The tryphosine genera Photosella gen. nov. and Tryphosella Bonnier, 1893 (Crustacea: Amphipoda: Lysianassoidea: Lysianassidae: Tryphosinae) in Australian waters 2956 Lowry, J. K. Stoddart, H. E. Zootaxa 2011 2011-07-08 2956 1 1 76 59GG3 Lowry & Stoddart, 2011 Lowry & Stoddart 2011 [151,435,1730,1756] Malacostraca Tryphosidae Tryphosella Animalia Amphipoda 65 66 Arthropoda species tuckanarra sp. nov.  ( Figs 47–49)   Type material.  HOLOTYPE, female, ovigerous, 3.1 mm, AMP.70523, Vancouver Peninsula, near Mistaken Island, King George Sound, Western Australia, Australia, 35°04'S 117°56'E,  2 m, grey sponge with crinoids,  13 December 1983, R. T. Springthorpe, stn WA-114.  PARATYPES: 1 immature, AMP.70524, same data as Holotype;  1 juvenile, AMP.70498, Vancouver Peninsula, near Mistaken Island, King George Sound, Western Australia, Australia, 35°04'S 117°56'E,  2 m, sponges,  13 December 1983, J.K. Lowry, stn WA-101;  1 male, 3.1 mm, AMP.70499, same data, stn WA-103;  4 specimens, AMP.70500, same locality, 2 m, soft corals and  Caulerpa, 13 December 1983, J.K. Lowry, stn WA-106;  1 specimen, AMP.70525, same locality,   2 m, sponge covered with short green alga (?  Caulerpa),  13 December 1983, R. T. Springthorpe, stn WA-119; 1 ovigerous female, 3.4 mm, AMP.70526 and 1 ovigerous female, AMP.70527, same locality,   6 m, sea grass,  13 December 1983, R. T. Springthorpe, stn WA-121.  Additional material examined.  Western Australia: 1 ovigerous female and  1 immature, AMP.70528, off Possession Point, King George Sound, 35°02.5'S 117°55'E,  7 m, mixed brown algae, bryozoans, sponge,  R. T. Springthorpe& J.K. Lowry,  14 December 1983, stn WA-133;  2 juveniles, AMP.70529, rocks near Migo Island, Port Harding, Torbay Bay, 35°04'S 117°39'E,  6–7 m, small branched alga with compound tunicate on underside of branches,  15 December 1983,  R. T. Springthorpe& J.K. Lowry, stn WA-152.    Typelocality. Vancouver Peninsular, near Mistaken Island, King George Sound, Western Australia, Australia, 35°04'S 117°56'E,  2 mdepth.   Etymology.Named for the tiny village of Tuckanarra located about about 580 kmnorth-northeast from Perth.  Diagnostic description. Headlateral cephalic lobe subtriangular, apically subacute; eyes apparently absent. Antenna 1peduncular articles 1 and 2 without anterodistal lobe; accessory flagellum not forming cap, terminal article not offset; flagellum with weak 2-field callynophore, robust setae absent from proximal articles; calceoli absent. Antenna 2peduncular article 3 short; articles 4 and 5 not enlarged; flagellum short, calceoli absent. Epistome/Upper lipepistome slightly produced beyond upper lip, narrowly rounded; upper lip slightly produced, straight. Mandiblemolar with asymmetrically reduced column, proximally setose, distally triturating; palp attached about midway, article 3 with proximal A3-seta. Maxilla 1outer plate setal-tooth 7 left and right asymmetrical, left cuspidate along most of sinusoidal inner margin, right cuspidate along most of curved inner margin; palp distal margin with apical robust setae. Maxillipedouter plate with 2 short apical robust setae.  Gnathopod 1subchelate; coxa medium length (length 1.5 x breadth), subtriangular, tapering distally; basis moderately setose along anterior margin; ischium short; carpus long, subequal in length to propodus, without posterior lobe; propodus small, margins subparallel, sparsely setose along posterior margin, palm slightly acute, entire, straight. Gnathopod 2palm transverse. Pereopod 4coxa without distinct lateral ridge, with a well-developed posteroventral lobe. Pereopod 5coxa without distinct lateral ridge, without umbo; basis about as long as broad, not posteroproximally excavate, posterior margin not serrate. Pereopod 7basis posterodistally produced less than halfway along merus, not posterodistally excavate.   FIGURE 48.  Tryphosella tuckanarra  sp. nov.Holotype, female, 3.1 mm, AM P.70523; paratype male, 3.1 mm, AM P.70499; King George Sound, Western Australia. Scales represent 0.1 mm.   FIGURE 49.  Tryphosella tuckanarra  sp. nov.Holotype, female, 3.1 mm, AM P.70523, King George Sound, Western Australia. Scales represent 0.1 mm.  Pleonites 1–3without mid-dorsal carina, not produced dorsodistally. Epimeron 3posterior margin smooth, posteroventral corner subquadrate. Urosomite 1not projecting over urosomite 2, with rounded boss. Uropods 1 and 2rami with distal stout striated robust setae. Uropod 2inner ramus without constriction. Uropod 3peduncle without dorsolateral flange, with 2 short apicomedial robust setae; inner and outer rami well developed, outer ramus article 2 short, without plumose setae on rami.  Telsondeeply cleft, with dorsal robust setae, with 1 apical robust seta on each lobe.  Male(sexually dimorphic characters). Antenna 1calceoli present. Antenna 2flagellum long, calceoli present. Unfortunately a fully-developed male is not present in the collections.   Remarks.  Tryphosella tuckanarrais one of the group of six Australian species with stout striated robust setae on the outer rami of uropods 1 and 2 (see Remarks under  T. fortescue).  Tryphosella tuckanarrais very similar to  T. fortescuein having a transverse palm on gnathopod 2, 2 short apicomedial robust setae on the peduncle of uropod 3 and 2 apical robust setae on each lobe of the telson.  Tryphosella tuckanarrais distinguished from  T. fortescueby the rounded lateral cephalic lobe (subtriangular in  T. fortescue), the less reduced coxa 1 and the presence of 2 apical robust setae on the outer plate of the maxilliped ( 3 in  T. fortescue).   Tryphosella tuckanarrais known only from King George Sound and TorbayBay in shallow-water. It has never been taken in traps.   Distribution.South-western Western Australia, 2–7 mdepth. 1983-12-13 AM, R, T Australia King George Sound 2 -35.066666 Vancouver Peninsula 1195 117.933334 Mistaken Island 65 66 1 Western Australia holotype [707,1432,1903,1927] 1983-12-13 AM Australia King George Sound 2 -35.066666 Vancouver Peninsula 1195 117.933334 Mistaken Island 65 66 1 1 Western Australia paratype 1983-12-13 AM J. K. Lowry Australia King George Sound 2 -35.066666 Vancouver Peninsula 1195 117.933334 Mistaken Island 65 66 1 1 Western Australia paratype 1983-12-13 AM J. K. Lowry Australia King George Sound 2 -35.066666 Vancouver Peninsula 1195 117.933334 Mistaken Island 65 66 1 1 Western Australia paratype [436,906,2011,2035] 1983-12-13 AM J. K. Lowry Australia King George Sound 2 -35.066666 Vancouver Peninsula 1195 117.933334 Mistaken Island 65 66 4 Western Australia paratype [508,953,151,175] 1983-12-13 AM J. K. Lowry Australia King George Sound 2 -35.066666 Vancouver Peninsula 1195 117.933334 Mistaken Island 66 67 1 Western Australia paratype 1983-12-13 R, T, AM Springthorpe Australia King George Sound 2 -35.066666 Vancouver Peninsula 1195 117.933334 Mistaken Island 66 67 1 Western Australia paratype [674,1425,223,248] 1983-12-13 R, T Springthorpe Australia 6 66 67 1 Western Australia paratype AM 7 -35.041668 King George Sound 763 117.916664 Possession Point 66 67 1 1 Western Australia 1983-12-14 R, T J. K. Lowry Springthorpe 66 67 1 Western Australia 1983-12-15 AM Migo Island 7 -35.066666 Torbay Bay 1195 117.65 Port Harding 66 67 2 2 Western Australia [491,1005,404,428] R, T J. K. Lowry Springthorpe 66 67 1 Western Australia Australia King George Sound 2 -35.066666 Vancouver Peninsular 1195 117.933334 Mistaken Island 66 67 1 Western Australia holotype