Hippomedon miersi Stebbing, 1888: 631 Anonyx Miersii . Della Valle, 1893: 813 Tmetonyx miersi Stebbing, 1906: 74 Sheard, 1937: 19 Barnard, 1958: 100 Barnard, 1962: 29 Thurston & Allen, 1969: 369 Tryphosella miersi Barnard & Karaman, 1991: 537 Galathella miersi Lowry & Stoddart, 2003: 280 The tryphosine genera Photosella gen. nov. and Tryphosella Bonnier, 1893 (Crustacea: Amphipoda: Lysianassoidea: Lysianassidae: Tryphosinae) in Australian waters 2956 Lowry, J. K. Stoddart, H. E. Zootaxa 2011 2011-07-08 2956 1 1 76 4GT9M (Stebbing, 1888) Stebbing 1888 [151,569,1260,1287] Malacostraca Tryphosidae Photosella Animalia Amphipoda 6 7 Arthropoda species miersi  ( Figs 2–4)      Hippomedon miersi Stebbing, 1888: 631, pl. 10.    Anonyx Miersii.—  Della Valle, 1893: 813.    Tmetonyx miersi. ―  Stebbing, 1906: 74(key), 75. —  Sheard, 1937: 19. —J.L.  Barnard, 1958: 100. —J.L.  Barnard, 1962: 29, 30 (key). —  Thurston & Allen, 1969: 369.    Tryphosella miersi. —  Barnard & Karaman, 1991: 537.    Galathella miersi. —  Lowry & Stoddart, 2003: 280.     Typelocality. East of Moncoeur Island, Bass Strait, Australia, 39°10'30''S 146°37'00''E,  69.5 mdepth.  Material examined.   New South Wales: 43 specimens, AM P.58468, north-east of Coffs Harbour, 30°14'38"S 153°27'41"E,  100 m, baited trap,  12–13 August 1993, P.B. Berents, R. T.  Springthorpe& W. Vader, MV   Cheryl Lee, stn NSW-884;  3 specimens, AM P.52619, north-east of Coffs Harbour, 30°15'45"S 153°21'59"E,  100 m, baited trap,  12–13 August 1993, P.B. Berents, R. T.  Springthorpe& W. Vader, MV   Cheryl Lee, stn NSW-886;  11 specimens, AM P.52651, same data, stn NSW-887;  17 specimens, AM P.55969, north-east of Coffs Harbour, 30°15'56"S 153°21'54"E,  100 m, baited trap,  11–12 August 1993, P.B. Berents, R. T.  Springthorpe& W. Vader, MV   Cheryl Lee, stn NSW-870;  1 specimen, AMP.57620 and  14 specimens, AM P.57621, same data, stn NSW-872;  6 specimens, AM P.48195, north-east of Coffs Harbour, 30°15'56"S 153°21'54"E,  100 m, baited trap,  8–9 September 1994, J.K. Lowry& K. Dempsey, MV   Carrie Anne, stn NSW-984;  20 specimens, AM P.48184, same data, stn NSW-985;  3 specimens, AM P.48428, same data,  9–10 September 1994, stn NSW-1006;  1 female, AM P.70483, south-east of Port Stephens, 32°52’S 152°32’E,  145 m, dredge,  6 December 1978, FRV  Kapala, stn K78-26-07;  1 male, 13.2 mm, AM P.70484, east of Long Reef Point, 33°43’S 151°46’Eto 33°44’S 151°46’E,  174 m, epibenthic sled,  19 December 1985, 2230–2300 hrs, J.K. Lowry& R. T.  Springthorpe, FRV  Kapala, stn K85-21-08;  1 male, AM P. 1825123 kmeast of Cape Banks, 33°58'24"S 151°29'E,  150 m, orange-peel grab,  18 June 1962, CSIRO  Fisheries, HMAS  Gascoyne, stn G2/62/56.   Victoria: 1 specimen, MV J62295, 31 km south-south-west of Cape Otway, western Bass Strait, 39°08'S 143°24'E,  77 m, medium sand, Smith-McIntyre grab,  8 October 1980, G.C.B. Poore, HMAS  Kimbla, stn BSS-56G;  1 specimen, MV J62294, 32 km south-south-west of Cape Otway, western Bass Strait, 39°09'S 143°26'E,  85 m, coarse carbonate sand, dredge,  8 October 1980, G.C.B. Poore, HMAS  Kimbla, stn BSS-55D;  1 specimen, MV J62299, 51 km south-south-west of Cape Otway, western Bass Strait, 39°18'S 143°03'E,  113 m, coarse carbonate sand, Smith-McIntyre grab,  10 October 1980, G.C.B. Poore, HMAS  Kimbla, stn BSS-69G;  1 specimen, MV J62296, 26 km south-west of Cape Otway, western Bass Strait, 39°01.0'S 143°22.1'E,  84 m, medium sand, dredge,  31 January 1981, M. Gomon et al., FRV  Hai Kung, stn BSS-120D;  3 specimens, MV J62297, 38 km south-west of Cape Paterson, central Bass Strait, 38°56.4'S14516.6'E,  70 m, fine sand, epibenthic sled,  12 November 1981, R.  Wilson, RV  Tangaroa, stn BSS-155S;  1 specimen, MV J62298, 44 km north-north-east of Cape Wickham, King Island, central Bass Strait, 39°16.0'S 144°05.4'E,  82 m, sandy shell, rock dredge,  23 November 1981, R.  Wilson, RV  Tangaroa, stn BSS-204RD.   Tasmania: 1 male, AMP.70485,  1 female, AMP.70486 and  4 specimens, AM P.70487, about  80 moutside Hannants Bight, north side of Cape Sorell, 42°11.5'S 145°11'E,  18 m, baited trapon sand and detritus,  26–27 April 1991, J.K. Lowry& S.J. Keable, stn TAS-277;  1 specimen, AM P.70488, about  400 moutside Hannants Bighton a line toward large bluff of Ocean Beach, north side of Cape Sorell, 42°11.4'S 145°11.1'E,  30 m, baited trapon sand,  26–27 April 1991, J.K. Lowry& S.J. Keable, stn TAS-279;  5 specimens, AM P.70489, about  50 moffshore and just east of Hannants Bighton a line toward large bluff of Ocean Beach, north side of Cape Sorell, 42°11.5'S 145°11.4'E,  18 m, baited trapon sand,  26–27 April 1991, J.K. Lowry& S.J. Keable, stn TAS-281;  6 specimens, AM P.45776, east of Fortescue Bay, north of Hippolyte Rocks, 43°06'42"S 148°03'27"E,  100 m, baited trap,  16–17 April 1993, J.K. Lowry& P. Freewater, MV   Tasmanian Enterprise, stn TAS-353;  68 specimens, AM P.51085, same data, stn TAS-354;  6 specimens, AMP.58244 and  1 specimen, AM P.70490, same data,  8–9 April 1994, J.K. Lowry& K. Dempsey, MV   Martrudan, stn TAS-389;  38 specimens, AM P.51090, same data, stn TAS-390;  40 specimens, AM P.51191, same data, stn TAS-391;  38 specimens, AM P.51332, same data,  9–10 April 1994, stn TAS-408;  20 specimens, AM P.51318, same data, stn TAS- 409;  1 specimen, AM P.45747, mouth of Fortescue Bay, 43°07'46"S 145°59'28"E,  100 m, baited trap,  16–17 April 1993, J.K. Lowry& P. Freewater, MV   Tasmanian Enterprise, stn TAS-350;  2 specimens, AM P.51314, same data,  9–10 April 1994, J.K. Lowry& K. Dempsey, MV   Martrudan, stn TAS-406.   South Australia: 1 female, AM P.70491, Marino, Gulf St Vincent, approx. 35°03'S 138°31'E, 1910, W.H. Baker.   Western Australia: 1 specimen, P.18195, east of Albany, 34°32'S 121°31'E, orange-peel grab,  9 July 1962, CSIRO  Fisheries, HMAS  Gascoyne, stn G2/62/109;  1 specimen, AM P.70492, north-west of Port Hedland, North West Shelf, 19°05.3'S 118°54.0'Eto 19°05.1'S 118°54.2'E,  82–83 m, epibenthic sled,  15 February 1983, FRV  Soela, CSIROstn NWS 01.B4.S;  1 female, AM P.70493, north-west of Port Hedland, North West Shelf, 19°05.4'S 118°53.9'Eto 19°05.3'S 118°54.0'E,  80 m, beam trawl,  30 October 1983, FRV  Soela, CSIROstn NWS 05.B4.BT;  3 specimens, AM P.70494, north-west of Port Hedland, North West Shelf, 19°04.8'S 118°50.7'Eto 19°05.2'S 118°50.4'E,  81 m, epibenthic sled,  30 October 1983, FRV  Soela, CSIROstn NWS 05.B5.S;  4 specimens, AM P.70495, north-west of Port Hedland, North West Shelf, 19°04.1'S 118°47.8'Eto 19°04.7'S 118°47.2'E,  82 m, epibenthic sled,  30 October 1983, FRV  Soela, CSIROstn NWS 05.B6.S;  2 specimens, AM P.70496, north-west of Port Hedland, North West Shelf, 19°02.0'S 118°47.2'Eto 19°01.7'S 118°47.1'E,  82 m, epibenthic sled,  7 December 1982, FRV  Soela, CSIROstn NWS 06.B6.S   FIGURE 2.  Photosella miersi( Stebbing, 1888). Hannants Bight, north side of Cape Sorell, Tasmania.  Diagnostic description. Headlateral cephalic lobe subtriangular, apically subacute; eyes (fading in alcohol), large, subtriangular with ventral concave lens. Antenna 1flagellum with strong 2-field callynophore, robust setae absent from proximal articles; calceoli absent. Antenna 2peduncular articles 4 and 5 not enlarged; flagellum short, calceoli absent. Epistome/Upper lipepistome less produced than upper lip, straight. Mandiblemolar with asymmetrically reduced column, proximally setose, distally triturating. Maxilla 1setal-tooth 7, left and right symmetrical, cuspidate along most of straight inner margin; palp distal margin with apical robust setae. Maxillipedouter plate with 2 short apical robust setae.  Gnathopod 1subchelate; coxa large (length 1.8 x breadth), subrectangular, distally subovate; basis moderately setose along anterior margin; ischium short (length 1.4 x breadth); carpus long (length 2 x breadth), shorter than propodus, without posterior lobe; propodus margins subparallel, palm moderately acute. Gnathopod 2propodus palm moderately obtuse.  Pereopod 4coxa without distinct lateral ridge.  Pereopod 5coxa without distinct lateral ridge; basis about as long as broad, with photophore.  Epimeron 3posterior margin smooth, posteroventral corner broadly rounded. Urosomite 1with anterodorsal notch and slightly rounded boss. Uropod 3outer ramus article 2 short, with plumose setae on both rami.  Telsondeeply cleft, with dorsal robust setae, with 1 or 2 apical robust setae on each lobe.  Male(sexually dimorphic characters). Antennae 1 and 2calceoli present. Antenna 2peduncular article 4 enlarged.   Remarks.Although Stebbing (1888)did not record or illustrate a photophore on the basis of pereopod 5 of his specimen we feel sure the present material is the same species. It is possible that Stebbing overlooked the structure, as it can be very difficult to discern in material that is de-calcified by preservatives.  Photosella miersiis the only species in the genus without ridging on the coxa of pereopods 4 or 5.   Distribution.South-eastern and southern Australiato Port Hedland in north-western Australia; 18–174 mdepth. [151,1295,1587,1612] Australia 70 -39.175003 East of Moncoeur Island 19 146.61667 Bass Strait 6 7 1 holotype [442,867,1623,1648] AM New South Wales 6 7 43 New South Wales 1993-08-12 1993-08-13 1993-08-12 R, T P. B. Berents 100 -30.243889 Coffs Harbour 20 153.46138 6 7 1 New South Wales [959,1299,1659,1683] MV W. Vader Springthorpe 6 7 1 New South Wales Cheryl Lee 6 7 1 New South Wales [323,526,1695,1719] AM 6 7 3 New South Wales 1993-08-12 1993-08-13 1993-08-12 R, T P. B. Berents 100 -30.2625 Coffs Harbour 20 153.3664 6 7 1 New South Wales [663,1013,1731,1755] MV W. Vader Springthorpe 6 7 1 New South Wales [1022,1320,1731,1756] Cheryl Lee 6 7 1 New South Wales AM 6 7 11 New South Wales [275,657,1767,1791] AM 6 7 1 New South Wales [667,877,1767,1791] AM 6 7 17 New South Wales 1993-08-11 1993-08-12 1993-08-11 R, T P. B. Berents 100 -30.265556 Coffs Harbour 20 153.365 6 7 1 New South Wales [959,1300,1803,1827] MV W. Vader Springthorpe 6 7 1 New South Wales Cheryl Lee 6 7 1 New South Wales [320,509,1839,1863] AM 6 7 1 New South Wales [665,878,1839,1863] AM 6 7 14 New South Wales [887,1277,1839,1863] AM 6 7 1 New South Wales AM 6 7 6 New South Wales 1994-09-08 1994-09-09 1994-09-08 MV J. K. Lowry & K. Dempsey 100 -30.265556 Coffs Harbour 20 153.365 6 7 1 New South Wales [472,786,1912,1936] Carrie Anne 6 7 1 New South Wales [798,1011,1912,1936] AM 6 7 20 New South Wales [1020,1409,1912,1936] AM 6 7 1 New South Wales AM 6 7 3 New South Wales [336,1002,1947,1971] 1994-09-09 1994-09-10 1994-09-09 AM 6 7 1 New South Wales [1014,1171,1947,1971] AM 6 7 1 1 New South Wales 1978-12-06 145 7 8 -32.866665 Port Stephens 1208 152.53334 6 7 1 New South Wales AM 7 8 1 1 New South Wales 1985-12-19 R, T J. K. Lowry 174 -33.733334 Long Reef Point 1203 151.76666 7 8 1 New South Wales [777,1276,1439,1463] Springthorpe 7 8 1 New South Wales [1293,1436,1439,1463] AM 7 8 1 1 New South Wales [151,1321,1476,1500] 1962-06-18 CSIRO 150 -33.973335 Cape Banks 767 151.48334 7 8 1 New South Wales Gascoyne Fisheries 7 8 1 New South Wales J62295 [523,944,1513,1537] MV Victoria 7 8 31 1 Victoria 1980-10-08 G. C. B. Poore 77 -39.133335 western Bass Strait 1171 143.4 Cape Otway 7 8 1 Victoria J62294 [509,837,1588,1612] MV 7 8 32 1 Victoria 1980-10-08 G. C. B. Poore 85 -39.15 western Bass Strait 1170 143.43333 Cape Otway 7 8 1 Victoria J62299 [280,625,1663,1687] MV 7 8 51 1 Victoria 1980-10-10 G. C. B. Poore 113 -39.3 western Bass Strait 1170 143.05 Cape Otway 7 8 1 Victoria J62296 [273,593,1737,1761] MV 7 8 26 1 Victoria 1981-01-31 M. Gomon & Hai Kung 84 -39.016666 western Bass Strait 116 143.36833 Cape Otway 7 8 1 Victoria J62297 MV 7 8 38 3 Victoria 1981-11-12 R 70 -38.94 central Bass Strait Cape Paterson 7 8 1 Victoria [414,544,1849,1873] RV Wilson 7 8 1 Victoria J62298 [846,1170,1849,1873] MV 7 8 44 1 Victoria 1981-11-23 R Cape Wickham 82 -39.266666 central Bass Strait 116 144.09 King Island 7 8 1 Victoria [296,426,1924,1948] RV Wilson 7 8 1 Victoria [748,1004,1924,1948] AM Tasmania 7 8 1 1 Tasmania [1115,1271,1924,1948] AM 7 8 1 1 Tasmania AM 7 8 4 Tasmania 1991-04-26 1991-04-27 1991-04-26 J. K. Lowry & S. J. Keable 18 -42.191666 Cape Sorell 691 145.18333 7 8 1 Tasmania [1243,1436,1999,2023] AM 7 8 1 Tasmania 1991-04-26 1991-04-27 1991-04-26 J. K. Lowry & S. J. Keable 30 -42.19 Cape Sorell 114 145.185 Ocean Beach 8 9 1 Tasmania [151,353,226,250] AM 8 9 5 Tasmania 1991-04-26 1991-04-27 1991-04-26 J. K. Lowry & S. J. Keable Hannants Bight 50 -42.191666 Cape Sorell 114 145.19 Ocean Beach 8 9 1 Tasmania [567,766,301,325] AM 8 9 6 Tasmania 1993-04-16 1993-04-17 1993-04-16 MV J. K. Lowry & P. Freewater 100 -43.111664 Hippolyte Rocks 19 148.05751 Fortescue Bay 8 9 1 Tasmania Tasmanian Enterprise 8 9 1 Tasmania [381,594,375,399] AM 8 9 68 Tasmania [602,980,375,399] AM 8 9 1 Tasmania [993,1192,375,399] AM 8 9 6 Tasmania AM 8 9 1 Tasmania [273,1092,413,438] 1994-04-08 1994-04-09 1994-04-08 MV J. K. Lowry & K. Dempsey 8 9 1 Tasmania [1102,1392,413,437] Martrudan 8 9 1 Tasmania AM 8 9 38 Tasmania [336,712,450,474] AM 8 9 1 Tasmania [725,937,450,474] AM 8 9 40 Tasmania [945,1321,450,474] AM 8 9 1 Tasmania AM 8 9 38 Tasmania [279,864,487,511] 1994-04-09 1994-04-10 1994-04-09 AM 8 9 1 Tasmania [878,1092,487,511] AM 8 9 20 Tasmania AM 8 9 1 Tasmania [208,395,525,549] AM 8 9 1 Tasmania 1993-04-16 1993-04-17 1993-04-16 MV J. K. Lowry & P. Freewater 100 -43.129444 Fortescue Bay 19 145.99112 8 9 1 Tasmania [589,996,562,586] Tasmanian Enterprise 8 9 1 Tasmania [1008,1206,562,586] AM 8 9 2 Tasmania 1994-04-09 1994-04-10 1994-04-09 MV J. K. Lowry & K. Dempsey 8 9 1 Tasmania [762,1054,599,624] Martrudan 8 9 1 Tasmania [1069,1436,599,624] AM South Australia 8 9 1 1 South Australia [151,1068,637,661] W. H. Baker -35.05 Gulf St Vincent 1195 138.51666 Marino 8 9 1 South Australia 1962-07-09 CSIRO -34.533333 Albany 1198 121.51667 8 9 1 Western Australia Gascoyne Fisheries 8 9 1 Western Australia [289,484,711,735] AM 8 9 1 Western Australia 1983-02-15 CSIRO Soela 83 -19.085 North West Shelf 127 118.903336 Port Hedland 8 9 1 Western Australia AM 8 9 1 1 Western Australia 1983-10-30 CSIRO Soela 80 -19.088333 North West Shelf 127 118.9 Port Hedland 8 9 1 Western Australia [1011,1204,823,847] AM 8 9 3 Western Australia 1983-10-30 CSIRO Soela 81 -19.086666 North West Shelf 127 118.84 Port Hedland 8 9 1 Western Australia [687,880,898,922] AM 8 9 4 Western Australia 1983-10-30 CSIRO Soela 82 -19.078333 North West Shelf 127 118.78667 Port Hedland 8 9 1 Western Australia [326,524,973,997] AM 8 9 2 Western Australia 1982-12-07 CSIRO Soela 82 -19.028334 North West Shelf 127 118.785 Port Hedland 8 9 1 Western Australia