Cyana bianca malayana Bucsek, 2012 Erebidae, Arctiinae A review of the Cyana bianca (Walker, 1856) species-group with descriptions of a new species and a new subspecies from Indochina (Lepidoptera, Erebidae, Arctiinae) Volynkin, Anton V. Černý, Karel Ivanova, Maria S. BFDWK Bucsek, 2012 Volynkin & Černý & Ivanova 2018 Bucsek 2012 [151,526,151,178] Insecta Erebidae Cyana GBIF Animalia Lepidoptera 12 Arthropoda species malayana stat. nov.    Cyana bianca malayana Bucsek, 2012,  Erebidae, Arctiinaeof Malay Peninsula – Malaysia 2012: 100, pl. 17, figs. 248, 248а, 248b ( Typelocаlity: “ Mаlаysiа, Pаhаngdistr., Cаmeron Highlаnds, Kаmpung Kuаlа Boh vill. env., 850–1050 m, 4°27,9’N, 101°34,8’E”).     Typematerial examined.  Holotype( Figs. 11, 31): ♂, Malaysia, Pahangdistr., Cameron Highlands, Kampung Kuala Bohvill. env., 4°27,9’N, 101°34,8’E,  850–1050 m,  26.III.–3.IV.2001, K. Bucsekleg., slide MWM 33517 Volynkin( Coll.MWM/ ZSM).   Paratypes: 3 ♂, same data as in the holotype, slides MWM 31806, MWM 33483 Volynkin (Coll. MWM/ ZSM).   Other material examined:  MYANMAR: 1 ♂,  Burmamerid., Tenasserim, Khao Yai,  1000 m, 99’30’’E, 13’14’’N, Lf,  VII.1995, lg. Steinke & Lehmann, slide MWM 33488 Volynkin( Coll.MWM/ ZSM);  1 ♀, Burmah, Adanusn. / Rothschild Bequest B.M.1939.-I. ( Coll. NHMUK);   THAILAND: 1 ♂, W Thailand, Kanchanaburi,  SriSawat, Ban Dong Lek,  10.V. 2005, 735 m, leg. K. Sukhumalind, slide AV 3855 ♂ Volynkin( Coll.CKC);  6 ♂, SW Thailand,  Ranong,  543 m, 10°02’01’’N, 98°40’17,66”E,  7.IV.2013, leg. K. Černý, slides AV 3852 ♂, AV 3853 ♂ Volynkin( Coll.CKC);  8 ♂, 1 ♀, SW Thailand, Ranong[prov.],  Ranong,  380 m, 10°01’32”N, 98°40’13”E,  3– 4.XII.2005, leg. K. Černý, slide AV 3854 ♂ Volynkin( Coll.CKC);  1 ♂, 4 ♀, SW Thailand, Chumphon, Pa Toh, Ban Lang Tang,  162 m, 9°46’5”N, 98°46’59”E, 2006 ex ovo, K. Černý( Coll.CKC);  1 ♂, SW Thailand,  Chumphon, Pa Toh, Ban Lang Tang,  162 m, 9°46´5”N, 98°46´59”E, 19,  21.XI.2006, leg. K. Černý, slide AV3851 ♂ Volynkin( Coll.CKC);  4 ♂, SW Thailand, Chumphon, Pa Toh, Ban Lang Tang,  162 m, 9°46’5”N, 98°46’59”E,  7.XII.2005, leg. K. Černý( Coll.CKC);  1 ♂, Muang Thai, Changwat Chumphon, Amphoe Pa Toh,  10 kmW Baan Lang Tang, primary forest,  160 m,  11–12.VII.2007, lg. Manghong Jantharamphorn & Lechner, ex coll. Lechner( Coll.CKC);  2 ♂, SW Thailand, Nakhon Si Tammarat, Khao Luang region, Nopphitamenv.,  70 m, 8°44.043’N, 99°41.450’E,  12–13.IV.2013, leg. K. Černý( Coll.CKC);  3 ♂, 2 ♀, SW Thailand, Nakhon Si Tammarat, Amphoe Ron Pibun, Ang Ka Tai Hill, 8°14.16’N, 99°48.12’E,  9.IV.2013leg. K. Černý( Coll.CKC);   MALAYSIA: 6 ♂, 2 ♀,  Malaysia, Prov. Perak, Taiping, VI–  V.1978, coll. Dr. W. Thomas, slides ZSM Arct.145/2017 ♂, MWM 33485 ( ♂),  ZSMArct. 146/2017 ♀, MWM 33514 ( ♀) Volynkin( Coll.MWM/ ZSM);  1 ♂, Malaysia, Taiping, coll. Dr. W. Thomas( Coll.MWM/ ZSM);   INDONESIA: 1 ♂, W Sumatra, Padang Panjang, Lembah Anai,  300 m, 0°27’S, 100°23’E,  I.2004, leg. Jakl, slide MWM 33486 Volynkin( Coll.MWM/ ZSM);  1 ♂, W Sumatra, Mt. Sanggul, Landai,  1200–1300 m, 0°00’, 100°38’E,  I–II.2004, leg. St.Jakl, slide MWM 33487 Volynkin( Coll.MWM/ ZSM);  1 ♂, Indonesia, North Sumatra, Via Sindar Raya – Pematang Raya, 11 kmof SR,  400 m,  24.III.1986, E. Diehlleg. ( Coll.CKC);  1 ♂, Sumatra, Via Sindar Raya, 11 kmof SR,  400m, 3°53’N, 98°51’E  16.III.1991, E. Diehlleg. ( Coll.CKC).   Remarks. 1.The male and female genitalia of  malayana( Figs. 31–33, 40, 41) have significant differences from those of  C. bianca( Fig. 25–27, 37). Therefore we treat this taxon as a good species and not a subspecies of  C. bianca. 2.The typeseries from Cameron Highlands has reduced black borders of the ante- and postmedial lines ( Fig. 11), while the specimens from other parts of the species’ range north and south of Cameron Highlands have the well-developed black borders of the transverse lines ( Figs. 12–16). Because of the absence of genital differences ( Figs. 31–33), we treat this population as a local color form only. 3.The female paratypesof the species ( Fig. 42) are not conspecific to the male holotypeand paratypesand belong to  Cyana determinata sausae Bucsek, 2012described by males also from Pahang.   Diagnosis. Forewing length is 14.5– 18 mmin males and 19–20 mmin females. Externally, C. malayana( Figs. 11–16) has no significant differences from  C. bianca( Figs. 1–4) and  C. indosinica  sp. nov.( Figs. 5–10). The male genitalia of  C. malayana( Figs. 31–33) differ clearly from those of  C. bianca( Fig. 25–27) in the rounded lateral lobes of the vinculum, the larger and more robust ampulla, the much larger medial posterior, medial anterior and distal diverticula, and the broader medial dorsal diverticulum; from  C. indosinica  sp. nov.( Figs. 28–30) they differ in the significantly larger and more robust ampulla, the narrower medial posterior and distal diverticula, which have narrower fields of cornuti, the smaller medial anterior diverticulum, and the basally narrower medial dorsal diverticulum. The female genitalia of  C. malayana( Figs. 40, 41) differ from those of  C. bianca( Fig. 37) in the longer ductus bursae, the larger appendix bursae, the shorter and stronger rugose posterior sclerotized plate of the corpus bursae, the weakly sclerotized medial-lateral sclerotized band of the corpus bursae, and the much smaller signa; from  C. indosinica  sp. nov.( Figs. 38, 39) they differ in the more weakly sclerotized and much less rugose posterior plate of the corpus bursae, and the presence of only two signa in the corpus bursae (while in  C. indosinica  sp. nov.there are three signa in the corpus bursae).   Distribution( Fig. 46). South Myanmar( Hampson 1900), South-West and South Thailand(provinces Kanchanaburi, Chumphon, Ranongand Nakhon Si Tammarat), Malaysia(Malay Peninsula), Indonesia( SumatraIsl.). 1914299851 2001-03-26 2001-04-03 2001-03-26 ZSM K. Bucsek & Volynkin & Coll. Malaysia 950 4.465 Kampung Kuala Boh 130 101.58 Cameron Highlands 12 1 1 Pahang holotype 1914299841 2001-03-26 2001-04-03 2001-03-26 ZSM K. Bucsek & Volynkin & Coll. Malaysia 950 4.465 Kampung Kuala Boh 130 101.58 Cameron Highlands 12 3 3 Pahang paratype 1914299864 [199,750,530,555] Myanmar MYANMAR 12 1 1 1914299843 1995-07 ZSM Volynkin & Coll. Myanmar 1000 Lf 12 1 1914299856 NHMUK Rothschild Bequest B. M. Myanmar Coll. 12 1 1 1914299858 [865,1392,602,627] W Thailand Thailand Kanchanaburi 12 1 1 Kanchanaburi 1914299842 2005-05-10 K. Sukhumalind & Volynkin & Coll. Thailand Sri 12 1 1 Sri 1914299865 Thailand Thailand 12 6 6 1914299849 2013-04-07 K. Cerny & Volynkin & Coll. Thailand 543 10.033611 Ranong 15 98.67157 12 2 2 Ranong 1914299846 [423,892,710,735] Thailand Ranong 12 9 1 8 Ranong 1914299860 2005-12-03 2005-12-04 2005-12-03 K. Cerny & Volynkin & Coll. Thailand 380 10.025556 Ranong 21 98.67027 12 1 1 Ranong 1914299844 K. Cerny & Coll. Thailand 162 9.768055 Ban Lang Tang 21 98.78306 Pa Toh 12 5 4 1 Chumphon 1914299862 [1204,1431,782,807] Thailand Thailand 12 1 1 1914299855 2006-11-21 K. Cerny & Volynkin & Coll. Thailand 162 9.768055 Ban Lang Tang 21 98.78306 Pa Toh 12 1 1 Chumphon 1914299848 2005-12-07 K. Cerny & Coll. Thailand 162 9.768055 Ban Lang Tang 21 98.78306 Pa Toh 12 4 4 Chumphon 1914299861 2007-07-11 2007-07-12 2007-07-11 Lechner & Coll. Thailand 160 Amphoe Pa Toh Muang Thai 12 1 1 Changwat Chumphon 1914299863 2013-04-12 2013-04-13 2013-04-12 K. Cerny & Coll. Thailand Nakhon Si Tammarat 70 8.73405 Nopphitam 1 99.690834 Khao Luang region 12 2 2 1914299845 2013-04-09 K. Cerny & Coll. Thailand Nakhon Si Tammarat 8.236 Ang Ka Tai Hill 13 99.802 Amphoe Ron Pibun 12 5 2 3 1914299854 Malaysia MALAYSIA 12 8 2 6 1914299868 1978-05 ZSM Dr. W. Thomas & Arct. Malaysia Taiping 12 2 2 Perak 1914299852 [205,1021,1106,1131] ZSM Malaysia Coll. Volynkin 12 2 2 Perak 1914299859 ZSM Dr. W. Thomas Malaysia Coll. Taiping 12 1 1 1914299850 2004-01 ZSM Jakl & Volynkin & Coll. Indonesia 300 -0.45 Lembah Anai 1308 100.38333 Padang Panjang 12 1 1 Sumatra 1914299847 2004-01 2004-02-31 2004-01 ZSM St. & Volynkin & Coll. Indonesia 1250 Landai 100.63333 Mt. Sanggul 12 1 1 Sumatra 1914299853 1986-03-24 E. Diehl & Coll. Indonesia 400 Via Sindar Raya - Pematang Raya 12 1 1 North Sumatra 1914299857 1991-03-16 E. Diehl & Coll. Indonesia 400 3.8833332 Via Sindar Raya 1307 98.85 12 1 1 Sumatra