 150.4 64 65 ANIC29035760 1 1 New South Wales 2571544706 1986-11-23 ANIC Kangaroo Park & Manly & Cotoneaster & D.S. Horning Australia -33.803722 151.29477 64 65 ANIC29035752 1 1 New South Wales 2571544693 1991-12-16 AM G. Williams Australia -31.758753 152.53426 64 65 2 1 1 New South Wales 2571544697 1952-10-14 CNC S.J. Paramonov Australia -35.65846 150.17775 64 65 ANIC29034901 3 1 2 New South Wales 2571544728 [464,1107,836,863] 1953-10-20 ANIC S.J. Paramonov Australia 64 65 ANIC29034890 1 1 New South Wales 2571544700 1959-09-29 ANIC S.J. Paramonov Australia 64 65 ANIC29034900 1 1 New South Wales 2571544690 [490,1191,872,899] 1952-10-22 CNC S.J. Paramonov Australia 64 65 1 1 New South Wales 2571544691 1952-10-20 CNC Australia 64 65 1 1 New South Wales 2571544737 [652,1368,908,935] 1952-10-22 CNC S.J. Paramonov Australia 64 65 1 1 New South Wales 2571544698 1984-11-06 QM Bald Rock N.P. & D.K. Yeates & Reg Australia -28.860783 152.05568 64 65 1 1 New South Wales 2571544715 ANIC S.J. Paramonov Australia -34.633957 150.55591 64 65 ANIC29034897 1 1 New South Wales 2571544695 AM N.W. Rodd Australia -33.50026 150.53299 64 65 1 1 New South Wales 2571544730 1978-02-24 AM Burralow Swamp & N.W. Rodd Australia -33.548344 150.59204 64 65 1 1 New South Wales 2571544745 1953-11-12 CNC F.M. Hull & Reg Australia -35.645004 150.29144 64 65 5 2 3 New South Wales 2571544689 1950-12-01 ANIC Mannus & Tumbarumba & T.G. Campbell Australia -35.808334 147.93579 64 65 ANIC29034891 1 1 New South Wales 2571544736 1975-11-25 GDCB Mooney Mooney Ck & Bosford & G. Daniels & G. Daniels Collection Australia -33.437893 151.24646 64 65 1 1 New South Wales 2571544731 1978-10-26 AM N.W. Rodd Australia -33.549614 150.41603 64 65 15 10 5 New South Wales 2571544748 [300,1129,1412,1439] 1926-01-26 QM Tooloom & Reg Australia -28.592394 152.45337 64 65 1 1 New South Wales 2571544743 1991-11-29 AM G. Williams Australia -31.758753 152.53426 64 65 1 1 New South Wales 2571544741 2001-09-16 AM Willoughby & Sydney & D.K. McAlpine Australia -33.926376 151.12593 64 65 1 1 New South Wales 2571544747 1999-10-03 QM G. Monteith & C. Burwell & Cook, S Australia 64 65 3 3 Queensland 2571544732 1996-01-24 AM Clarke Range & G. & A. Daniels Australia -20.805279 148.48334 64 65 1 1 Queensland 2571544692 1999-10-03 ANIC D.H. Colless & G. Monteith & C. Burwell & Cook, S Australia -19.0 146.09 64 65 ANIC29034484 1 1 Queensland 2571544717 1960-03-22 QM Gatton & Reg Australia -27.563583 152.27998 64 65 1 1 Queensland 2571544725 1997-10-03 QM Scrub Road & Brisbane Forest Park & S. Winterton & N. Power & D. White & Reg Australia -27.416666 152.83333 64 65 1 1 Queensland 2571544739 2009-03-17 AM Emu Vale & G. Daniels Australia -28.235 152.37611 64 65 1 1 Queensland 2571544733 2014-12-19 CNC Chelmer's Road & J.H., A.M. and A.W. Skevington Australia -21.132822 148.49268 64 65 CNC451698 1 1 Queensland 2571544696 [694,1436,1880,1907] 1914-12-28 ANIC Geeveston & G.H. Australia -43.1665 146.9334 64 65 Tasmania 2571544694 1924-01 ANIC Hobart & G.H. Hardy Australia -42.882137 147.3272 64 65 ANIC29034898 1 1 2571544742 [244,1359,1952,1979] 1967-02-17 ANIC Huon-Picton & E.F. Riek Australia -43.13969 146.71475 64 65 ANIC29034893 1 1 2571544740 [1371,1436,1953,1979] 1916-01 ANIC Australia -43.488235 147.29543 64 65 Tasmania 2571544729 1998-01-14 QM Warra Long Term & Research Site & Reg Australia 65 66 -43.1 146.68333 64 65 1 1 Tasmania 2571544746 1955-03 SAMA A. Simson & Gunyah & Bornemissza Australia -38.525604 146.25511 65 66 ANIC29034892 2 1 1 Victoria 2571544754 ANIC Springvale & L.B. Thorn Australia -37.947174 145.15306 65 66 ANIC29034489 1 1 Victoria 2571544735 2006-01-27 QM Tarra Bulga Visitors Centre & W. van Steenis & Jeff & Skevington & Specimen Australia -38.429577 146.56618 65 66 1 1 Victoria 2571544755 1951-03-06 GDCB B. McMillan & Reg & F.M. Hull Australia -40.93419 148.23528 65 66 2 1 1 Victoria