A Catalogue and Historical Study of the Odonata of Ecuador
Mauffray, William F.
Tennessen, Kenneth J.
Zootaxa
2019
2019-07-03
4628
1
1
265
32ML
Rambur, 1842
Rambur
1842
[151,392,1690,1716]
Insecta
Coenagrionidae
Argia
Animalia
Odonata
35
36
Arthropoda
genus
Argiais the largest genus of damselflies in the New World, with over 138 valid species names and at least seven species yet to described ( Garrison & von Ellenrieder 2018). Adults range from small to large and are variable in color pattern, the thoracic dorsum being black with blue or purple stripes (rarely orange) to entirely metallic red or copper, the thoracic sides usually blue, and the abdomen from mostly blue with black markings to mostly black with small blue markings ( Figs. 16a & 16b), rarely largely orange. Species identification relies on Calvert (1902), and Garrison & von Ellenrieder (2015, 2017, 2018). Thirty-two species have been recorded in Ecuador, of which three are considered endemic. Habitat of most species in Ecuadoris small streams to mid-sized rivers, with a few found in pooled areas of slow streams, seepage streams or sheet flow over boulders. FIGURE 15. Anisagrion inornatum ♂, ECUADOR, Loja Province., seep, trib. of River Chamba, Vilcabamba, 30 iii 2008; photo K. J. Tennessen. Argia acridens Garrison & von Ellenrieder, 2018. Publ. Recs. ES:* Garrison & von Ellenrieder (2018); MN:* Garrison & von Ellenrieder (2018— Holotype: Flavio-Alfaro Canton: 79kmwest of Santo Domingo de los Colorados ( -0.3333, -79.6666, 260m), 7 v 1975, leg. Paul J. Spangler et al.[USNM]); SD: * Garrison & von Ellenrieder (2018). Note: Endemic to Ecuador’s Pacific slope. Argia adamsiCalvert, 1902. Publ. Recs. ECU: (P11), (H14); LR: Garrison & von Ellenrieder (2018); MN: Garrison & von Ellenrieder (2018); MS: Garrison & von Ellenrieder (2018); NA:* Tennessen (1996b), Daigle (1996), Garrison & von Ellenrieder (2018); OR: Garrison & von Ellenrieder (2018); PA: Garrison & von Ellenrieder (2018); PC: Garrison & von Ellenrieder (2018); SD: Garrison & von Ellenrieder (2018); TU: Garrison & von Ellenrieder (2018). Also CEN, PER, VEN: (H08), (P11); Plus COL: (H08), P&P (2011), (P16), and BOL, BRA: Garrison & von Ellenrieder (2018). Addl. Data. ES, San Lorenzo:[FSCA. JJD, KJT]; LR, Quevedo:[FSCA, JTJ, KJT]; MS, Gualaquiza:[USNM]; Huambaya:[KJT, RWG]; NA, Tena:[FAL, FSCA, UTIC, JJD, KJT, RWG, TWD, USNM]; OR, Francisco de Orellana:[FSCA, JJD, KJT, RWG, SWD]; Loreto[FSCA, JTJ, KJT, RWG, USNM]; PA, Pastaza:[FSCA, JJD, JTJ, KJT, RWG, USNM]; SD:[RWG]; SU, Lago Agrio:[SWD]; Shushufindi:[UTIC]. Note: In east slope form the light coloration on the thorax is blue, and in some west slope forms it is lighter blue or pale yellowish, especially from northwest Ecuador. According to Garrison & von Ellenrieder (2018), L. K. Gloyd considered A. adamsifrom Ecuadorwest of the Andes to represent a new species to which she had applied the manuscript name A. lorena. However, Garrison & von Ellenrieder (2018)found no significant differences in male caudal appendages nor female mesostigmal plates. FIGURE 16a. Argia fulgida♂, ECUADOR, Pichincha Province, small stream, Mashpi Road, 2 x 2018; photo K. J. Tennessen. FIGURE 16b. Argia pulla♂, ECUADOR, Bolivar Province., stream 5 km E of Caluma, 5 iii 2010; photo K. J. Tennessen. Argia collataSelys, 1865. Publ. Recs. ECU: *Tennessen (1996b—as A. thespis), (L06—as A. thespis), (H08), (H14), Garrison & von Ellenrieder (2018); NA:* Garrison & von Ellenrieder (2015); OR:* Garrison & von Ellenrieder (2015); SU: * Garrison and von Ellenrieder (2015). Also BRA: (L06—as A. thespis), Garrison & von Ellenrieder (2015), (H08), (P11—as A. thespis), GUF: (H08); plus COL, PER, VEN: Garrison & von Ellenrieder (2015); and BOL (P16). Addl. Data. EO, Santa Rosa: 1 ♂, stream 12 kmE of Arenillas, Hwy. 25 ( -3.5244, -79.9831, 30m), 29 iii 2008, leg. & det. KJT—as A. thespis[FSCA]; GY, Balao: 1 ♂, stream, banana plantation, 2 kmE of Hwy. 25 & Balao Jct. ( -2.9100, -79.6919, 70m), 28 iii 2008, leg. & det. KJT—as A. thespis, [FSCA]; NA, Tena: 4 ♂♂, 1 ♀, Río Chinchipino, 27.7 kmE of Puerto Napo( -1.0728, -77.5864, 385m), 13 xi 1997, leg. & det. J. Daigle—as A. thespis[JJD]; OR, Franciscode Orellana: 1 ♂, stream 2.3 kmE of Yasuni Sci. Res. Sta., P.N.Y., 9 vi 1995, leg. & det. KJT—as A. thespis[KJT]; 1 ♂, same data but leg. & det. WFM—as A. thespis[FSCA]; 1 ♂, seepage 2.1 kmE of first road past research station, YasuniPreserve, 15 xi 1997, leg. & det. JJD—as A. thespis[JJD]; 1 ♀, stream WaoraniRoad, 2.0 km past cutoff, YasuniPreserve, 16 xi 1997, leg. & det. JJD—as A. thespis[JJD]; SU, Cascales: 1 ♀, stream 50 kmW of Lago Agrío( 395m), 8 vi 1995, leg. & det. KJT—as A. thespis[KJT]; Shushufindi: 1 ♂, stream about 3 kmNW of Shushufindi( -0.1664, -76.6167, 280m), 14 v 2009, leg. & det. KJT—as A. thespis[KJT]. Note: According to Garrison (pers. comm. 2011), “ A. collatawas thought to be a synonym of A. thespis, but it has been elevated back to distinct species. Specimens labeled and/or published as A. thespisfrom Ecuadorshould be this species.” Argia cuspidata Garrison & von Ellenrieder, 2018. Publ. Recs. ES:* Garrison & von Ellenrieder (2018— ♀para- type, from tandem pair), San Lorenzo:seeps 1.1 kmNW of Lita at bridge, 5:00 pm [ 0.8850, -78.4755, 470m], 2 ii 1997, leg JJD, [JJD]); SD:* Garrison & von Ellenrieder (2018— Holotype: 19 kmE of Santo Domingo de los Colorados [ -0.3137, -79.0289, 740m], 7 v 1975, leg. A. Langley & J. Cohen [USNM]). Note: Endemic to limited area of Pacific slope. Argia difficilisSelys, 1965. Publ. Recs. ECU: (H08); (P11); MS:Garrison & von Ellenrieder (2007); NA:Garrison & von Ellenrieder (2007); OR:Garrison & von Ellenrieder (2007); PA: Garrison & von Ellenrieder (2007); SU:Garrison & von Ellenrieder (2007). Also BRA, CEN, COL, PER, VEN, T&T: (H08). Addl. Data. MS, Gualaquiza:[RWG, UMMZ, USNM]; NA, Tena:[DRP, FSCA, RWG, UMMZ]; OR, Francisco de Orellana:[FSCA, KJT, NVE, SWD, TWD, USFQ, USNM]; Loreto:[KJT, RWG, UMMZ]; PA, Mera:[FSCA]; Pastaza: [RWG]; SU, Shushufindi: [DRP, FSCA, RWG, SWD]. Argia dives Förster, 1914. Publ. Recs. ECU:(L06), (H08), (P11), (H14); MS: * Tennessen (1998), Garrison & von Ellenrieder (2018); NA: Garrison & von Ellenrieder (2018); OR: Garrison & von Ellenrieder (2018); PA: Tennessen (2006), Garrison & von Ellenrieder (2018); SU:( Garrison & von Ellenrieder, 2018); TU: Garrison & von Ellenrieder (2018). Also PER: (L06), (P11); plus BRA: (L06); and COL: P&P (2011), (P16). Addl. Data. MS, Huamboya:[JJD]; Morona:[RWG]; Gualaquiza:[FSCA]; NA, Archidona:[RWG]; Carlos Julio Arsemena Tola: [RWG]; Tena:[FSCA, KJT, RWG, USNM]; OR, La Joya de los Sachas:[FSCA]; Loreto:[DRP, FSCA, JJD, JTJ, KJT, RWG, TWD, USNM]; PA, Mera:[FSCA, JJD, KJT, RWG, USNM]; Pastaza:[FAL, FSCA, JJD, KJT, RWG, SWD, SWD]; SU, Shushufindi:[FSCA, RWG, SWD]; ZC, El Pangui:[FSCA, JJD, KJT]. Argia euphorbiaFraser, 1946. Publ. Recs. ECU:(P16), Garrison & von Ellenrieder (2018); * OR: von Ellenrieder & Garrison (2011), Garrison & von Ellenrieder (2015). Addl. Data. OR, Franciscode Orellana: 1 ♂, T. B.S.( -0.6333, -76.1500, 220m), date?, leg. K. Jackson, D. Inward, det. RWG-2003[BMNH] AlsoPER: ( H08), (P11); plus ARG, BRA: ( H08); plus COL, VEN: ( P16). Note: The Argentinarecord is an error ( Garrison & von Ellenrieder 2015). Argia fraudatricula Förster, 1914. Publ. Recs. ECU:(*H08), (P11), (H14), Garrison & von Ellenrieder (2018). Also PER: (H08), (P11). Addl. Data. NA, Tena: 1 ♂, Jatun Yacu, near mouth of Ilapulin, 8 iii 1937, leg. WCM, det. JJD [FSCA]; same but 4 ♂♂, det. L. Gloyd, from RWG [RWG, UMMZ]; 3 ♂♂, Jatun Yacu, Río Napowatershed ( -1.0167, -77.8332, 700m), 4 iii 1937, leg.WCM, det. L. Gloyd, from UMMZ [RWG, UMMZ]; same but 3♂♂, 2 ♀♀, 21 iii 1937, [RWG, UMMZ]; PA: 4 ♂♂, Partidero Rivertrail [ El Partidero, Indian village on east bank of Río Anzu, jct. of trails from Mera and Puyo northward to Napo], Río Napowatershed ( -1.0167, -78.000, 1000m), 16 ix 1935, leg. WCM, det. L. Gloyd, from RWG [RWG, UMMZ]; ZC, Nangaritza: 7 ♂♂, 2 ♀♀, Río Nangaritza(-4.55, -78.8167, 1400m), 2 xii 1941, leg. D. Laddey, det. L. Gloyd[RWG, UMNZ]; same but 1♂, 1 ♀, Mayaicu, Río Nangaritza( - 3.9894, -78.6328, 1136 m) [RWG]; same but 2 v 1941[RWG]. Note: There are no recent records of this species despite numerous field surveys within that part of Ecuadorwhere it was recorded more than 75 years prior to this report. Argia fulgida Navás, 1934. Publ. Recs. BO:* Garrison & von Ellenrieder (2017, 2018); EO:* Garrison & von Ellenrieder (2017, 2018); ES:* Garrison & von Ellenrieder (2017, 2018); LR:* Garrison & von Ellenrieder (2017, 2018); MN:* Garrison & von Ellenrieder (2017, 2018); PC: Garrison & von Ellenrieder (2018); SD:* Garrison & von Ellenrieder (2017, 2018);. Also CEN, COL: Garrison & von Ellenrieder (2017). Addl. Data. BO, Caluma: 4 ♂♂, rocky stream, 5 kmE of Caluma, Guaranda Road ( -1.5942, -79.2308, 480m), 5 iii 2010, leg. JJD, det. RWG [JJD, RWG]; 4 ♂♂, same data but KJT [FSCA, KJT]; CN, Cañar: 2 ♂♂, Cochancay 20.2 kmE of Prov. Border on Hwy.70 rocky stream ( -2.4686, -79.2853, 195m), 3 xi 1997, leg, det. JJD [FSCA]; EO, Portovelo: 1 ♂, Portovelo( -3.7167, -79.6167, 650m), 1 vii 1941, leg. D. Laddey, det. RWG [RWG]; ES, San Lorenzo: 1 ♂, rocky stream 36.5 kmNW of Lita, near Urbina, 3 ii 1997, leg. & det. JJD; 1 tan. pair, 1 ♂, [San Antonio], nearby, 36.5 kmWNW of Litaon [old] San Lorenzo Hwy, pools/seeps/stream (0.95, -78.68, 200m), 3 ii 1997, leg. & det. RWG [FSCA]; LR, Babahoyo 2 ♂♂, 2 ♀♀, Playas de Juan Montalvo, near Balzapamba( -1.6833, -79.2833, 15m), 28 ii 1938, leg. WCM, det. RWG, from UMMZ [RWG]; 1 ♂, 1 ♀, same data but 8 iv 1938[RWG]; 2 ♂, same data but 15 iv 1938[RWG]; 2 ♂♂, same data but 17 iv 1938[RWG]; 1 ♂, same data but 31 vii 1941[RWG]; PC, Quito: 6 ♂♂, upper Mashpi Road, Rio Huaycuyacu, above waterfall ( 0.1952, -78.8657, 850m), 2 ix 2018, leg. & det. WFM [FSCA, USFQ]; 2 ♂♂, 1 tan. pair, same data but leg & det. KJT [KJT]; SD: 3 ♂♂, 1 ♀, Río Pove, Santo Domingode los Colorados(-0.25, -79.15, 600m), 27 ix 1940, leg. D. Laddey, det. RWG, from UMMZ [RWG]; 1 ♂, 1 ♀, same data but 28 ix 1940[RWG]; 1 ♂, Río Mulaute, near Santo Domingode los Colorados; virgin forest, 3 iii 1973, leg. M. Deyrup, det. RWG, from UMMZ [RWG]; 1 ♂, small stream, Hwy.30, near km marker 71, near La Unión del Toachi( -0.3139, -78.9436, 900m), 25 ii 2010, leg. & det. JTJ [JTJ]. Note: All records are from Pacific slope. Argia fumigataHagen inSelys, 1865. Publ. Recs. ECU:(P16), Garrison & von Ellenrieder (2018); * OR: Garrison & von Ellenrieder (2015). Also: BRA, GUF, GUY, SUR, VEN: (P16). Addl. Data. OR: Franciscode Orellana: 1 ♀, Reserva Étnica Waorani, 1 kmS Okone Gare Camp, canopy fogging project, Lot#1546, TR3 #6, terra firme forest ( -0.6569, -76.4528, 216 m), 21 vi 1996, leg. T.L. Erwin et al., det. RWG [USNM]. Note: This record is the westernmost for this predominately Guianashield species. Argia gerhardi Calvert, 1909. Publ. Recs. ECU: (H08), (P11), (H14). Also BOL, COL, PER: (H08), (P11). Addl. Data. Provided by R. Garrison (pers. comm., 25 vi 2018): LJ, Loja: 1 ♂, 1 ♀, Estación Cientifica San Francisco, Río San Francisco, small creek, oviposition in soil ( -3.9744, -79.0778, 1870m), 28 x 2014, leg. C. Willigalla, det. RWG [ C. Willigalla]; NA, Archidona: 4 ♂♂, 1 ♀, seepages near river, 43.5 kmW Loreto( -0.7212, -77.6410, 880m), 14 vi 1995, leg & det. SWD [RWG]; 1 ♂, seep 44 kmeast of Narupajct., Loreto Road, 14 xi 1997, leg. KJT, det. RWG [RWG]; PA, Mera: 1 ♀, 20 kmon Puyoto Baños Hwy.( -1.4543, -78.1221, 1130m), 19 x 1997, leg. M. Kippenrand, det. RWG [RWG]. SU, Gonzalo Pizarro: 1 ♂, 1 ♀, San Rafael Falls, Río Quijos, 52 kmnortheast of Baeza( -0.1020, -77.5890, 1290m), 22 i 2009; leg DLW, WAH, Michael Thomas, Fred Morrison& Ronald Vargas, det. RWG [WAH]; TU, Baños: 1 ♀, Abitagua, Río Pastazawatershed, Hda. Guillermina( -1.4431, -78.1665, 1200m), 7 xii 1939, leg. WCM, det. L. Gloyd[RWG]; same but, 1 ♂, 9 xii 1939[RWG]; ZC, Zamora: 1 ♂, seepage near Río Sabanillo, W of Zamora( -4.0453, -79.0147, 1225m), 1 iv 2008, leg. & det. JTJ [JTJ]; 1 ♂, 1 ♀, tributary of Bombuscara Riveron road towards Zamora( -4.0428, -79.0142, 1280m), 18 viii 2006, leg. JCA, det. RWG [UTIC]. Notes: According to Rosser Garrison (pers. comm. 25-vi-2018), “The identification within the Argia gerhardi/ kokama/ nigriorcomplex is difficult. I have seen (and illustrated) typesof all three and have thus far done a lot of work on this group....Putting names on various populations is not always easy as I have had to compare various characters from Ecuadorean and Peruvian populations with the types( Bolivia, Peru) and the characters I have thus far seen to be of diagnostic value do not seem to be concordant. In particular, males of Argia kokamaapproach those of A. nigriorclosely and the females of A. gerhardiand A. nigriorare difficult to separate. All of this is going to require more work.....the records I have for Argia gerhardifrom Ecuadorare listed with the proviso that not all of the records may pertain to that species.” All records are from relatively high elevation, from 880–1870 m. Argia hamulataFraser, 1946. Publ. Recs. ECU: * Garrison & von Ellenrieder (2018), Also BOL (RWG, pers. comm.), PER: (H08). So far known from only one record: SD:1 tan. pair, stream, Tinalandia, E of Santo Domingo, Hwy. 30 ( -0.3000, -79.0667, 800m), 31 i 1997, leg. KJT, det. RWG, data provided by RWG [KJT]. Argia huanacina Förster, 1914. Publ. Recs. ZC:stream, north side of Guaguayme Bajo, 4 iv 2008, leg. & det. KJT [FSCA] * Tennessen & Johnson (2008), Garrison & von Ellenrieder (2018). Also BOL, PER (H08) (P11), Garrison & von Ellenrieder (2018). Notes: From Rosser Garrison (pers. comm., 2018): “This species was treated and illustrated in Garrison & von Ellenrieder (2015). von Ellenrieder & Garrison (2007)followed by Heckman (2008)illustrated the appendages of a male in the RBINS with the manuscript name “ lateralis” by Selys as A. huanacina. This is incorrect. Subsequent examination of these specimens showed the manuscript name lateralisSelysto represent A. infrequentulaas can be seen by comparing Figs. 6l-n in von Ellenrieder & Garrison (2007)and Fig. 3.1. 427 in Heckman (2008)to Figs. 10l-2 and 10 p- 2 in Garrison & von Ellenrieder (2015).” This southern Ecuadorrecord seems to be the northernmost record for this species. Argia incultaHagen inSelys, 1865. Publ. Recs. ECU:(*H08), (P11), (H14), Garrison & von Ellenrieder (2018). Also PER: (H08), (P11). Addl. Data. ZC: 2 ♂♂, 1 ♀, Pacocha ( -1.0577, -80.8472, 120m), 27 iv 1946, leg. L. Gomez Alonzo, det. L. Gloyd [RWG, UMMZ]; EO, Arenillas: 1 tan. pair, 2 ♂, clay-bottom stream, 14 kmS of Arenillas, road to Alamar ( -3.6656, -80.0869, 95m), 10 iv 2008, leg. & det. KJT [FSCA]; 3 ♂♂, same data but leg. & det. JTJ [JTJ]; 1 tan pair, farm stream, 11 kmS of Arenillas, road to Alamar ( -3.6436, -80.0989), 10 iv 2008, leg. & det. KJT [FSCA]; Piñas: 1 ♀, Piñas (-3.6667, -79.65, 1000m), 23 vii 1941, leg. D. Laddey, det. L. Gloyd, from CHK/ UMMZ [RWG]; Portoviejo: 9 ♂♂, 2 ♀♀, Cerro de Hojas ( -1.0432, -80.4311, 200m), 19 iv 1946, leg. L. Gomez Alonzo, det. MJW [CSCA, RWG, UMMZ], same but, 21 iv 1946, 10 ♂♂, 3 ♀♀, [CSCA, RWG, UMMZ]; same but, 16 ♂♂, 2 ♀♀, [CSXA, RWG, UMMZ]; 1 ♀, concrete ditch, Hwy. E40, E of Portoviejo ( -1.0672, -80.1350, 420m), 4 iii 2010, leg. & det. KJT [FSCA]; Santa Rosa: 1 ♂, stream, 12 kmE of Arenillas, Hwy. 25 ( -3.5244, -79.9831, 30m), 29 iii 2008, leg. & det. KJT [FSCA]; 2 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, same data but 9 iv 2008, leg. & det. KJT [FSCA]; LJ, Catamayo: 1 ♂, 1 ♀, small stream and tilapia pond on old road from Malacatos to Catamayo, ( -4.0312, -79.3558, 1290 m), 18 i 2014, leg. F. Morrison, R. Vargas, & E. Wyman, det. RWG [RWG]; MN, Chone: 2 ♂♂, 7.2 kmNNE of Chone, Boca de Mosquito, between Río Grande and Río Mosquito ( -0.6833, -80.1000, 110m), 24 x 1978, leg. J. Anderson, det. RWG, from USNM [JJD, RWG]; 1 ♂, small stream/trib. 3 kmN of Canuto( -0.7678, -80.1231, 60m), 3 iii 10, leg. & det. JTJ [JTJ]; 9 ♂♂, same data but leg. & det. JJD [JJD]; Flavio Alfaro: 1 ♂, borrow pit/small stream about 4 kmN of Flavio Alfaro( -0.3750, -79.8875, 295m), 3 iii 2010, leg. & det. JTJ [JTJ]; Jipijapa: 1 ♂, [Playa] San Jose, 4 milesinland (-1.38, -80.75, 60m), 12 iii 1970, leg.? det. KJT [FSCA]; 2 ♂♂, Puerto Lopez, 20 kmto Salinas, 5 iv 1998, leg. E. Tapia, det. JJD [JJD]; Junin: 2 ♂♂, 1 ♀, stream just S of Junin( -0.9317, -80.2256), 3 iii 2010, leg. & det. KJT [FSCA]; Manta: 1 ♂, Santa Rosa ( -1.0576, -80.8472, 20m), 11 v 1946, leg. L. Gomez Alonzo, det. L. Gloyd[RWG]; same but, 6 ♂♂, 2 ♀♀, 12 v 1946[CSCA, RWG, UMMZ]; same but, 6 ♂♂, 1 ♀, 14 v 1946[CSCA, RWG, UMMZ]; Pedernales: 2 ♂, small stream about 2 kmE of Pedernales( 0.0836, -80.0139, 90m), 28 ii 2010, leg. & det. KJT [FSCA]; 1 ♂, same data but leg. & det. JTJ [JTJ]. Note: Collection data indicate that A. incultais a low elevation west slope species. Argia indicatrixCalvert, 1902. Publ. Recs. ECU:(L06), (H08), (P11), (H14), Garrison & von Ellenrieder (2018); CN: Garrison & von Ellenrieder (2015); EO: Garrison & von Ellenrieder (2015); * ES: Daigle et al.(1997), Garrison & von Ellenrieder (2015); LR: Garrison & von Ellenrieder (2015); NA: Garrison & von Ellenrieder (2015); OR: von Ellenrieder & Garrison (2011), Garrison & von Ellenrieder (2015); PC: Garrison & von Ellenrieder (2015); * SD: Daigle et al.(1997) Garrison & von Ellenrieder (2015); SU: Garrison & von Ellenrieder (2015). Also CEN, COL, PER, VEN: (L06), (H08), (P11) plus BRA: (L06), (H08) and BOL: (P16). Addl. Data: BO, San Miguel:[RWG]; CN, La Troncal:[RWG, SWD]; EO, Santa Rosa:[FSCA, JTJ, KJT]; ES, San Lorenzo:[FAL, FSCA, JA, JJD, KJT, RWG, SWD]; LR, Babahoyo:[RWG]; Buena Fé:[DRP, SWD]; MN, Flavio Alfaro:[FSCA, KJT]; OR, Francisco de Orellana:[JJD, KJT, RWG, SWD]; Loreto:[JTJ, KJT]; PC, San Miguel Bancos:[RWG]; SD: [DRP, FAL, FSCA, JJD, KJT, RWG, SWD]; SU, Lago Agrío:[RWG]. Argia infrequentulaFraser, 1946. Publ. Recs. ECU:(H08), (P11), (H14), Garrison & von Ellenrieder (2018); NA: *Daigle (1996). Also COL: (H08, P11, P&P); and PER: (P16). Addl. Data. MS, Morona: [UMMZ]; NA, Tena:[BMNH, CSCA, FSCA, JJD, KJT, RWG, UMMZ]; OR, Francisco de Orellana:[FSCA, KJT, TWD, USNM]; Loreto:[JJD, KJT, USNM]; PA, Mera:[JJD, KJT, RWG, UMMZ]; Pastaza:[BMNH, CSCA, RWG, UMMZ]; Santa Clara:[JJD, RWG]; SU, Cascales:[FSCA, USNM]; Lago Agrio:[RWG]; Sushufindi:[FAL, FSCA, JJD, JTJ, KJT, RWG]; TU, Baños: [UMMZ]; ZC, Zamora:[RWG]. Note: Some of these older records need to be rechecked as they may be Argia selysi Garrison & von Ellenrieder (2018). Argia jocosaHagen inSelys, 1865. Publ Recs. S U: 1 ♂, Bellavista River, 32kmSE of Lago Agrio, 11 i 2014, leg. D. Wagner, W. Haber, F. Morrison, det. RWG [RWG] Garrison & von Ellenrieder (2018). Also COL, VEN: ( Garrison & von Ellenrieder 2018). Argia kokama Calvert, 1909. Publ. Recs. ECU: Garrison & von Ellenrieder (2018); MS: * Tennessen (2006); OR: von Ellenrieder & Garrison (2011, 2017); SU: Garrison & von Ellenrieder (2017); Also PER: (H08), (P11); plus COL: P&P (2011), (P16). Addl. Data. MS, Huamboya: 6 ♂♂, 1 tan. pair, Macas, 58 kmN on Hwy. 45 ( 70 kmN of Sucua), roadside ditch and tribs. near Río Pasanac ( -1.9447, -77.8536, 762m), 16 ix 2005, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, 1 ♀, same but: 10 xi197, leg. & det. JJD—as “ A. makoka”[JJD]; 1 ♂, same data but, leg. TWD, det. JJD—as “ A. makoka” [JJD]; Morona; 8 ♂♂, Macas 15.7 kmW on Macas-Guamote Roadsmall stream ( -2.2689, -78.1936, 1463m), 17 ix 2005, leg. & det. WFM [FSCA, QZAC, USNM]; 8 ♂♂, Macas 22.9 kmW on Guamote Road, 3.3 kmW on Río Albanicorocky stream and flowing roadside ditches ( -2.2342, -78.218, 1690m), 18 ix 2005, 8 ♂♂, leg. & det. WFM [FSCA, QZAC, USNM]; OR, Franciscode Orellana, 1 ♀, T. B.S., USFQ, open grassy area behind din- ing room, surrounded by bushes and trees and some seepage areas ( -0.6383, -76.1492, 220m), 14 i 2009, leg. & det. RWG & NVE [RWG]; 2 tan. pairs, same data but 15 i 2009; 1 tan. pair, 1 ♂, same data but, 16 i 2009, 1 ♂, same data but 16 i 2009, 1 ♂, 1 ♀, same data but 18 i 2009; 1 ♂, T. B.S., USFQ, black water stream crossing Guacamayotrail at marker 2050, 16 i 2009, 1 ♂, same data but T. B.S., USFQ, Mauritiapalm swamp off Guacamayotrail at marker 2100, 20 i 2009, 2 ♂♂, 1 ♀, same data but T. B.S., USFQ, Guacamayotrail between markers 450–2100, 22 i 2009, leg. & det. RWG & NVE [USFQ]; 1 ♂, 1 ♀, General Plaza Gutierrez: [about 7 kmN] & 22.5 kmN of Indanza on Hwy.45, roadside ditch, 10 i 2009, leg. & det. RWG & NVE [RWG]; PA, Santa Clara: El Partidero, Río Napowatershed ( -1.2833, -77.9000, 800m), 4 ix 1935, leg. WCM, det. RWG, from C.H. Kennedy[UMMZ]; Mera; 5 ♂♂, Merabetween Meraand Shell, Los Copalestrails and forest N of Río Metolo( -1.4886, -78.0708, 1021m), 20 ix 2005, leg. det. WFM [FSCA, QZAC, USNM]. Argia limitataNavás, 1924. Publ. Recs. -NA: * Garrison & von Ellenrieder (2018); OR:*Garrison & von Ellenrie- der (2018); PA: * Garrison & von Ellenrieder (2018); SU:* Garrison & von Ellenrieder (2018); ZC: * Garrison & von Ellenrieder (2018). Also PER: (H08), (P11), (H14). Addl. Data: NA, El Chaco:[FSCA]; Tena:[JJD, KJT, RWG, SWD, SWD]; OR, Loreto:[JTJ, KJT]; PA, Mera:[JJD]; Pastaza: [FAL, JJD, KJT, RWG, SWD]; SU, Cascales: [SWD]; Lago Agrio:[RWG]. Note: Often referred to as “near” A. risi,then in the mid 1990’s it was referred to as “ A. dunklei,” a manuscript name since it was thought to be a new species. Many specimens in various museums around the world are labeled “ A. dunkleims.” Another manuscript name that was proposed more recently was “ A. upano;” Rosser Garrison (pers. comm., 2017) has determined that most of these refer to A. limitata ,a very common, mid-elevation east slope species. A few specimens labeled “ A. upanoms.” turned out to be A. philipi Garrison & von Ellenrieder, 2018. Argia loutoni Garrison & von Ellenrieder, 2015. Publ. Recs. ECU:(P16), Garrison & von Ellenrieder (2018); SU, Shushufindi: * Garrison & von Ellenrieder (2015). Also BRA, COL, PER, VEN: Garrison & von Ellenrieder (2015). Addl. Data. SU, Shushufindi: 1 ♀, small stream about 3 kmW of Shushufindi( -0.1647, -76.6878, 286m), 28 x 2013, leg. KJT, det. RWG [KJT]. Argia mauffrayi Garrison & Ramón Cabrera, 2019. Publ. Recs. ECU: PC, San Miguel de Los Bancos Cantón: Garrison & Ramón Cabrera (2019— Holotype ♂, Recinto Milpe, Milpe Bird Sanctuary, Mirador Uno Trail ( 0.0333, -78.8661, 1101m), 4 ix 2018, leg. WFM [USFQ]. ES, Quinindé: Paratype ♂, Reserva Canande ( 0.5203, -79.0866, 470m), 15 v 2012, leg. Jessica Ware & Melissa Sanchez [RWG]). Note: Known only from these two localities at present. Argia medullarisHagen inSelys, 1865. Publ. Recs. ECU:(H08), (P11), (H14); IM: Garrison & von Ellenrieder (2018); LJ: Garrison & von Ellenrieder (2018); MN: Garrison & von Ellenrieder (2018); MS: Garrison & von Ellenrieder (2018); NA:* Tennessen (1996b), Daigle (1996), Garrison & von Ellenrieder (2018); PA: Garrison & von Ellenrieder (2018); PC: Garrison & von Ellenrieder (2018); SD: Garrison & von Ellenrieder (2018); TU: Garrison & von Ellenrieder (2018). Also CEN, COL, VEN: (H08), (P11), Garrison & von Ellenrieder (2018). Addl. Data. CT, Pangua:[FSCA, KJT]; Sigchos:[FSCA]; IM, Cotacachi:[FSCA, KJT, USFQ]; San Miguel Urcuq:[JTJ]; LJ, Catamayo:[JTJ]; MS, Morona:[FSCA, JJD, RWG]; NA,Archidona: [FSCA, JTJ, KJT, SWD]; Quijos:[FAL, FSCA, JJD, KJT, RWG, TWD, USNM]; PC, Mejía:[FSCA, JJD, USNM]; San Miguel Bancos:[FSCA, USNM]; SD:[FAL, KJT]; SU, Gonzalo Pizarro:[FSCA]; TU, Baños:[RWG]; ZC, Zamora:[JTJ]. Argia nigrior Calvert, 1909. Publ. Recs. ECU: (H08), (P11), (H14); Garrison & von Ellenrieder (2018); ZC:* Tennessen & Johnson (2008). Also PER: (H08), (P11). Note: Rosser Garrison (pers. comm., 2017) wrote “A pair labeled Argiasp. “D”: ECUADOR: Zamora-Chinchipe Province, 5.6 kmeast of Zamora, 3.7 kmsouth of Timbara [“small river with boulders, tributary of Río Zamora, end of gravel road, trickle on roadside and along trail bordering river, secondary forest and pasture”], 4.0658, -78.9044; 4 iv 2008; collected by Dave Wagner, William A. Haber, Michael Thomas, Fred Morrison & Ronald Vargas matches well the typeof A. nigriorfrom Bolivia and have so labeled it this species.” Argia oculataHagen inSelys, 1865. Publ. Recs. ECU:*Calvert (1902), Calvert (1902b), Ris (1918), Soukop (1954), Paulson (1983), Tsuda (1991), Steinmann (1997a), (L06), (H08), (P11), (H14); CR: Campos (1922); CB: Calvert (1907), Campos (1922); EO: Campos (1927); LR: Campos (1922); AZ, BO, CT, ES, LJ, LR, PC, SD: Garrison & von Ellenrieder (2015), MS: Garrison & von Ellenrieder (2007); NA:Daigle (1996), Garrison & von Ellenrieder (2007); OR: Tennessen (1996), Garrison & von Ellenrieder (2007, 2011); PA: Garrison & von Ellenrieder (2007); SU:Garrison & von Ellenrieder (2007). Also BRA, COL, PER—as A. difficilis, TTO as A. difficilis, VEN: (L06), (H08), (P11); plus CEN, PER, TTO: (H08); and BOL, GUF, GUY, SUR: (P16). Addl. Data. AZ, Santa Isabel:[FSCA, KJT, RWG, SWD]; BO, Caluma:[FSCA, KJT, JTJ]; CB, Camada:[RWG]; CN, Cañar:[FSCA]; La Troncal:[JJD, KJT, SWD]; San Miguel:[FSCA, KJT]; CT, Pangua:[FSCA, JTJ, KJT]; Sigchos:[FSCA]; EO, Arenillas:[FSCA]; Piñas:[FSCA, JTJ, KJT]; Santa Rosa:[JTJ, KJT]; ES, San Lorenzo:[FSCA, JJD, KJT, RWG, USNM]; GY, Naranjal:[FSCA, JTJ, KJT]; IM, Ibarra: [FSCA]; LJ, Catamayo:[FSCA, JTJ, KJT]; Loja:[FSCA, JTJ, KJT, UTIC]; LR, Buena Fé:[SWD]; Quevedo:[JTJ, KJT]; Montalvo:[KJT]; MN, Flavio Alfaro:[KJT]; Pedernales:[JTJ, KJT]; PC, Mejía:[FSCA]; San Miguel Bancos:[FSCA, KJT, RWG, USFQ]; SD:[FSCA, JJD, JTJ, KJT, RWG, SWD, USNM]. Note: Many specimens of A. oculatain various collections are labeled “ Argiacf. oculata,” “ A. oculata?,” “ A. difficilis,” “ A. forficula” and “ A. elipitca”. Garrison & von Ellenrieder (2015)defined A. oculataand discussed its varability. It is very common in Ecuadoron the Pacific slope. Argia philipi Garrison & von Ellenrieder, 2018. Publ. Recs. MS:* Garrison & von Ellenrieder (2018); NA:* Garrison & von Ellenrieder (2018); OR:* Garrison & von Ellenrieder (2018); SU:* Garrison & von Ellenrieder (2018). Also BOL, COL, PER: Garrison & von Ellenrieder (2018). Addl. Data. OR, Loreto: 2 ♂♂, 1 cop. pair, forest stream about 12 kmNE of Loreto( -0.6217, -77.2600, 365m), 12 x 2009, leg. KJT [KJT]; same data but 1 ♂, small streams and forest trails 12 kmE of Loreto, 2+ km S of Hwy.45, leg. JTJ, det. RWG [JTJ]; PA, Mera: 1 ♀, near Shell, trails, forest, and Río Metolo, at Los Copales( -1.4919, - 78.0725, 1070m), 20 ix 2005, leg. KJT, det. RWG [KJT]; SU, Shushufindi: 1 ♂, stream about 3 kmNW of Shushu- findi ( -0.1664, -76.6917, 290m), 13 x 2009, leg. KJT, det. RWG [KJT]. Argia popolucaCalvert, 1902. Publ. Recs. ECU: (H14), * Garrison & von Ellenrieder (2018). Also COL (as A. variata): (H08), (P11), (P&P). Addl. Data. BO, Caluma: 1 ♂, tributary of rocky stream 5 kmE of Caluma, Gauranda Road ( -1.5942 -79.2308, 480m), 5 Mar 2010, leg. JJD, det. RWG [JJD]; ES, Quinindé: 1 ♂, Río Canande Reserve, 35 kmNE of Quininde, tributary of Río Hoja Blanca, small gravel stream in forest ( 0.5264, -79.2122, 345m), 20 x 2012, leg. WAH, det. RWG, data provided by RWG (2018) [WAH]; PC, Quito: 2 ♂♂, 2 ♀♀, Río Tulambi, Forestry StationMaquipucuna (about 5 kmE of Nanegalito) ( -0.1169, -78.6297, 1344m), 5 ix 1990, leg. OSF, det. RWG [RWG, USNM]; 1 ♂, 2 kmNE of Mindo, Septimo Paraiso Lodge, small stream in forest ( 0.0308, -78.7633, 1555m), leg. WAH, det. RWG, data provided by RWG (2018) [WAH]. Argia pullaHagen inSelys, 1865. Publ. Recs. ECU:(L06), von Ellenrieder & Muzón (2007), (H08), (P11), (H14), Garrison & von Ellenrieder (2018); NA: *Daigle (1996). Also BRA, CEN, COL, PER, TTO, VEN: (L06), (H08), (P11); plus ARG: (H08); and SUR: (P16). Addl. Data. ES, Esmeraldas:[DRP]; MN, Chone:[JJD, JTJ]; Flavio Alfaro:[JTJ]; Junin:[JTJ, KJT]; MS, Santiago:[SWD]; NA, El Chaco:[SWD]; Tena:[FSCA, JTJ, KJT, SWD, TWD, USNM]; OR, Francisco de Orellana: [FSCA, JJD, KJT, RWG, TWD, SWD, USNM]; Loreto:[JTJ]; SD:[FSCA, JJD, KJT, USNM]; SU, Lago Agrio:[SWD, USNM]; Shushufindi:[FSCA, KJT, SWD, USNM]. Argia schneideri Garrison & von Ellenrieder, 2017. Publ. Recs. MS:* Garrison & von Ellenrieder (2017); NA:* Garrison & von Ellenrieder (2017— Holotype: Las Palmas, on Río Anzu in Río Napowatershed, 11 xii 1936, leg. WCM, [UMMZ]; PA:* Garrison & von Ellenrieder (2017); TU:* Garrison & von Ellenrieder (2017). Also COL: Bota-Sierra et al.(2018). Addl. Data. OR, Loreto: 1 cop. pair, forest stream about 12 kmNE of Loreto( -0.6217, -77.2600, 365m), 12 x 2009, leg. & det. KJT [KJT]; 1 ♂, same data but 15 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, small streams and forest trails 12 kmE of Loreto, 2+ km S of Hwy.45 ( -0.6217, -77.2600, 365m), 15 x 2009, leg. & det. JTJ [JTJ]; SU, Shushufindi: 1 ♂, trails, small streams 3 kmW of Shushufindi, 2+ km N off Hwy. 26 at km marker 17 ( -0.1664, -76.6917, 290 m), 13 x 2009, leg. & det. JTJ [JTJ]. Note: The manuscript name “ camura” was originally applied to this species. Argia selsyi Garrison & von Ellenrieder, 2018. Publ. Recs. MS:* Garrison & von Ellenrieder (2018); NA:* Garrison & von Ellenrieder (2018— Holotype: [ Tena]:Jatun Yacu, NapoRiver watershed [ -1.0166, -77.8333, 700m], 18 iv 1935, leg. WCM [UMMZ]); OR:* Garrison & von Ellenrieder (2018); PA:* Garrison & von Ellenrieder (2018); TU:* Garrison & von Ellenrieder (2018). Note: Endemic but common when found. Argia talamanca? Calvert, 1907. Publ. Recs. ECU:* Tennessen (1998), (P11), (H14). Also CEN, COL, VEN: (H08), (P11). Addl. Data. BO, San Miguel: 1 ♂, Balzapamba (-1.77, - 79.18W, 732m), ix 1935, leg. R. Campos, det. L. Gloyd [FSCA]; CN, La Troncal: 2 ♂♂, small stream and waterfall, 25 kmE of Cochancay, Hwy. 70, ( -2.4692, -79.1697, 1540m), 3 xi 1997, leg. & det. JJD [FSCA]; 4 ♂♂, same data but leg. & det. KJT [FSCA]; 5 ♂♂, 2 ♀♀, same data but leg. & det. TWD [RWG, TWD, FSCA]; 3 ♂♂, same data but leg. & det. WFM [FSCA, LSAM, USNM]; CT, Sigchos: 3 ♂♂, San Francisco de las Pampas Otonga Reserve (-0.42, -79.0, 1700m), 14–16 ii 1997, leg. & det. Elicio Tapia, [FSCA, QZAC]; ES, San Lorenzo: 1 ♂, 5 kmNW of Lita, seeps along RR tracks, 5 ii 1997, leg. & det. JJD [FSCA]; MN, Jipijapa: 3 ♂♂, 3 ♀♀, Puerto Lopez “ 20 kmto Salinas” (-1.57, -80.82, 61m), 5 iv 97, leg. & det. Elicio Tapia [FSCA, QZAC]; PC, Mejía: 1 ♂, Manuel Cornejo 2 kmS of Tandapi, waterfalls (-0.43, -78.78, 1737m), 21 x 1980, leg. & det. MJW [FSCA]; 1 ♂, Manual Cornejo E near Tandapi, “east of Santo Domingo de los Colorados” ( -0.4167, -78.7833, 1585m), 21 x 1980, leg. & det. MJW [FSCA]; 7 ♂♂, same data but 24 x 1980. [FSCA]; TU, Baños: 1 ♂, La Ventana, upper Río Pastazawatershed, ix 1935, leg. & det. F Campos R, [FSCA]. Note: South American specimens may represent a new undescribed species, since the true A. talamancamay occur only in Central America (R. Garrison, pers. comm., 2018). Very similar to A variegata. Argia talamancaappears to be a west slope species whereas A. variegatais mainly an east slope species. Argia tennesseni Garrison & von Ellenrieder, 2018. Publ. Recs. NA: * Garrison & von Ellenrieder (2018); OR:* Garrison & von Ellenrieder (2018— Holotype: [ Loreto]: stream 8.5 kmE of Loreto[ -0.6183, -77.2950, 400m], 14 xi 1997, leg. KJT [FSCA]; ♀ Paratype: stream on road to Waorani Reserve, 6.2 kmsouth of main Yasuni Road [ - 0.7120, -76.4744, 230m], 10 vi 1995, leg. KJT [FSCA]; SU:* Garrison & von Ellenrieder (2018). Also BRA, PER: Garrison & von Ellenrieder (2018). Addl. Data. OR, Franciscode Orellana: 3 ♂♂, 1 ♀, stream, road to Waorani Reserve, 6.2 kmS of Yasuni Sci. Res. Sta., 10 vi 1995, leg. KJT [KJT, RWG]; 1 ♀, P.N.Y., on road to Waorani Reserve, 1.8 kmW of WaoraniRe- serve Road, on Markus Oil Co. RoadPC 29-30, small stream ( -0.7000, -76.4667, 200m), leg. WFM, det. RWG & NVE [RWG]; 1 ♂, 2 ♀♀, T. B.S.( -0.6333, -76.1500, 220m), 20 ix 2000, leg. Kelly Jackson& D. Inward, det. RWG [BMNH]; 1 ♀, T. B.S., USFQ, stream crossing Matapalotrail at marker 450 ( -0.6369, -76.1458, 245m), 14 i 2009, leg. & det. RWG & NVE [RWG]; 1 ♂, 4 ♀♀, T. B.S., USFQ, stream crossing Chorongotrail at marker 225 ( -0.6350, -76.1503, 220m), 16 i 2009, leg. & det. RWG & NVE [RWG]; 1 ♂, 4 ♀♀, same data but 18 i 2009[RWG]; 2 ♀♀, T. B.S., USFQ, black water stream crossing Guacamayotrail at marker 2050 ( -0.6383, -76.1492, 220m), 17 i 2009, leg. & det. RWG & NVE [RWG]; 2 ♀♀, T. B.S., USFQ, Guacamayotrail between markers 450–2100 ( -0.6350, - 76.1533, 237m), 22 i 2009, leg. & det. RWG & NVE [RWG]; Loreto: 3 ♂♂, small streams and forest trails 12 kmE of Loreto, 2+ km S of Hwy.45 ( -0.6271, -77.2600, 365m), 12 x 2009, leg. JTJ [JTJ]; 1 ♂, 2 ♀♀, 1 reared, same data but leg. KJT [KJT]. Argia translataHagen inSelys, 1865. Publ. Recs. ECU: (H08), (P11). CR, GY, LR: * Campos (1922), EO: Campos (1927). Also ARG, CEN, COL, GUF, PER, TTO, VEN: (H08), (P11); plus BOL, GUY, SUR: (P16). Addl. Data. EO, Piñas:[FSCA, JTJ]; Santa Rosa:[KJT]; GU, Balao:[FSCA, KJT]; Naranjal:[FSCA]; LR, Buena Fé:[DRP, FSCA, RWG, SWD]; Quevedo:[FSCA, KJT]; Vantanas:[FSCA, KJT]; MN, Chone:[USNM]; Manta:[RWG, USNM]; NA, Tena:[FSCA, JJD, RWG, SWD, TWD, USNM]; SD: [RWG]; SU, Gonzalo Pizarro:[USNM]. Argia variegata Förster, 1914. Publ. Recs. ECU:* Förster (1914- Description, no specific typelocality), Davies & Tobin (1984), Tsuda (1986, 1991), Bridges (1993), Steinmann (1997a), Garrison et al.(2003), (H08), (P11), (H14); MS: Tennessen (2006); NA: Daigle (1996), Tennessen (1998); ZC: Tennessen & Johnson (2008). Also COL, PER: (H08), (P11). Addl. Data. MS, Huamboya:[FAL, FSCA, JJD, KJT, TWD, USNM]; Morona:[FSCA]; NA, Archidona:[FAL, FSCA, JJD, JTJ, KJT, SWD, TWD, USNM]; Carlos Julio Arsemena Tola:[RWG]; El Chaco:[SWD]; Quijos:[JJD]; Tena:[SWD]; LJ, Loja[JTA ]; OR, Loreto:[DRP, FSCA, JJD, KJT, RWG, TWD, USNM]; PA, Mera:[FSCA, JJD, KJT, RWG, UMMZ, USNM]; PC, San Miguel de Los Bancos:[FSCA, USFQ]; SU, Gonzalo Pizarro:[FSCA]; TU, Baños:[CSCA, FSCA, RWG, UMMZ, USNM]; ZC, Zamora:[FSCA, JTJ, KJT, USNM]. Note: This species is closely related to A. talamancaand may be found to be a junior synonym of it; the issue is currently under review by Rosser Garrison. Argiaunresolved:There are numerous specimens and records for Argiawhich we have not listed here. Many belong to the Argiagerhardi-kokama-nigriorgroup that still needs to be resolved. Museum specimens are often labeled by alphabetical designation (e.g. “ Argiasp. g”). Garrison & von Ellenrieder have been working on this complex for some time and hopefully someday will provide a solution for these unresolved Argia.