Trichotichnus ( Trichotichnus ) subangulatus Ito, 2021b: 71 New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies Kataev, Boris M. Hongbin, Liang Wrase, David W. Zootaxa 2022 2022-06-30 5159 3 301 353 Ito, 2021 Ito 2021 [151,700,151,178] Insecta Carabidae Trichotichnus true Animalia Coleoptera 32 333 Arthropoda species subangulatus  ( Figs. 55, 74, 77–83& 87)      Trichotichnus( Trichotichnus) subangulatus Ito, 2021b: 71.   Material examined.   China. YUNNAN: 1 male, 3 females, Gongshan, Cikai Town, 0.5 kmof Kongdang, 27.88111°N 98.34063°E,  1500 m,  25.X.2004night, D. Kavanaughleg. ( IOZ, CAS);  3 males, 2 females, Gongshan, Dulongjiang,  2.8 kmS Longyuan Vill., 28.00905°N 98.32204°E,  1660 m,  31.X.2004 Night, D. Kavanaugh, D.Z. Dongleg. ( IOZ, CAS);  1 female, Gongshan, Dulongjiang, Maku, Wood, vegetation, 27.68875°N 98.31303°E,  1677 m,  3.IX.2006, Liu Yeleg. ( IOZ);  2 males, 1 female, Baoshan Pref., Gaoligong Shan,  65 kmNNE Tengchong,  1750 m, 25°35'20"N 98°40'21"E, sec. mixed forest, overgrown stone debris, litter and moss sifted,  27.VIII.2009, D.W. Wraseleg. [10] (cWR); 1 f, Santaishan, 30 kmSW Mangshi,  1200 m,  18.V.1955, U Loleg. ( ZIN);  1 male, 1 female, Jinping: Shilicun Forest Station,  1960 m, 22°46'28.4"N 103°17'49.3"E,  16.XII.2003, B. Kataevleg. [5] ( ZIN).  GUANGXI: 1 male, 1 female, Longsheng, Huaping Nature Reserve, Hongtan, light trap,  4.X.2007, night, Liu Chunxiangleg. ( IOZ).    Vietnam. LAO CAI: 2 males,  40 kmWNW of Lao Cai, env. Y Ty, Bat Hat Natn. Park,  1600–1900 m, 22°37′– 37.6′N 103°37.3′–28.3E,  16–21.X.2018, D. Fedorenkoleg. ( SIEE);  1 female, Sa Pa, at light,  23.IV.–6.V.2013, A. Prosvirovleg. ( SIEE).  CAO BANG: 1 male, vic. Vin Den, Nui Pia Oac Nature Res., 22°33′53′′N 105°52′53′′E,  900–1300 m,  06–10.V.2013, A. Weigelleg. ( NME);  1 female, 40 kmW of Cao Bang Phia Oac Mt., h ~  1650–1700 m, deciduous forest,  3–11.X.2018, D. Fedorenkoleg. ( SIEE).  Re-description. Body length 7.7–9.6 mm. Habitus as in Fig. 74.   FIGURES 74–76.  Trichotichnusspp., habitus. 74,  T. subangulatusIto, 2021(NNE Tengchong); 75. T.?  subangulatusIto, 2021(S Longyuan); 76,  T. noctuabundus Habu, 1954(Vietnam). Scale bar = 1.0 mm. Morphological character states are as described for  T. schmidti  sp. n., but differing as follows. Ventral side of body blackish brown, with medial portions in most specimens paler. Labrum markedly concave anteriorly. Left mandible in frontal view wide and truncate at tip ( Fig. 55). Pronotal sides either rounded along entire length, more widely in basal half than in apical half, or very shallowly sinuate before basal angles; apical margin markedly emarginate; basal angle obtusangular, sharp at apex, in most specimens with a very small obtuse denticle at tip. Last visible abdominal sternite ( VII) in male rounded or subtruncate at apex. Median lobe of aedeagus ( Figs. 77–81) in lateral view arcuate, with apical third rather strongly, somewhat angularly curved ventrally; ventral margin in basal two thirds almost straight; apex thin, not prominent dorsally or ventrally; in dorsal view median lobe comparatively wide, only slightly narrowed apically, with terminal lamella ( Fig. 77) very wide and short, roundly narrowed posteriorly. Ventral side edged laterally in apical part and depressed between margins. Apical orifice in dorsal position, wide apically. Internal sac with narrow spiny patches along sides apically.   FIGURES 77–86.  Trichotichnus subangulatusIto, 2021(77–79, 82 & 83, Yunnan; 80 & 81, Vietnam; 84–86, T.?  subangulatus, Yunnan, Elidang Vill.); 77, 84, terminal lamella of aedeagus; 78–81, 85 & 86, median lobe of aedeagus; 82, gonocoxite; 83, laterotergite, gonosubcoxite, and gonocoxite. Dorsal (77, 79, 81, 84 & 86), lateral (78, 80, 82 & 85) and ventral (83) views. Scale bars: A = 0.5 mm(77, 82–84), B = 1.0 mm (78–81, 85 & 86). Female genitalia ( Figs. 82 & 83): gonocoxite moderately wide in ventral view, about 0.6 times as long as gonosubcoxite, with wide base. Proportions ( 6 malesand 8 femalesmeasured): HWmax/PWmax 0.68–0.74; HWmin/PWmax 0.51–0.59; HWmax/HWmin 1.24–1.40; PWmax/PL 1.45–1.63; PWmax/PWmin 1.16–1.27; EL/EW 1.47–1.62; EL/PL 2.98–3.29; EW/PWmax 1.24–1.38.  Comparison.In addition to the distinctive features of the male genitalia, this species can be distinguished from  T. doiinthanonensis,  T. schmidti  sp. n.,  T. fedorenkoi  sp. n.and  T. noctuabundusby having labrum more deeply concave anteriorly, left mandible in frontal view wider and truncate at apex, and pronotum in most specimens with small denticle at apex.   Distribution( Fig. 87). Described from North Vietnam( typelocality: Sapa). Known from North Vietnam( Lao Caiand Cao Bangprovinces) and China(northwestern and southern Yunnanand Guangxi); it is the first record for China. The species occurs at altitudes of about 1400–2000 m.   Remarks.The identity of the following four malesis obscure and requires further study.     China. Yunnan: 1 male, Gongshan, Dulongjiang, Elidang Village, beach 28.00287°N, 98.32145°E,  1640 m,  2004.11.3 Night D. Kavanaugh, D.Z. Dong ( IOZ);  1 male, Gongshan, Dulongjiang,  2.8 kmS Longyuan Vill., 28.00905°N 98.32204°E,  1660 m,  31.X.2004, D. Kavanaugh, D.Z. Dongleg. ( CAS);  1 male, Gongshan, Dulongjiang,  2.3 kmS Longyuan, 28.00532°N 98.32145°E,  1685 m,  2.XI.2004, D. Kavanaughleg. ( ZIN).    Vietnam. Lao Cai: 1 male, 6 kmNW Sa Pa, 22°21′39′′N 103°47′19′′E,  1810 m, pasture margin,  7.VIII.2013, V. Assing[8+2] ( MFNB). These males differ from other examined males in the median lobe of the aedeagus ( Figs. 84–86) which in lateral view is less strongly curved, with the apical third more evenly bent ventrally. In other characters including the armature of the internal sac (in all examined specimens, the internal sac is partly everted), they are not distinguishable from other specimens of  T. subangulatus( Fig. 75). All four maleswere collected together with typical specimens of this species ( Fig. 88). The presence of several specimens with similar characteristics of the median lobe indicates that they can belong to a separate sympatric species, but additional material and more detail studies are needed to completely determine their taxonomic status. Their proportions (all 4 malesmeasured): HWmax/PWmax 0.71–0.73; HWmin/PWmax 0.54–0.55; HWmax/ HWmin 1.29–1.36; PWmax/PL 1.47–1.53; PWmax/PWmin 1.17–1.24; EL/EW 1.48–1.58; EL/PL 3.09–3.13; EW/ PWmax 1.31–1.39. Body length 8.1–8.9 mm. 3821894302 2004-10-25 IOZ, CAS D. Kavanaugh China Gongshan 1500 27.88111 Kongdang 1 98.34063 Cikai Town 32 333 4 3 1 Yunnan 3821894485 2004-10-31 IOZ, CAS D. Kavanaugh & D. Z. Dong China 1660 28.00905 Dulongjiang 1 98.32204 Gongshan 32 333 5 2 3 Yunnan 3821894406 2006-09-03 IOZ Liu Ye China Gongshan 1677 27.68875 Maku 1 98.31303 Dulongjiang 32 333 1 1 Yunnan 3821894521 1955-05-18 2009-08-27 1955-05-18 ZIN D. W. Wrase China Baoshan Pref. 1750 25.58889 Mangshi 21 98.6725 Gaoligong Shan 32 333 3 1 2 Yunnan 3821894430 [493,1339,542,569] 1955-05-18 2009-08-27 1955-05-18 ZIN U. Lo China Baoshan Pref. 1750 25.58889 Mangshi 21 98.6725 Gaoligong Shan 32 333 3 1 2 Yunnan 3821894515 2003-12-16 ZIN B. Kataev China 1960 22.774555 Shilicun Forest Station 1 103.29703 Jinping 32 333 2 1 1 Yunnan 3821894461 2007-10-04 IOZ Liu Chunxiang China Longsheng Hongtan Huaping Nature Reserve 32 333 2 1 1 Guangxi 3821894468 2018-10-16 2018-10-21 2018-10-16 SIEE Y Ty & D. Fedorenko Vietnam 1750 Bat Hat Natn. Park 28.3 32 333 2 2 Lao Cai 3821894459 2013-04-23 2013-05-06 2013-04-23 SIEE A. Prosvirov Vietnam Sa Pa 32 333 1 1 Lao Cai 3821894541 2013-05-06 2013-05-10 2013-05-06 NME A. Weigel Vietnam 1100 22.564722 Nui Pia Oac Nature Res. 21 105.88139 Vin Den 32 333 1 1 Cao Bang 3821894361 2018-10-03 2018-10-11 2018-10-03 SIEE D. Fedorenko Vietnam 1675 Cao Bang Phia Oac Mt. 32 333 1 1 Cao Bang 3821894514 2004-11-03 IOZ Night D. Kavanaugh, D. Z China Gongshan 1640 28.00287 Elidang Village 1 98.32145 Dulongjiang 34 335 1 1 Yunnan 3821894350 2004-10-31 CAS D. Kavanaugh & D. Z. Dong China 1660 28.00905 Dulongjiang 1 98.32204 Gongshan 34 335 1 1 Yunnan 3821894357 2004-11-02 ZIN D. Kavanaugh China 1685 28.00532 Dulongjiang 1 98.32145 Gongshan 34 335 1 1 Yunnan 3821894400 2013-08-07 MFNB V. Assing Vietnam 1810 22.360834 Sa Pa 21 103.78861 34 335 1 1 Lao Cai