A synoptic revision of the genus Lepisanthes Blume (Sapindaceae) in Madagascar
Buerki, Sven
Callmander, Martin W.
Lowry Ii, Porter P.
Phillipson, Peter B.
Adansonia
2009
3
2009-12-31
31
2
301
309
2. Lepisanthes perrieri (Choux) Buerki, Callm. & Lowry, comb. et stat. nov. Manongarivea perrieri Choux, Comptes rendus hebdomadaires des Séances de l’Académie des Sciences(Paris) 182: 713. (1926); Mémoires de l’Académiemalgache4: 36 (1927). — Aphania senegalensisf. perrieri(Choux) Capuron, Mémoires du Muséum d’Histoirenaturelle(Paris), sér. B, 19: 82. — Type: Madagascar, Prov. Mahajunga, Manongarivo(Ambongo), [ 16°16’S, 45°22’E], X.1909, fl., y.fr., Perrier de la Bâthie1812(lecto- [here designated], P 00624104!; isolecto-, K 000426188!, P 00624103!, TAN!]. NOTES. — As indicated above, Capuron (1969)used one of the syntypesof Manongarivea perrieri( Perrier de la Bâthie 1744)as the type of his name Aphania senegalensissubsp. chrysotricha, implicitly leaving only Perrier de la Bâthie 1812associated with Choux’s name. Here we formalize this choice, designating one of the two sheets in the Paris Herbarium as the lectotypeof M. perrieri. ADDITIONAL MATERIAL EXAMINED. — Madagascar. Prov. Antsiranana, AnkaranaAP, 12°54’32”S, 49°06’35”E, 172 m, 7.XII.2006, y.fr., Andriamihajarivoet al. 1047( MO, P, TAN).— AnkaranaAP, 12°49’00”S, 49°01’17”E, 150 m, 12-20.X.1993, fl., Andrianantoanina368( MA, MO, P, TAN). — Ankarana PA, 2 kmavant le village d’Ambondromifehy, 12°52’36”S, 49°13’13”E, 276 m, 21.XI.1995, fr., Andrianantoanina& Bezara892( G, MO, P, TAN). — Montagne d’AmbreAP, c. 8 kmà l’est du village de Bobakilandy, 12°37’37”S, 49°6’26”E, 533 m, 5.XII.1995, fr., Andrianantoanina& Bezara896( G, MO, P, TAN). — Ankarana AP, 12°55’23”S, 49°05’11”E, 150 m, 20.X.2001, fl., Bardot-Vaucoulon& Andrianantoanina820( P[2 sheets], TAN). — Ankarana AP, 100 m, 12°57’35”S, 49°07’15”E, 30.X.2001, fr., Bardot-Vaucoulon& Andrianantoanina871( K, MO, P[2 sheets], TAN). — Ankarana AP, 12°58’52”S, 49°06’26”E, 125 m, 12.III.2001, fl., Bardot-Vaucoulon& Andrianantoanina901( K, MO, P[2 sheets], TAN). — Ankarana AP, 12°53’38”S, 49°06’47”E, 130 m, 9.XII.1997, fr., Bardot-Vaucoulon& Toly1042( K, MO, P[2 sheets], TAN). — Abattoir, 22.XII.1916, fl., Decary77( P). — AnkaranaAP, 12°54’40”S, 49°06’31”E, 180 m, 19.XI.1996, fl., y.fr., Labat2772( K, L, MO, P, TAN, WAG). — Daraina, forêt de Binara, 13°15’S, 49°37’E, 300 m, 5.XI.2001, fl., Gautier& Ravelonarivo4045( G, TEF, DARAINA, P, MO, K). — Daraina, forêt de Bekaraoka, 13°06’S, 49°42’E, 450 m, 22.XI.2006, fl., Gautier& Chatelain4912( G, TEF, DARAINA, P, MO, K). — AnkaranaAP, 100-200 m, 12°51’S, 49°04’E, 22- 26.XI. 1992, fl., Malcomberet al. 1821( K[2 sheets], G, MO, P, TAN). — Ankarana AP, 100-200 m, 12°51’S, 49°04’E, 22-26.XI. 1992, y.fr., Malcomberet al. 1823( K, MO, P). — Malcomberet al. 1882( MO). — AnkaranaAP, 100-200 m, 12°51’S, 49°04’E, 22-26.XI. 1992, fl., Malcomberet al.1886( MO, P, TAN, WAG).— AnkaranaAP, 100-200 m, 12°51’S, 49°04’E, 22-26.XI.1992, fr., Malcomberet al. 1887( MA, MO, P, TAN). — AnkaranaAP, 12°52’S, 49°14’E, 320 m, 24.XI.1993, fr., McPherson14525( MO, P, TAN, TEF). — Nosy Mitsiou, [ 12°54’S, 48°36’E], X.1936, fl., Perrier de la Bâthie18775( G, MO, P). — Ampondrabe, 12°58’18”S, 49°41’57”E, 200 m, 5.XI.2005, fl., Rakotonandrasanaet al. 953( CNARP, MO, P, TAN). — Befarafara, 13°05’26”S, 049°34’32”E, 100 m, 15.XI.2005, fl., Rakotonandrasanaet al. 1002( CNARP, MO, P, TAN). — Daraina, forêt de Bobankora, 13°11’4”S, 49°45’30”E, 200 m, 11.XI.2005, fl., Rakotondrajaonaet al. 378( CNARP, MO, P, TAN). — Daraina, forêt d’Antsahabe, 13°13’S, 49°33’E, 340 m, fr., 28. I.2006, Ranarisonet al. 1134( G, DARAINA, MO, K, P, TEF). — Ananjaka, 13°06’00”S, 049°10’07”E, 155 m, 6. XI. 2004, fl., Randrianaivoet al. 1093( CNARP, MO, P, TAN). — Antsisikala, forêt d’Analabe, 12°11’49”S, 049°11’51”E, 184 m, 11.XII.2005, fl., Randrianaivoet al. 1336( CNARP, MO, P, TAN). — AnkaranaAP, 12°58’57”S, 049°09’50”E, 127 m, 10.XII.2007, fr., Randrianasolo S. et al. 630( CNARP, MO, P, TAN). — Ankijomantsina, 13°07’40”S, 49°27’53”E, 100 m, 31.X.2005, fl., Ratovosonet al. 1044( CNARP, MO, P, TAN). — Antsisikala, 12°50’02”E, 049°12’38”S, 100 m, 3.XII.2007, fr., Ratovosonet al. 1416( CNARP, MO, P, TAN). — AnkaranaAP, 160-200 m, 12°54’S, 49°08’E, 10.XI.2006, fl., Z. S. Rogerset al. 1160( MO, P, TAN, G, K, US). — 23 km Sof Anivorano, 40 km Nof Ambilobe, 12°53’S, 49°12’E, 200 m, 24.X.1992, fl., Schatzet al. 2409( MO, P, TAN). — Massif de la Montagne d’Ambre, [ 12°30’S, 49°05’E], c. 300 m., 19.X.1958, fl., y.fr., Service Forestier11343( G, K, MO, P, TEF). — Windsor Castle, [ 12°14’S, 49°10’30’’E], c. 100 m., 4.XI.1955, fr., Service Forestier15168( TEF). — Ambondromifehy, [ 12°53’30’’S, 49°12’30’’E], c. 300 m, 24.X.1959, fl., y.fr., Service Forestier15189( TEF). — Près d’Ambohimagodra, [ 13°01’30”S, 49°08’E], c. 100 m., 12.XI.1958, fr., Service Forestier18951( TEF). — Ankarana, forêt de Marovato, massif de l’Ankerana, [ 12°48’30’’S, 49°09’E], 200 m, 14.XI.1958, fr., Service Forestier20008( P, TEF). — Sdu Pic Raynaud, [ 12°28’30”S, 49°26’30”E], 100-400 m, 9.XI.1961, fl., Service Forestier20364( P, TEF). — Sdu Pic Raynaud, [ 12°28’30”S, 49°26’30”E], 100-400 m, 9.XI.1961, fl., Service Forestier20365( TEF). — Nord de Vohémar, Mafokovo, [ 13°18’S, 49°51’E], [ 50-450 m], 18.XII.1966, y.fr., Service Forestier27370( P, TEF). Prov. Mahajanga, Manongarivo, Ambongo, [ 16°16’S 45°22’E], 2.X.1910, fl., y.fr., Perrier de la Bâthie1812( P). — Soalala, Ambongo, [ 16°06’S, 45°19’E], XII.1926, fl., y.fr., Perrier de la Bâthie17842( P[2 sheets]). — Bongolava, Borizini, 15°38’58”S, 47°35’03”E, 217 m, 16.XI.2004, fl., Ramananjanaharyet al. 133( MO, P, TEF). — Bongolava, Borizini, 15°34’00”S, 47°29’55”E, 115 m, 27.XI.2004, fr., Ramananjanaharyet al. 166( MO, P, TEF). — Prov. Mahajanga, AntsianitiaSTF, 20 m, 15°35’18”S, 46°26’01”E, 20. V.2001, fr., R. Randrianaivoet al. 645( CNARP, MO, P, TAN). — Ambondro, Analalava, [ 15°01’S, 47°16’E], 1.X.1958, fr., Service Forestier10778( P, TEF). — Massifforestier de Bora, [ 14°54’S, 48°13’E], 100-411 m, 29.XI.1961, fr., Service Forestier18496( P, TEF). — Forêt d’Ambondro-Ampasy, [ 15°01’S, 47°16’E], 50 m, 29.X-3.XI.1958, fr., Service Forestier18811( P, TEF). — Soalala, village le plus proche Anjiafitatra, [ 15°26’S, 46°37’E], 100 m, 13.X.1967, fl., Service Forestier21383( MO, P, TEF). — Nde Soalala(baie de Baly), [ 16°05’S, 45°09’E], 100 m, 22.XI.1965, fr., Service Forestier24248( TEF). — Plateau de Berivotra, [ 15°54’S, 46°34’30”E], 100 m, 24.XI.1965, fl., Service Forestier24293( P, TEF).
DISTRIBUTION AND ECOLOGY Lepisanthes perrierioccurs in semi-deciduous and deciduous forests from western Madagascar(Ambongo, Soalala) to the far north of the island (Montagne d’Ambre PN, Ankarana RS and Daraina) and appears to grow on limestone and basement rock ( Fig. 1).
REMARKS Lepisanthes perrierican easily be distinguished by its leaves usually with just a single or two (rarely three) pairs of leaflets (vs. consistently two pairs in L. chrysotrichaand three or rarely two or four in L. sambiranensis), a petiole and rachis that combined usually measure less than 2.5 (occasionally up to 11) cm (vs.4.5[-6]cm in L.chrysotrichaand7-15[-22] cm in L. sambiranensis), and a glabrous inflorescence (vs. golden indument covering the whole inflorescence in L. chrysotricha, but glabrous in L. sambiranensis).
CONSERVATION STATUS With an EOO of 74 179 km2, an AOO of 133 km2, and 19 subpopulations, six of which are situated within a protected area (Ankarana, Baie de Baly), Lepisanthes perrieriis assigned a preliminary status of Least Concern (LC).
[926,1089,1129,1153]
B
de l'Academie & du Museum d'Histoire
Seances de l'Academie des Sciences
Memoires de l'Academie
3
304
1
1909-10
TAN
Perrier de la Bathie
Madagascar
-16.266666
Manongarivo
1282
45.366665
3
304
P 00624104, K 000426188, P 00624103
1
2006-12-07
MO, P, TAN
Prov. & Ankarana & Andriamihajarivo
Madagascar
172
4
305
-12.908889
Antsiranana
21
49.109722
3
304
1
Antsiranana
1993-10-12
1993-10-20
1993-10-12
MA, MO, P, TAN
Ankarana & Andrianantoanina
Madagascar
150
-12.816667
21
49.02139
4
305
1
Antsiranana
1995-11-21
PA, G, MO, P, TAN
Ankarana & Andrianantoanina & Bezara
Madagascar
276
-12.876667
d'Ambondromifehy
21
49.22028
4
305
1
Antsiranana
1995-12-05
G, MO, P, TAN
Montagne d'Ambre & Bobakilandy & Andrianantoanina & Bezara
Madagascar
533
-12.626945
21
49.10722
4
305
1
Antsiranana
1916-12-22
2001-10-30
1916-12-22
P, TAN, K, MO
Ankarana AP & Bardot-Vaucoulon & Andrianantoanina & Toly & Abattoir & Decary
Madagascar
150
-12.893888
21
49.113052
4
305
1
Antsiranana
1996-11-19
K, L, MO, P, TAN, WAG
Ankarana & Labat
Madagascar
180
-12.911111
21
49.10861
4
305
1
Antsiranana
2001-11-05
G, TEF, DARAINA, P, MO, K
Daraina & Binara & Gautier & Ravelonarivo
Madagascar
300
-13.25
1291
49.616665
4
305
1
Antsiranana
2006-11-22
G, TEF, DARAINA, P, MO, K
Daraina & Bekaraoka & Gautier & Chatelain
Madagascar
450
-13.1
1291
49.7
4
305
1
Antsiranana
1992-11-22
1992-11-26
1992-11-22
K, G, MO, P, TAN
Ankarana & Malcomber & Ankarana AP
Madagascar
150
-12.85
1292
49.066666
4
305
1
Antsiranana
[233,513,1011,1035]
MO
Malcomber
Madagascar
4
305
1
Antsiranana
1992-11-22
1992-11-26
1992-11-22
MO, P, TAN, WAG
Ankarana & Malcomber
Madagascar
150
-12.85
1292
49.066666
4
305
1
Antsiranana
1992-11-22
1992-11-26
1992-11-22
MA, MO, P, TAN
Ankarana & Malcomber
Madagascar
150
-12.85
1292
49.066666
4
305
1
Antsiranana
1993-11-24
MO, P, TAN, TEF
Ankarana & McPherson
Madagascar
320
-12.866667
1292
49.233334
4
305
1
Antsiranana
1936-10
G, MO, P
Nosy Mitsiou & Perrier de la Bathie
Madagascar
-12.9
1292
48.6
4
305
1
Antsiranana
2005-11-05
CNARP, MO, P, TAN
Ampondrabe & Rakotonandrasana
Madagascar
200
-12.971666
21
49.69917
4
305
1
Antsiranana
2005-11-15
CNARP, MO, P, TAN
Befarafara & Rakotonandrasana
Madagascar
100
-13.090555
21
49.575554
4
305
1
Antsiranana
2005-11-11
CNARP, MO, P, TAN
Daraina & Bobankora & Rakotondrajaona
Madagascar
200
-13.184444
21
49.758335
4
305
1
Antsiranana
2006-01-28
I, G, DARAINA, MO, K, P, TEF
Daraina & d'Antsahabe & Ranarison
Madagascar
340
-13.216666
1291
49.55
4
305
1
Antsiranana
2004-11-06
CNARP, MO, P, TAN
Ananjaka & Randrianaivo
Madagascar
155
-13.1
21
49.168613
4
305
1
Antsiranana
2005-12-11
CNARP, MO, P, TAN
Antsisikala & d'Analabe & Randrianaivo
Madagascar
184
-12.196944
21
49.197502
4
305
1
Antsiranana
2007-12-10
S, CNARP, MO, P, TAN
Ankarana & Randrianasolo
Madagascar
127
-12.982499
21
49.16389
4
305
1
Antsiranana
2005-10-31
CNARP, MO, P, TAN
Ankijomantsina & Ratovoson
Madagascar
100
-13.127778
21
49.46472
4
305
1
Antsiranana
2007-12-03
CNARP, MO, P, TAN
Antsisikala & Ratovoson
Madagascar
100
-49.210556
18
12.833889
4
305
1
Antsiranana
2006-11-10
Z, S, MO, P, TAN, G, K, US
Ankarana & Rogers
Madagascar
180
-12.9
1292
49.133335
4
305
1
Antsiranana
1992-10-24
S, N, MO, P, TAN
Schatz
Madagascar
200
-12.883333
Anivorano
1292
49.2
4
305
1
Antsiranana
1958-10-19
G, K, MO, P, TEF
Massif de la Montagne d'Ambre & Service Forestier
Madagascar
300
-12.5
1293
49.083332
4
305
300
Antsiranana
1955-11-04
TEF
Windsor Castle & Service Forestier
Madagascar
100
-12.233334
925
49.175003
4
305
100
Antsiranana
1959-10-24
TEF
Ambondromifehy & Service Forestier
Madagascar
300
-12.891666
21
49.208336
4
305
1
Antsiranana
1958-11-12
TEF
Pres d'Ambohimagodra & Service Forestier
Madagascar
100
-13.025
901
49.133335
4
305
100
Antsiranana
1958-11-14
P, TEF
Ankarana & Marovato & l'Ankerana & Service Forestier
Madagascar
200
-12.808333
902
49.15
4
305
1
Antsiranana
1961-11-09
S, P, TEF
Service Forestier
Madagascar
250
-12.474999
S du Pic Raynaud
21
49.44167
4
305
1
Antsiranana
1961-11-09
S, TEF
Service Forestier
Madagascar
250
-12.474999
S du Pic Raynaud
21
49.44167
4
305
1
Antsiranana
1966-12-18
P, TEF
Mafokovo & Service Forestier
Madagascar
250
-13.3
Nord de Vohemar
1291
49.85
4
305
1
Antsiranana
1910-10-02
P
Perrier de la Bathie
Prov.
-16.266666
Ambongo
1282
45.366665
Manongarivo
4
305
1
Mahajanga
1926-12
2004-11-16
1926-12
P, MO, TEF
Perrier de la Bathie & Ramananjanahary
Soalala
217
-15.649445
Bongolava
21
47.584164
Ambongo
4
305
1
Mahajanga
2004-11-27
MO, P, TEF
Ramananjanahary
115
-15.566667
Borizini
21
47.49861
Bongolava
4
305
1
Mahajanga
2001-05-20
V, R, CNARP, MO, P, TAN
Randrianaivo
20
-15.588333
Antsianitia
21
46.433613
Prov.
4
305
1
Mahajanga
1958-10-01
P, TEF
Service Forestier
-15.016666
Analalava
1286
47.266666
Ambondro
4
305
1
Mahajanga
1961-11-29
P, TEF
Service Forestier
256
-14.9
Bora
1286
48.216667
Massif
4
305
1
Mahajanga
1958-10-29
1958-11-03
1958-10-29
P, TEF
Service Forestier
50
-15.016666
Foret d'Ambondro-Ampasy
1286
47.266666
4
305
1
Mahajanga
1967-10-13
MO, P, TEF
Service Forestier
100
-15.433333
Anjiafitatra
1285
46.616665
Soalala
4
305
1
Mahajanga
1965-11-22
N, TEF
Service Forestier
100
-16.083334
Baly
1283
45.15
4
305
1
Mahajanga
1965-11-24
P, TEF
Service Forestier
100
-15.9
Plateau de Berivotra
925
46.575
4
305
1
Mahajanga