Psilota coerulea Macquart, 1846: 268 Merodon muscaeformis Walker, 1852: 244 Revision of the Psilota Meigen, 1822 flower flies (Diptera: Syrphidae) of Australia Young, Andrew D. Skevington, Jeffrey H. Steenis, Wouter Van Zootaxa 2020 2020-02-17 4737 1 1 127 4PJPN Macquart, 1846 Macquart. A 1846 [151,552,1336,1363] Insecta Syrphidae Psilota Animalia Diptera 40 41 Arthropoda species coerulea      Psilota coerulea Macquart, 1846: 268. Typelocality: Tasmania, Australia     Merodon muscaeformis Walker, 1852: 244. Typelocality: Tasmania, Australia   Diagnosis:Metallic blue species with frons pale pilose, pleuron shiny and black pilose. Hind femur with a distinct rounded apicoventral setulose carina. Hind tibia simple. Calypter entirely brown. Tergite 4 white pilose. Body Length: 9.4–14.0 mm.   Description: MALE: Head:Head black, face densely pale pollinose. Face and frons yellow pilose, vertex black pilose. Antenna usually entirely orange, rarely dull brown dorsally on postpedicel. Eye densely yellow pilose.   Thorax:Thorax dark, metallic blue. Legs mostly dark brown, with apices of femora, and entire fore and mid tibia sometimes pale brown/orange. Hind femur with a distinct rounded apicoventral setulose carina. Hind tibia simple, with no basoventral spur. Pleuron shining, pollinosity free. Pile on pleuron, scutum, and scutellum entirely black. Wing colourless, with cells c, r, bm, cua, and the anal lobe mostly bare. Cells r 4+5, r 2+3, and dm with bare areas basally. Alula completely microtrichose. Halter dark brown. Calypter pale brown to dark brown, always with a dark brown border.  Abdomen:Abdomen metallic bluish-green. Tergites 2 and 3 with white pile on basal half and black pile on apical half, tergite 4 entirely white pilose. Sternites shining, white pilose.  Genitalia:Epandrium compact, about as long as tall. Cercus approximately triangular. Outer lobe of surstylus inserted just above mid-height of inner lobe, curved downwards sharply at one-third its length. Inner lobe of surstylus strongly projected ventrally at base forming a triangular plate, slightly contricted at midlength, then broadening slightly towards a rounded apex. Phallapodeme with a small ventral projection. Gonostylus thick, darkened.   FIGURE 16.  P. coeruleaMacquart. A) male dorsal habitus, B) male anterolateral face, C) male lateral habitus. Scale bar =1 mm.  FEMALE:Identical except for normal sexual dimorphism.  Specimens examined:  LECTOTYPE: AUSTRALIA(  Psilota coerulea):  Tasmania: Bearslabels reading “ HOLOTYPE”, “MNHN, Paris ED9367”, “13 44”, “ Psilota coerulea  n. sp.Macq.”. ( 1♀, MNHN, photograph). Bears label “Type” [green circular label], “Muscae,” “VDL,” and “ Merodon/ muscaeformis / Wlk” [in E. E. Austen’shandwriting], J. Skevington Specimen_45634, ( 1♀BMNH, photograph). Other material:   Australian Capital Territory: Black Mountain, -35.273675 149.097583,  25.ii.1957, I.F.B. Common, light trap, ANIC29034817( 1♀, ANIC);  Mount Ainslie, -35.266667 149.166667,  28.xi.1996, C.J. Burwell, QM_ REG._NO._T223174 ( 1♂, QM);   New South Wales:  24 kmW South Grafton, -29.616667 152.733333,  19.ix.1981;  8.ix.1983;  3.x.1985;  26.ix.1986;  29.ix.1986;  29.ix.1989, R. Eastwood, GDCBReg_19565; GDCBReg_19667-19668; GDCBReg_19694-19695; GDCBReg_19697; GDCBReg_19774; UQIC_ Reg94590; UQIC_ Reg94707 ( 9♂♂, GDCB, QM);  Congo, - 35.959786 150.135839,  3.x.1981, M.S. Upton, ANIC29034820( 1♀, ANIC);  Gosford, -33.426686 151.341658,  x.1924, ANIC29034815( 1♀, ANIC);  Jenolan Caves, -33.819039 150.022917,  609m, ANIC29034458( 1♀, ANIC);  Katoomba, -33.713133 150.313267,  18.ii.1954, AustralianMuseumK410150 ( 1♀, AM);  Mount Tomah, Blue Mountains, -33.549614 150.416033,  29.ix.1977;  18.x.1980, N.W. Rodd, AustralianMuseumK407026; AustralianMuseumK410291 ( 1♂, 1♀, AM);  Narrabeen, -33.719683 151.297125,  16.ix.1934, D. F.W., ANIC29034814( 1♂, ANIC);  Sydney, -33.926375 151.125936, 1925, Dateis ambiguous reads ”2.1.25”, ANIC29034818( 1♀, ANIC);   Queensland: Carnarvon N.P., Summit Of“Fly Hill”, near West Branch Camp, -24.974167 147.992778,  900m,  12.x.2002, J. Skevington; J.E. O’Hara, CNC_ Diptera254989-254990; CNC_  Diptera254994-254997 ( 5♂♂, CNC); Kenniff’s Lookout, Mt Moffattsect., Carnarvon Nat Pk, -25.165167 147.840419,  25.ix.1986, D.K. Yeates, UQIC_ Reg94661 ( 1♂, QM);  Kuranda, -16.816878 145.633497, ANIC29034453( 1♀, ANIC);  Mount MoffatNa- tional Park, Mount Ruggedsummit, -24.900000 148.000000,  1130m,  24.xi.1995, C.J. Burwell, QM_ REG._NO._ T223173 ( 1♂, QM); Mount Moffatt, Mt Moffattsect., Carnarvon Nat. Pk., -25.165167 147.840419,  26.ix.1986, D.K. Yeates, UQIC_ Reg94660 ( 1♂, QM);  Mount Tamborine, -27.923058 153.171667, 1923, W.H. Davidson, QM_ REG._NO._T223175 ( 1♀, QM); Near Rosedale, -24.631128 151.913664,  27.x.1974, H. Frauca, ANIC29034916( 1♂, ANIC);   Tasmania: Mole Creek, -41.558628 146.404517,  4.ii.1910, ANIC29034457( 1♀, ANIC);  MountWil- liam National Park, Summit of Mount William, -40.909722 148.186389,  23.xii.1998, J. & A. Skevington, UQIC_ Reg94553-94554 ( 2♂♂, QM);  Swansea, -42.134133 148.058614, Lea, ANIC29034808; ANIC29034816( 2♀♀, ANIC);   Victoria: Beaconfield, -38.048778 145.371733, A.M. Wade, ANIC29034821( 1♀, ANIC);  Broadwater National Park, -29.050325 153.427539,  6.ix.1981, D. Yeates, U.Q.I.C. Loan#1036, UQIC_ Reg94582 ( 1♂, QM).   Discussion: Lectotypefemale in MNHN and labeled: “ HOLOTYPE” [red label], “13 /44” [circular label], “MNHN, Paris/ ED9367,” and “ Psilota/ coerulea / n. sp.Macq.” Only a fragment remains in a microvial, along with a badly dermestid eaten remnant on the pin. The original species description of  Psilota coeruleadoes not list a holotypeor syntypeseries but appears to be based on a female specimen described as “De le Tasmanie. Muséum.”. A specimen in the Paris Muséum National d’Histoire Naturelle bears labels reading “ HOLOTYPE” [red label], “13 /44” [circular label], “MNHN, Paris/ ED9367,” and “ Psilota/ coerulea / n. sp.Macq.”, and appears to be the specimen that Macquart originally described. This specimen is herewith designated lectotypeand has so been labeled. The specimen has been badly eaten by dermestids, with only parts of the thorax and a fragment in a genitalia vial remaining. Likewise, the original description of  Merodon muscaeformisis based on a female in BMNH and labeled “Type” [green circular label], “Muscae,” “VDL [Van Dieman’s Land]” and “ Merodon/ muscaeformis / Wlk” [in E. E. Austen’s handwriting]. This specimen is herewith designated lectotypeand has been so labeled. 2565887924 MNHN E. E. Austen's & J. Skevington Specimen Australia Bears 42 43 2 2 Tasmania lectotype 2565883894 1957-02-25 light trap ANIC I. F. B. Common -35.273674 Black Mountain 1 149.09758 42 43 ANIC29034817 1 1 Australian Capital Territory 2565887963 [246,1052,368,395] 1996-11-28 QM C. J. Burwell Australia -35.266666 Mount Ainslie 1 149.16667 42 43 1 Tasmania 2571544657 [1066,1432,369,395] QM Australia 42 43 1 1 2571544718 1985-10-03 GDCB Eastwood & Reg & Reg Australia -29.616667 152.73334 42 43 9 9 New South Wales 2571544661 1981-10-03 ANIC M.S. Upton Australia -35.959785 150.13583 42 43 ANIC29034820 1 1 New South Wales 2565887938 1924-10 ANIC Australia -33.426685 Gosford 1 151.34166 42 43 ANIC29034815 1 1 New South Wales 2565887949 ANIC Australia 609 -33.81904 Jenolan Caves 1 150.02292 42 43 ANIC29034458 1 1 New South Wales 2571544704 [248,1251,584,611] 1954-02-18 AM Australia -33.713135 150.31326 42 43 1 1 2571544668 1980-10-18 AM N.W. Rodd Australia -33.549614 150.41603 42 43 2 1 1 2571544652 1934-09-16 ANIC F.W. Australia -33.719685 151.29712 42 43 ANIC29034814 1 1 2565888001 ANIC Date Australia -33.926376 Sydney 1 151.12593 42 43 ANIC29034818 1 1 New South Wales 2565888032 2002-10-12 CNC Carnarvon N. P. & Summit Of & J. Skevington & O'Hara Australia 900 -24.974167 West Branch Camp 1 147.99278 42 43 1 Queensland 2571544654 1986-09-25 CNC D.K. Yeates & Reg Australia -25.165167 147.84042 42 43 5 5 2571544674 [488,1213,836,863] ANIC Australia -16.816877 145.6335 42 43 ANIC29034453 1 1 2565888010 1995-11-24 QM C. J. Burwell Australia Mount Moffat 1130 -24.9 Mount Rugged 1 148.0 Park 42 43 1 Queensland 2571544651 1986-09-26 QM D.K. Yeates & Reg Australia -25.165167 147.84042 42 43 1 1 2571544667 [628,1422,944,971] QM W.H. Davidson Australia -27.923058 153.17166 42 43 2565888035 1974-10-27 QM, ANIC H. Frauca Australia -24.631128 Near Rosedale 1 151.91367 42 43 ANIC29034916 2 1 1 Queensland 2565887903 [300,1294,1016,1043] 1910-02-04 ANIC Australia -41.558628 Mole Creek 1 146.40451 42 43 ANIC29034457 1 1 Tasmania 2571544659 1998-12-23 QM J. & A. Skevington & Reg Australia -40.90972 148.18639 42 43 2 2 2571544656 ANIC Lea Australia -42.134132 148.05861 42 43 ANIC29034808, ANIC29034816 2 2 2565887905 [248,1290,1124,1151] ANIC A. M. Wade Australia -38.04878 Beaconfield 1 145.37173 42 43 ANIC29034821 1 1 Victoria 2565887958 1981-09-06 QM D. Yeates & U. Q. I. C. Loan & Reg Australia -29.050325 Broadwater National Park 1 153.42754 42 43 1 1 Victoria