Revision of Thai Dissomphalus Ashmead, 1893 (Hymenoptera, Bethylidae), with description of twenty four new species
MUGRABI, DANIELE F.
AZEVEDO, CELSO O.
Zootaxa
2013
2013-05-22
3662
1
1
73
Mugrabi & Azevedo
Mugrabi & Azevedo
[151,636,1375,1400]
Insecta
Bethylidae
Dissomphalus
GBIF
Animalia
Hymenoptera
25
26
Arthropoda
species
verus
sp. nov.
( Figs 27–30, 104, 159–161) Description.MALE. Body length 3.65 mm. LFW 2.51 mm. Maximum wing width 1.07 mm. Color. Head and mesosoma black; metasoma dark castaneous. Head. Mandible with five apical teeth. Clypeus with median lobe subtrapezoidal; median tooth rounded; median carina complete apically or nearly so, angled in profile. Ratio of first four antennal segments about 14:7:6:5. Frons strongly coriaceous, punctures large. LH 0.95 × WH, WF 0.63 × WH, WF 1.26 × HE, OOL 1.23 × WOT, DAO 0.35 × WOT, posterior ocelli distant from vertex crest 1 × DAO, frontal angle of ocellar triangle right, VOL 0.68 × HE. Vertex crest weakly concave. Occipital carina not visible in dorsal view. Mesosoma. Pronotal disc strongly coriaceous, anterior margin ecarinate. Notauli deep, complete, widening posterad, converging posterad. Propodeal disc 0.55 × as long as wide; basal triangle rugulose, posterolateral area rugulose; median carina complete; lateral margins strongly convex, parallel; posterior carina well-defined, straight. Lateral of propodeum mostly striolate. Propodeal declivity mostly rugulose-strigate. Metasoma. Tergal process with deep, subtriangular and sublateral pair of depressions; depressions with external lateral margin higher than internal lateral margin, 0.83 × as long as tergite II, diverging posterad, distant on top with short and dorsad tuft of setae ( Figs 27–30). Hypopygium with median stalk 1.15 × as long as hypopygeal plate; median stalk narrow; corner of anterior margin angular; lateral stalk of hypopygium 0.11 × as long as median stalk; posterior margin strongly concave; corner angalute ( Fig. 104). Genitalia( Figs 159–161). Genital ring produced, each half convex in dorsal view. Basiparamere with dorsal surface much wider apicad, much narrower than ventral surface. Paramere with basal area wider than basiparamere apex; apex rounded; apical margin weakly concave in ventral view, not curved; inner margin of ventral surface with triangular spine at median region. Basivolsella setose, with several thin, short and apical setae. Cuspis biramous. Aedeagus 3.72 × as long as wide; dorsal body of aedeagus wider basally, narrowing abruptly apicad, lateral margin strongly convex basally, parallel, not curved dorsad, apex with two pair of outer lobes; dorsal pair 1 × as long as wide, straight, with apex subangulate and dorsad, outer lateral with laminar process, elliptical, with lateral surface wider than dorsal and ventral surfaces; ventral pair tubular, short, slightly arched, laterad, with apical margin rounded and surface smooth, much shorter than dorsal lobe of aedeagus; base of dorsal body of aedeagus strongly sclerotized, weakly concave, with median region bilobate; ventral ramus of aedeagus as high as dorsal body, lateral margin straight, evenly narrow and base as narrow as apex. Apodeme extending beyond genital ring. FEMALE unknown.
Material examined. Holotype, ♂. THAILAND. Chaiyaphum: Pa Hin Ngam NP, Mixeddeciduous forest (Thepana waterfall), 15°38.920’N 101°25.077’E, 614 m, Malaise trap, 1-7.x.2006, Katae Sa-nog& Buakaw Adnafaileg., T656. Paratypes: THAILAND. Malaise trap, Mae Hong Son : 1 ♂ Namtok Mae SurinNP, E/ Huai Fai Korreservoir, 19°20.616’N 97°59.3’E, 27.i-3.ii.2008, Kamkoon;A leg., T3496. Chiang Mai: 1 ♂ Doi Inthanon NP, Vachiratharn Falls, 18°32.311’N 98°36.048’E, 700 m, 30.viii-6.ix.2006, Y. Areeluckleg., T236. Loei: Phu Ruea NP, 1 ♂ Subhnonghin, 17°28.772’N 101°21.308’E, 860 m, 5-12.vii.2006, Patikhom Tamtipleg., T310; 1 ♂behind checkpoint, 17°27.829’N 101°21.360’E, 691 m, 5-8.vi.2007, Patikhom Tumtipleg., T2514; Phu Kradueng NP, Sutin Glong-lasae leg., 1 ♂%deciduous%, 16°56.569’N 101°49.045’E, 274 m, 18-25.ix.2006, T950, 1 ♂ Dryevergreen forest ( Pongneep forest unit), 16°56.589’N 101°42.074’E, 273 m, 4-11.xi.2006, T792; 1 ♂ Bamboo forestat Lam Huay Taadat Loeiforest unit 2 (E-lerd), 16°56.565’N 101°48.896’E, 273 m, 25.ix- 2.x.2006, Sutin Glong-lasaeleg., T954; 1 ♂ Koke Hin Ngam, 16°51.817’N 101°50.704’E, 270 m, 16-23.viii.2006, Sutin Khonglasaeleg., T486; 1 ♂, Forestprotection unit Loei.5 ( Phakbung), 16°50.540’N 101°41.663’E, 406 m, 13-19.ii.2007, Wuthicahikwanjam leg., T1499; 1 ♂ Roadto Ta Krongwaterfall of Na Noy ForestUnit, 16°48.913’N 101°47.634’E, 265 m, 26.xi-2.xii.2006, Suthin Gong-lasaeleg., T1079. Sakon Nakhon: Phu Phan NP, 1 ♂ Kam Homwaterfall at Haew Sinchai, 17°7.415’N, 104°1.179’E, 347 m, 16-22.ix.2006, Winlon Khongnaraleg., T616; Nam Hom Waterfall# Sao Hi, 17°7.340’N 104°0.788’E, 344 m, Sailom Tongboonchaileg., 2 ♂ 4-10.iii.2007, T2368; 1 ♂ 10-17.iii.2007, T2371; Behindoffice, 17°3.543’N 103°58.452’E, 312 m, Winlon Kongnaraleg., 4 ♂ 8-14.vii.2006, T197; 1 ♂ 27.vii-2.viii.2006, T206; Sailom Tongboonchai leg., 1 ♂ Northof well, 17°3.543’N 103°58.452’E, 312 m, 11-17.i.2007, T1519; 1 ♂ Westof well, 17°3.521’N 103°58.450’E, 322 m, 11-17.i.2007, leg., T1520; 17°3.488’N 103°58.497’E, 318 m, Sailom Tongboonchai leg., 1 ♂ Behindoffice, 15- 21.vii.2006, T202; Behind national parkoffice, 3 ♂ 17-23.i.2007, T1524; 1 ♂ 23-30.i.2007, T1527; Phu Pha YonNP, Channel, 16°55.639’N 104°10.748’E, 295 m, Manop Ngoyjansri& Chatree Cheaukamjanleg., 3 ♂ 5- 11.vii.2006, T289; 1 ♂ 17-23.vii.2006, T295; 1 ♂ Deciduousdipterocarp forest, 16°55.449’N 104°10.757’E, 295 m, 17-23.vii.2006, T297; 1 ♂ Forestnear Huay Nam Pung Dam, 16°54.713’N 103°54.294’E, 289 m, 19- 25.xi.2006, Sailom Tongboonchaileg., T1101. Phitsanulok: Thung Salaeng LuangNP, Mixeddeciduous forest ( Gang Sopawatrfall), 16°52.642’N 100°49.441’E, 481 m, Pongpitak Praneeleg., 3 ♂, Pan trap, 16-17.x.2006, T756; Malaise trap, 1 ♂, 4-12.xi.2006, T771; 1 ♂ 16°52.464’N 100°49.665’E, 501 m, 28.x-4.xi.2006, T769; 2 ♂ Moistevergreen, 16°50.641’N 100°52.894’E, 557 m, 18-25.viii.2006, T569; 1 ♂ Mixeddeciduous forest, 16°50.563’N 100°51.757’E, 481 m, 18-25.ix.2006, T930; Dry Evergreen forest, 16°50.277’N 100°52.917’E, 486 m, 2 ♂ 1-8.iv.2007, Pongpitak& Pranee& Sathitleg., T2394, 1 ♂ 25.ix-2.x.2006, Pongpitak Praneeleg., T932; 1 ♂, Deciduous forest, 16°50.563’N 100°51.757’E 481 m, 8-15.iv.2007, Pongpitak& Pranee& Sathitleg. T2398; Khao Kho NP, Nursery, 520 m, Somchai Chatchumnanand Saink Singtongleg., 1 ♂ 16°52.581’N 101°8.060’E, 12-19.x.2006, T808; 1 ♂ 16°52.573’N 101°8.077’E, 12-19.x.2006, T806; 1 ♂ 16°52.573’N 101°8.077’E, 26.x- 2.xi.2006, T812; 2 ♂ Savanaat nursery, 16°52.568’N 101°8.104’E, 520 m, 5-12.x.2006, T804; 1 ♂ Checkpoint, 16°43.695’N 101°33.797’E, 921 m, 1-8.iii.2007, T2285; 1 ♂ Mixdeciduous, 16°39.589’N 101°8.185’E, 168 m, 5- 101°8.123’E, 230 m, 1 ♂ 26.ii-5.iii.2007, T1610; 1 ♂ 19-26.ii.2007, T1607; 1 ♂office, 16°39.550’N 101°8.123’E, 230 m, 12-19.vii.2006, T167; Mixeddeciduous forest at Ta Phol river, 16°32.561’N 101°2.479’E, 242 m, 1 ♂ 5- 12.xi.2006, T968; 1 ♂ 12-19.xi.2006, T971; 2 ♂ 19-26.xi.2006,. T974; 1 ♂ 26.xi-2.xii.2006, T977; 1 ♂ Mixeddeciduous forest, 16°32.546’N 101°2.501’E, 560 m, 26.xii.2006- 2.i.2007, T1189. Khonkaen: Nam Pong NP, office, Khamphol Jaideeleg., 2 ♂ 16°37.201’N 102°34.481’E, 218 m, 19-26.vii.2006, T113; 16°37.341’N 102°34.467’E, 324 m, 2 ♂ 5-12.vii.2006, T115; 1 ♂ 11-18.vii.2006, T105; 16°37.377’N 102°34.454’E, 344 m, 1 ♂ 5-12.vii.2006, T106; 1 ♂ 12-19.vii.2006, T110; 1 ♂ 19-26.vii.2006, T111; Chaiyaphum: Tat Tone NP, Tawit Jaruphanleg., 1 ♂ Drydipterocarp forest near swamp at Sabsomboon forest unit, 16°1.059’N 101°58.603’E, 674 m, 5-12.xi.2006, T1137; Lawnnear Sab Somboon forestunit, 16°0.792’N 101°58.472’E, 648 m, 2 ♂ 5-12.xi.2006, T1136; 1 ♂ 26.xi-3.xii.2006, T1145; 16°0.738’N 102°1.342’E, 195 m, Tawit Jaruphan&Orawan Budsawong leg., 4 ♂ Forestfire Protection station, 3-9.vi.2006, T2535; Chaiyapoom forestfire station, 1 ♂ 19-23.xii.2006, T1374; 1 ♂ 26.xii.2006- 2.i.2007, T1377; Dry Dipterocarp Forest, 15°59.037’N 102°2.103’E, 250 m, 1 ♂ 19-26.vi.2006, Boonnak Nongpeuleg., T22; 1 ♂ 21-28.vi.2006, Leng Chanteableg., T32; 1 ♂ 23-26.vi.2006, Lumyai Ittichanleg., T27; 1 ♂ 25-27.vi.2006, M. Sharkeyleg., T10; 1 ♂ 26-29.vi.2006, Lumyai Ittichanleg., T30; Tawit Jaruphan & Orawan Budsawong leg., 1 ♂ Nearstream, 15°58.771’N 102°2.397’E, 305 m, 26.vii-7.viii.2006, T227; Phuhang sing, 15°58.723’N 102°2.231’E, 290 m, 1 ♂ 5-12.vii.2006, T220; 1 ♂ 12-19.vii.2006, T223; Nurserynear head water, 15°58.344’N 102°2.169’E, 257 m, 2 ♂ 5-12.viii.2006, T547; 1 ♂, 26.ii-2.iii.2007, T1737; 1 ♂ Entranceto Pa Eangwaterfall, 15°57.520’N 101°54.442’E, 297 m, 12-19.x.2006, T681; 2 ♂ Watertank at Tat Fahwaterfall, 15°56.468’N 102°5.855’E, 245 m, 19-26.iii.2007, T2354; Streamsideat Tat Fahwaterfall, 15°56.463’N 102°5.953’E, 242 m, 1 ♂ 19-26.iii.2007, T2352; 2 ♂ 26.iii-2.iv.2007, T2355; 1 ♂ Oficerhouse at Tat Fahwaterfall, 15°56.461’N 102°5.955’E, 19-26.iii.2007, T2353; Pa Hin Ngam NP, Katae Sanog & Buakaw Adnafai leg., 4 ♂, Savana( Thepanawaterfall), 15°38.948’N 101°25.625’E, 604 m, 19-25.x.2006, T666; ( Thepanawaterfall), 2 ♂ Mixeddeciduous Forest, 15°38.920’N 101°25.077’E, 614 m, 1-7.x.2006, T656; Dryevergreen forest, 15°38.884’N 101°25.840’E, 605 m, 1 ♂ 7-13.x.2006, T661; 2 ♂ 13-19.x.2006, T664; 1 ♂ 19-25.x.2006, T667; 15°38.438’N 101°23.576’E, 780 m, 1 ♂car park at Tung Dok Grajeaw, 6-12.vii.2006, T330; 2 ♂ Thung Dok Kra Jeow in Dryevergreen, Pan trap, 6-7.i.2007, T1451; 15°38.391’N 101°23.609’E, 750 m, Malaise trap, 1 ♂creek at Tung Dok Grajeaw, 6-12.vii.2006, T329; 1 ♂ Carpark at Thung Dok Kra Jeow, 13-19.i.2007, T1459; ecotone between mix deciduous and dipterocarp forest, 15°38.132’N 101°23.922’E, 698 m, 1 ♂ Pan trap, 3- 4.ii.2007, T1636; 1 ♂ Malaise trap, 19-25.ii.2007, T1652; 2 ♂ Dipterocarp forest, 15°38.099’N 101°23.921’E, 698 m, 1-7.ii.2007, T1644; 1 ♂ Naturetrail at Lan Hin Nor, 15°37.615’N 101°23.436’E, 668 m, 9-15.iii.2007, T2328. Suphanburi: 1 ♂, Pu Toei NP, Phu Toeihill top/road, 14°57.32’N 99°26.972’E, 650 m, 1-7.vii.2008, Saunbua.L.leg., T3138. Ubon Ratchathani: Pha Taem NP, 1 ♂ Tung Na Meaung Waterfall, 15°30.914’N 105°35.369’E, 120 m, 6-12.iii.2007, Porntip Tonsu& Bunlu Sapsirileg., T2144; 1 ♂ Wildflower field 1, 15°27.336’N 105°34.870’E, 232 m, 18-25.xi.2006, Sorawitand Thongdeeleg., T1060; 1 ♂ Firstlevel-Huay Sa-nhomwaterfall, 15°27.407’N 105°34.867’E, 230 m, 2-9.v.2007, Sorawit Mingmanleg., T2185 14°28.524’N 101°22.928’E, 757 m; 1 ♂ Dong Ngu How area, 5-12.xii.2006, Pong Sandowleg., T1303; 2 ♂ Cobrazone near Fireprotection office, 5-12.vi.2007, Pong Sandaoleg., T2221. Petchaburi: 1 ♂ Kaeng Krachan NP, Panernthung/km27/water pump, 12°49.151’N 99°22.483’E, 970 m, 5-12.ix.2008, Sirichai& Chusakleg., T4377. Nakhon Nayok: 1 ♂ Khao Yai NP, Naturetrail in secondary forest, 14°24.522’N 101°22.434’E, 750 m, 12-19.viii.2006, Pong Sandaoleg., T402. Chanthaburi: Khao Khitchakut NP, NE/ Prabaht Unit, Suthida& Charoenchaileg., 1 ♂, 50 m, 12°48.779’N 102°9.181’E, 107 m, 7-14.vii.2008, T2976, 2 ♂ 100 m, 12°48.842’N 102°9.144’E, 203 m, 7-14.vii.2008, T2973; Campground/ Prabaht Unit, 12°48.852’N 102°9.204’E, 99 m, 1 ♂ 11-18.viii.2008, T2982, 1 ♂ 28.vii-3.viii.2008, T2985. Prachuab Khiri Khan: 1 ♂, Khao Sam Roi Yot NP, Nursery, 12°7.58’N 99°57.478’E, Pan trap, 3-4.vii.2008, Amnad& Yaileg., T3039. Surat Thani: Khao Sok NP, Pongphanleg., 1 ♂, Klong Mog Unit, 8°53.725’N 98°39.025’E, 87 m, 31.vii-7.viii.2008, T3067; Bang Huaraed, 8°54.555’N 98°30.522’E, 122 m, 1 ♂, 7-14.iv.2009, T4569, 1 ♂ 10- 17.viii.2008, T3361. Nakhon Si Thammarat: Namtok Yong NP, 1 ♂ Roadto Khao Mhen. 150 m, from Nern499, 8°16.959’N 99°39.149’E 499 m, 15-22.vii.2008, Samnaokan.S.leg., T3096; 1 ♂, Klong Jangwaterfall, 8°16.283’N 99°38.702’E, 154 m, 29.xii.2008- 5.i.2009, Samnaokan;S. leg., T4290; Behindcampground lavatory, 8°10.434’N 99°44.508’E, 80 m, U.prai.K. leg., 2 ♂ 12-19.viii.2008, T3080, 1 ♂ 29.vii-5.viii.2008, T3072, 1 ♂ 19- 26.viii.2008, T3074. metasoma varying between dark castaneous with castaneous spots or fully castaneous; frontal with punctures small, shallow and irregularly sparse; vertex crest straight; lower tooth of mandible sub-angulate; last flagellomere with angulate apex; placoid sensillae elliptical; eye gibbous; scutellar groove slightly convex; ventral outer lobe of aedeagal dorsal body straight, varying between very short (apical margin of ventral pair not reaching basal region of ventral pair) and short (apical margin of ventral pair aligned with median region of ventral pair).
Remarks. Dissomphalus verus sp. nov.is similar to D. brownibecause of the following characteristics: hypopygium with long and narrow median stalk, lateral margins slightly convergent posterad, posterior margin with angulate corners, dorsal outer lobe of aedeagal dorsal body with laminar process, ventral outer lobe of dorsal body short and arched; aedeagal ventral ramus as long as dorsal body, apex with triangular cross-section. However, D. verus sp. nov.differs from D. brownias follows: lateral stalks of hypopygium triangular, apical region of median stalk with constriction, posterior margin slightly sinuous; base of paramere wider than basiparamere apex; base of aedeagal dorsal body wide, laminar process sickle-shaped with apical margin strongly concave; outer ventral lobe of dorsal body strongly arched with round apex.
Distribution. Thailand( Chaiyaphum, Mae Hong Son, Chiang Mai, Loei, Sakon Nakhon, Phitsanulok, Chaiyaphum, Suphanburi, Ubon Ratchathani, Petchaburi, Nakhon Nayok, Chanthaburi, Prachuab Khiri Khan, Surat Thaniand Nakhon Si Thammarat).
2006-10-01
2006-10-07
2006-10-01
T
Pa Hin Ngam NP & Katae Sa-nog & Buakaw Adnafai
614
15.648666
Mixed
1
101.41795
26
27
T656
1
1
Chaiyaphum
holotype
[382,1014,763,788]
Mae Hong Son
26
27
1
Mae Hong Son
paratype
2008-01-27
2008-02-03
2008-01-27
T
Kamkoon
19.3436
Huai Fai Kor
87
97.988335
Namtok Mae
26
27
T3496
1
1
Surin
paratype
2006-08-30
2006-09-06
2006-08-30
T
Doi Inthanon NP & Y. Areeluck
700
18.538517
Vachiratharn Falls
1
98.6008
26
27
T236
1
1
Chiang Mai
paratype
[151,375,871,896]
Phu Ruea NP
Loei
26
27
1
Loei
paratype
2006-07-05
2006-07-12
2006-07-05
T
Patikhom Tamtip
860
17.479534
Subhnonghin
1
101.35513
26
27
T310
1
1
Loei
paratype
[225,1381,908,932]
2007-06-05
2007-06-08
2007-06-05
T
Patikhom Tumtip
691
17.463818
1
101.356
26
27
T2514
1
1
Loei
paratype
2006-09-18
2006-09-25
2006-09-18
T
274
16.942816
1
101.81741
26
27
T950
1
1
Loei
paratype
[224,1411,980,1004]
2006-11-04
2006-11-11
2006-11-04
T
273
16.94315
Pong
1
101.70123
Dry
26
27
T792
1
1
Loei
paratype
2006-09-25
2006-10-02
2006-09-25
T
Sutin Glong-lasae
273
16.942749
Lam Huay Taad
1
101.81493
Bamboo forest
26
27
T954
1
1
Loei
paratype
2006-08-16
2006-08-23
2006-08-16
T
Sutin Khonglasae
270
16.863617
Koke Hin Ngam
1
101.84507
26
27
T486
1
1
Loei
paratype
2007-02-13
2007-02-19
2007-02-13
T
Wuthicahi
406
16.842333
Phakbung
1
101.69438
Forest
26
27
T1499
1
1
Loei
paratype
2006-11-26
2006-12-02
2006-11-26
T
Suthin Gong-lasae
Road
265
16.815216
Na Noy Forest
1
101.7939
Ta Krong
26
27
T1079
1
1
Loei
paratype
Phu Phan NP
Sakon Nakhon
26
27
1
Sakon Nakhon
paratype
2006-09-16
2006-09-22
2006-09-16
T
Winlon Khongnara
347
17.123583
Haew Sin
1
104.01965
Kam Hom
26
27
T616
1
1
Sakon Nakhon
paratype
[410,1431,1231,1255]
Sailom Tongboonchai
344
17.122334
Sao Hi
1
104.01313
Nam Hom Waterfall
26
27
1
Sakon Nakhon
paratype
[151,443,1268,1292]
2007-03-04
2007-03-10
2007-03-04
T
26
27
T2368
2
2
Sakon Nakhon
paratype
[457,762,1268,1292]
2007-03-10
2007-03-17
2007-03-10
T
26
27
T2371
1
1
Sakon Nakhon
paratype
Winlon Kongnara
312
17.05905
Behind
1
103.9742
26
27
1
Sakon Nakhon
paratype
[328,612,1303,1327]
2006-07-08
2006-07-14
2006-07-08
T
26
27
T197
4
4
Sakon Nakhon
paratype
[626,954,1303,1327]
2006-07-27
2006-08-02
2006-07-27
T
26
27
T206
1
1
Sakon Nakhon
paratype
2007-01-11
2007-01-17
2007-01-11
T
312
17.05905
North
1
103.9742
26
27
T1519
1
1
Sakon Nakhon
paratype
2007-01-11
2007-01-17
2007-01-11
T
322
17.058683
West
1
103.97417
26
27
T1520
1
1
Sakon Nakhon
paratype
2006-07-15
2006-07-21
2006-07-15
T
Behind
26
27
T202
1
1
Sakon Nakhon
paratype
[368,681,1412,1436]
Behind national park
26
27
1
Sakon Nakhon
paratype
[693,978,1412,1436]
2007-01-17
2007-01-23
2007-01-17
T
26
27
T1524
3
3
Sakon Nakhon
paratype
[991,1276,1412,1436]
2007-01-23
2007-01-30
2007-01-23
T
26
27
T1527
1
1
Sakon Nakhon
paratype
Manop Ngoyjansri & Chatree Cheaukamjan
295
16.927317
Channel
1
104.17913
Phu Pha Yon
26
27
1
Sakon Nakhon
paratype
2006-07-05
2006-07-11
2006-07-05
T
26
27
T289
3
3
Sakon Nakhon
paratype
[368,661,1484,1508]
2006-07-17
2006-07-23
2006-07-17
T
26
27
T295
1
1
Sakon Nakhon
paratype
2006-07-17
2006-07-23
2006-07-17
T
295
16.92415
Deciduous
1
104.17928
26
27
T297
1
1
Sakon Nakhon
paratype
2006-11-19
2006-11-25
2006-11-19
T
Sailom Tongboonchai
289
16.911884
Huay Nam Pung Dam
1
103.9049
Forest
26
27
T1101
1
1
Sakon Nakhon
paratype
Pongpitak Pranee
Thung Salaeng Luang
481
16.877367
Gang Sopa
1
100.82402
Mixed
26
27
1
Phitsanulok
paratype
2006-10-16
2006-10-17
2006-10-16
T
26
27
T756
3
3
Phitsanulok
paratype
[381,661,1628,1652]
2006-11-04
2006-11-12
2006-11-04
T
26
27
T771
1
1
Phitsanulok
paratype
[674,1379,1628,1652]
2006-10-28
2006-11-04
2006-10-28
T
501
16.8744
1
100.82775
26
27
T769
1
1
Phitsanulok
paratype
2006-08-18
2006-08-25
2006-08-18
T
557
16.844017
Moist
1
100.88157
26
27
T569
2
2
Phitsanulok
paratype
2006-09-18
2006-09-25
2006-09-18
T
481
16.842716
Mixed
1
100.86262
26
27
T930
1
1
Phitsanulok
paratype
486
16.83795
Dry Evergreen forest
1
100.88195
26
27
1
Phitsanulok
paratype
[187,846,1735,1759]
2007-04-01
2007-04-08
2007-04-01
T
Pongpitak & Pranee & Sathit
26
27
T2394
2
2
Phitsanulok
paratype
[858,1410,1735,1759]
2006-09-25
2006-10-02
2006-09-25
T
Pongpitak Pranee
26
27
T932
1
1
Phitsanulok
paratype
2007-04-08
2007-04-15
2007-04-08
T
Pongpitak & Pranee & Sathit
481
16.842716
Deciduous forest
1
100.86262
26
27
T2398
1
1
Phitsanulok
paratype
[151,1065,1807,1832]
Khao Kho NP & Somchai Chatchumnan & Saink Singtong
520
Nursery
26
27
1
Phitsanulok
paratype
2006-10-12
2006-10-19
2006-10-12
T
16.87635
1
101.13433
26
27
T808
1
1
Phitsanulok
paratype
[394,993,1844,1868]
2006-10-12
2006-10-19
2006-10-12
T
16.876217
1
101.13461
26
27
T806
1
1
Phitsanulok
paratype
2006-10-26
2006-11-02
2006-10-26
T
16.876217
1
101.13461
26
27
T812
1
1
Phitsanulok
paratype
[348,1233,1879,1904]
2006-10-05
2006-10-12
2006-10-05
T
520
16.876133
Savana
1
101.13506
26
27
T804
2
2
Phitsanulok
paratype
2007-03-01
2007-03-08
2007-03-01
T
921
16.72825
Checkpoint
1
101.563286
26
27
T2285
1
1
Phitsanulok
paratype
168
27
28
16.659817
Mix
1
101.13538
26
27
1
1
Phitsanulok
paratype
[391,696,151,175]
2007-02-26
2007-03-05
2007-02-26
T
27
28
T1610
1
1
Phitsanulok
paratype
[707,996,151,175]
2007-02-19
2007-02-26
2007-02-19
T
27
28
T1607
1
1
Phitsanulok
paratype
2006-07-12
2006-07-19
2006-07-12
T
230
16.659166
1
101.13538
27
28
T167
1
1
Phitsanulok
paratype
[494,1348,188,212]
242
16.542683
Ta Phol river
1
101.04131
Mixed
27
28
1
Phitsanulok
paratype
2006-11-05
2006-11-12
2006-11-05
T
27
28
T968
1
1
Phitsanulok
paratype
[364,655,223,247]
2006-11-12
2006-11-19
2006-11-12
T
27
28
T971
1
1
Phitsanulok
paratype
[669,967,223,247]
2006-11-19
2006-11-26
2006-11-19
T
27
28
T974
2
2
Phitsanulok
paratype
[981,1294,223,247]
2006-11-26
2006-12-02
2006-11-26
T
27
28
T977
1
1
Phitsanulok
paratype
2006-12-26
2007-01-02
2006-12-26
T
560
16.542433
Mixed
1
101.04169
27
28
T1189
1
1
Phitsanulok
paratype
Nam Pong NP & Khamphol Jaidee
Khonkaen
27
28
1
Phitsanulok
paratype
[519,1270,295,319]
2006-07-19
2006-07-26
2006-07-19
T
218
16.620016
1
102.574684
27
28
T113
2
2
Phitsanulok
paratype
[405,679,332,356]
2006-07-05
2006-07-12
2006-07-05
T
27
28
T115
2
2
Phitsanulok
paratype
[690,977,332,356]
2006-07-11
2006-07-18
2006-07-11
T
27
28
T105
1
1
Phitsanulok
paratype
2006-07-05
2006-07-12
2006-07-05
T
27
28
T106
1
1
Phitsanulok
paratype
[398,700,367,391]
2006-07-12
2006-07-19
2006-07-12
T
27
28
T110
1
1
Phitsanulok
paratype
[716,1016,367,391]
2006-07-19
2006-07-26
2006-07-19
T
27
28
T111
1
1
Phitsanulok
paratype
2006-11-05
2006-11-12
2006-11-05
T
674
16.01765
Sab
1
101.976715
Dry
27
28
T1137
1
1
Phitsanulok
paratype
[429,1226,439,463]
648
16.0132
Sab Somboon forest
1
101.97453
Lawn
27
28
1
Phitsanulok
paratype
2006-11-05
2006-11-12
2006-11-05
T
27
28
T1136
2
2
Phitsanulok
paratype
[237,553,476,500]
2006-11-26
2006-12-03
2006-11-26
T
27
28
T1145
1
1
Phitsanulok
paratype
2006-06-03
2006-06-09
2006-06-03
T
Protection station
Forest
27
28
T2535
4
4
Phitsanulok
paratype
[747,1092,511,535]
Chaiyapoom forest
27
28
1
Phitsanulok
paratype
[1104,1410,511,535]
2006-12-19
2006-12-23
2006-12-19
T
27
28
T1374
1
1
Phitsanulok
paratype
2006-12-26
2007-01-02
2006-12-26
T
27
28
T1377
1
1
Phitsanulok
paratype
[520,1200,548,572]
250
15.98395
Dry Dipterocarp Forest
1
102.03505
27
28
1
Phitsanulok
paratype
2006-06-19
2006-06-26
2006-06-19
T
Boonnak Nongpeu
27
28
T22
1
1
Phitsanulok
paratype
[494,1011,583,608]
2006-06-21
2006-06-28
2006-06-21
T
Leng Chanteab
27
28
T32
1
1
Phitsanulok
paratype
2006-06-23
2006-06-26
2006-06-23
T
Lumyai Ittichan
27
28
T27
1
1
Phitsanulok
paratype
[266,729,620,644]
2006-06-25
2006-06-27
2006-06-25
T
M. Sharkey
27
28
T10
1
1
Phitsanulok
paratype
[741,1253,620,644]
2006-06-26
2006-06-29
2006-06-26
T
Lumyai Ittichan
27
28
T30
1
1
Phitsanulok
paratype
[483,1377,655,679]
2006-07-26
2006-08-07
2006-07-26
T
305
15.979517
Near
1
102.03995
27
28
T227
1
1
Phitsanulok
paratype
290
15.978717
Phu
1
102.037186
27
28
1
Phitsanulok
paratype
[681,964,692,716]
2006-07-05
2006-07-12
2006-07-05
T
27
28
T220
1
1
Phitsanulok
paratype
[977,1274,692,716]
2006-07-12
2006-07-19
2006-07-12
T
27
28
T223
1
1
Phitsanulok
paratype
257
15.9724
Nursery
1
102.03615
27
28
1
Phitsanulok
paratype
[717,1020,727,751]
2006-08-05
2006-08-12
2006-08-05
T
27
28
T547
2
2
Phitsanulok
paratype
[1038,1370,727,751]
2007-02-26
2007-03-02
2007-02-26
T
27
28
T1737
1
1
Phitsanulok
paratype
2006-10-12
2006-10-19
2006-10-12
T
297
15.958667
Pa Eang
1
101.907364
Entrance
27
28
T681
1
1
Phitsanulok
paratype
2007-03-19
2007-03-26
2007-03-19
T
245
15.9411335
Tat Fah
1
102.09758
Water
27
28
T2354
2
2
Phitsanulok
paratype
242
15.94105
Tat Fah
1
102.09922
Streamside
27
28
1
Phitsanulok
paratype
[410,726,836,860]
2007-03-19
2007-03-26
2007-03-19
T
27
28
T2352
1
1
Phitsanulok
paratype
[743,1072,836,860]
2007-03-26
2007-04-02
2007-03-26
T
27
28
T2355
2
2
Phitsanulok
paratype
2007-03-19
2007-03-26
2007-03-19
T
15.941016
Tat Fah
1
102.09925
Oficer
27
28
T2353
1
1
Phitsanulok
paratype
2006-10-19
2006-10-25
2006-10-19
T
Thepana
604
15.649134
Thepana
1
101.427086
Savana
27
28
T666
4
4
Phitsanulok
paratype
[278,1256,943,967]
2006-10-01
2006-10-07
2006-10-01
T
614
15.648666
Forest
1
101.41795
Mixed
27
28
T656
2
2
Phitsanulok
paratype
605
15.6480665
Dry
1
101.430664
27
28
1
Phitsanulok
paratype
[649,916,980,1004]
2006-10-07
2006-10-13
2006-10-07
T
27
28
T661
1
1
Phitsanulok
paratype
[930,1211,980,1004]
2006-10-13
2006-10-19
2006-10-13
T
27
28
T664
2
2
Phitsanulok
paratype
2006-10-19
2006-10-25
2006-10-19
T
27
28
T667
1
1
Phitsanulok
paratype
[645,1295,1015,1040]
2006-07-06
2006-07-12
2006-07-06
T
Tung Dok Grajeaw
27
28
T330
1
1
Phitsanulok
paratype
2007-01-06
2007-01-07
2007-01-06
T
Thung Dok Kra Jeow in Dry
27
28
T1451
2
2
Phitsanulok
paratype
2006-07-06
2006-07-12
2006-07-06
T
Tung Dok Grajeaw
27
28
T329
1
1
Phitsanulok
paratype
[750,1431,1087,1111]
2007-01-13
2007-01-19
2007-01-13
T
Thung Dok Kra Jeow
Car
27
28
T1459
1
1
Phitsanulok
paratype
2007-02-03
2007-02-04
2007-02-03
T
27
28
T1636
1
1
Phitsanulok
paratype
[352,793,1159,1183]
2007-02-19
2007-02-25
2007-02-19
T
27
28
T1652
1
1
Phitsanulok
paratype
2007-02-01
2007-02-07
2007-02-01
T
698
15.634983
Dipterocarp forest
1
101.39868
27
28
T1644
2
2
Phitsanulok
paratype
[413,1433,1196,1220]
2007-03-09
2007-03-15
2007-03-09
T
668
15.626917
Lan Hin Nor
1
101.3906
Nature
27
28
T2328
1
1
Phitsanulok
paratype
2008-07-01
2008-07-07
2008-07-01
T
Pu Toei NP & Saunbua. L.
650
14.955334
Phu Toei
9
99.44953
27
28
T3138
1
1
Suphan Buri
paratype
[294,686,1267,1292]
Pha Taem NP
Ubon Ratchathani
27
28
1
Ubon Ratchathani
paratype
2007-03-06
2007-03-12
2007-03-06
T
Porntip Tonsu & Bunlu Sapsiri
120
15.515233
Tung Na Meaung Waterfall
1
105.589485
27
28
T2144
1
1
Ubon Ratchathani
paratype
2006-11-18
2006-11-25
2006-11-18
T
Sorawit & Thongdee
232
15.4556
Wild
1
105.58117
27
28
T1060
1
1
Ubon Ratchathani
paratype
2007-05-02
2007-05-09
2007-05-02
T
Sorawit Mingman
230
15.456783
Sa-nhom
1
105.581116
First
27
28
T2185
1
1
Ubon Ratchathani
paratype
2006-12-05
2006-12-12
2006-12-05
T
Pong Sandow
Dong Ngu How area
27
28
T1303
1
1
Ubon Ratchathani
paratype
2007-06-05
2007-06-12
2007-06-05
T
Pong Sandao
Fire
Cobra
27
28
T2221
2
2
Ubon Ratchathani
paratype
[463,599,1447,1472]
Petchaburi
27
28
1
Ubon Ratchathani
paratype
2008-09-05
2008-09-12
2008-09-05
T
Kaeng Krachan NP & Sirichai & Chusak
970
12.819183
Panernthung
1
99.37472
27
28
T4377
1
1
Ubon Ratchathani
paratype
2006-08-12
2006-08-19
2006-08-12
T
Khao Yai NP & Pong Sandao
750
14.4087
Nature
1
101.3739
27
28
T402
1
1
Nakhon Nayok
paratype
Khao Khitchakut NP & Prabaht Unit & Suthida & Charoenchai
Chanthaburi
27
28
1
Chanthaburi
paratype
2008-07-07
2008-07-14
2008-07-07
T
50
12.8129835
1
102.153015
27
28
T2976
1
1
Chanthaburi
paratype
[405,1183,1591,1615]
2008-07-07
2008-07-14
2008-07-07
T
100
12.8140335
1
102.1524
27
28
T2973
2
2
Chanthaburi
paratype
99
12.8142
Prabaht Unit
1
102.1534
Campground
27
28
1
Chanthaburi
paratype
[617,940,1628,1652]
2008-08-11
2008-08-18
2008-08-11
T
27
28
T2982
1
1
Chanthaburi
paratype
[956,1305,1628,1652]
2008-07-28
2008-08-03
2008-07-28
T
27
28
T2985
1
1
Chanthaburi
paratype
2008-07-03
2008-07-04
2008-07-03
T
Khao Sam Roi Yot NP & Amnad & Yai
12.126333
Nursery
9
99.95797
27
28
T3039
1
1
Chanthaburi
paratype
2008-07-31
2008-08-07
2008-07-31
T
87
8.895416
Klong Mog Unit
1
98.65041
27
28
T3067
1
1
Chanthaburi
paratype
[455,1015,1735,1760]
122
8.90925
Bang Huaraed
1
98.5087
27
28
1
Chanthaburi
paratype
[1030,1327,1735,1759]
2009-04-07
2009-04-14
2009-04-07
T
27
28
T4569
1
1
Chanthaburi
paratype
2008-08-10
2008-08-17
2008-08-10
T
27
28
T3361
1
1
Chanthaburi
paratype
[388,884,1771,1796]
Namtok Yong NP
Nakhon Si Thammarat
27
28
1
Nakhon Si Thammarat
paratype
2008-07-15
2008-07-22
2008-07-15
T
Samnaokan. S.
Road
150
8.28265
Nern
1
99.65248
Khao Mhen.
27
28
T3096
1
1
Nakhon Si Thammarat
paratype
2008-12-29
2009-01-05
2008-12-29
T
Samnaokan
154
8.271383
Klong Jang
1
99.645035
27
28
T4290
1
1
Nakhon Si Thammarat
paratype
80
8.1739
Behind
1
99.7418
27
28
1
Nakhon Si Thammarat
paratype
[683,994,1879,1903]
2008-08-12
2008-08-19
2008-08-12
T
27
28
T3080
2
2
Nakhon Si Thammarat
paratype
[1005,1338,1879,1903]
2008-07-29
2008-08-05
2008-07-29
T
27
28
T3072
1
1
Nakhon Si Thammarat
paratype
2008-08-19
2008-08-26
2008-08-19
T
27
28
T3074
1
1
Nakhon Si Thammarat
paratype