Revision of Thai Dissomphalus Ashmead, 1893 (Hymenoptera, Bethylidae), with description of twenty four new species MUGRABI, DANIELE F. AZEVEDO, CELSO O. Zootaxa 2013 2013-05-22 3662 1 1 73 Mugrabi & Azevedo Mugrabi & Azevedo [151,636,1375,1400] Insecta Bethylidae Dissomphalus GBIF Animalia Hymenoptera 25 26 Arthropoda species verus sp. nov.  ( Figs 27–30, 104, 159–161)  Description.MALE. Body length 3.65 mm. LFW 2.51 mm. Maximum wing width 1.07 mm. Color. Head and mesosoma black; metasoma dark castaneous.  Head. Mandible with five apical teeth. Clypeus with median lobe subtrapezoidal; median tooth rounded; median carina complete apically or nearly so, angled in profile. Ratio of first four antennal segments about 14:7:6:5. Frons strongly coriaceous, punctures large. LH 0.95 × WH, WF 0.63 × WH, WF 1.26 × HE, OOL 1.23 × WOT, DAO 0.35 × WOT, posterior ocelli distant from vertex crest 1 × DAO, frontal angle of ocellar triangle right, VOL 0.68 × HE. Vertex crest weakly concave. Occipital carina not visible in dorsal view.  Mesosoma. Pronotal disc strongly coriaceous, anterior margin ecarinate. Notauli deep, complete, widening posterad, converging posterad. Propodeal disc 0.55 × as long as wide; basal triangle rugulose, posterolateral area rugulose; median carina complete; lateral margins strongly convex, parallel; posterior carina well-defined, straight. Lateral of propodeum mostly striolate. Propodeal declivity mostly rugulose-strigate.  Metasoma. Tergal process with deep, subtriangular and sublateral pair of depressions; depressions with external lateral margin higher than internal lateral margin, 0.83 × as long as tergite II, diverging posterad, distant on top with short and dorsad tuft of setae ( Figs 27–30). Hypopygium with median stalk 1.15 × as long as hypopygeal plate; median stalk narrow; corner of anterior margin angular; lateral stalk of hypopygium 0.11 × as long as median stalk; posterior margin strongly concave; corner angalute ( Fig. 104).  Genitalia( Figs 159–161). Genital ring produced, each half convex in dorsal view. Basiparamere with dorsal surface much wider apicad, much narrower than ventral surface. Paramere with basal area wider than basiparamere apex; apex rounded; apical margin weakly concave in ventral view, not curved; inner margin of ventral surface with triangular spine at median region. Basivolsella setose, with several thin, short and apical setae. Cuspis biramous. Aedeagus 3.72 × as long as wide; dorsal body of aedeagus wider basally, narrowing abruptly apicad, lateral margin strongly convex basally, parallel, not curved dorsad, apex with two pair of outer lobes; dorsal pair 1 × as long as wide, straight, with apex subangulate and dorsad, outer lateral with laminar process, elliptical, with lateral surface wider than dorsal and ventral surfaces; ventral pair tubular, short, slightly arched, laterad, with apical margin rounded and surface smooth, much shorter than dorsal lobe of aedeagus; base of dorsal body of aedeagus strongly sclerotized, weakly concave, with median region bilobate; ventral ramus of aedeagus as high as dorsal body, lateral margin straight, evenly narrow and base as narrow as apex. Apodeme extending beyond genital ring. FEMALE unknown.   Material examined.   Holotype, ♂. THAILAND.  Chaiyaphum: Pa Hin Ngam NP, Mixeddeciduous forest (Thepana waterfall), 15°38.920’N 101°25.077’E,  614 m, Malaise trap,  1-7.x.2006, Katae Sa-nog& Buakaw Adnafaileg., T656.   Paratypes: THAILAND. Malaise trap, Mae Hong Son :  1 ♂ Namtok Mae SurinNP, E/ Huai Fai Korreservoir, 19°20.616’N 97°59.3’E,  27.i-3.ii.2008, Kamkoon;A leg., T3496.   Chiang Mai: 1 ♂ Doi Inthanon NP, Vachiratharn Falls, 18°32.311’N 98°36.048’E,  700 m,  30.viii-6.ix.2006, Y. Areeluckleg., T236.   Loei: Phu Ruea NP,  1 ♂ Subhnonghin, 17°28.772’N 101°21.308’E,  860 m,  5-12.vii.2006, Patikhom Tamtipleg., T310;  1 ♂behind checkpoint, 17°27.829’N 101°21.360’E,  691 m,  5-8.vi.2007, Patikhom Tumtipleg., T2514; Phu Kradueng NP, Sutin Glong-lasae leg.,  1 ♂%deciduous%, 16°56.569’N 101°49.045’E,  274 m,  18-25.ix.2006, T950,  1 ♂ Dryevergreen forest ( Pongneep forest unit), 16°56.589’N 101°42.074’E,  273 m,  4-11.xi.2006, T792;  1 ♂ Bamboo forestat Lam Huay Taadat Loeiforest unit 2 (E-lerd), 16°56.565’N 101°48.896’E,  273 m,  25.ix- 2.x.2006, Sutin Glong-lasaeleg., T954;  1 ♂ Koke Hin Ngam, 16°51.817’N 101°50.704’E,  270 m,  16-23.viii.2006, Sutin Khonglasaeleg., T486;  1 ♂, Forestprotection unit Loei.5 ( Phakbung), 16°50.540’N 101°41.663’E,  406 m,  13-19.ii.2007, Wuthicahikwanjam leg., T1499;  1 ♂ Roadto Ta Krongwaterfall of Na Noy ForestUnit, 16°48.913’N 101°47.634’E,  265 m,  26.xi-2.xii.2006, Suthin Gong-lasaeleg., T1079.   Sakon Nakhon: Phu Phan NP,  1 ♂ Kam Homwaterfall at Haew Sinchai, 17°7.415’N, 104°1.179’E,  347 m,  16-22.ix.2006, Winlon Khongnaraleg., T616;  Nam Hom Waterfall# Sao Hi, 17°7.340’N 104°0.788’E,  344 m, Sailom Tongboonchaileg.,  2 ♂  4-10.iii.2007, T2368;  1 ♂  10-17.iii.2007, T2371;  Behindoffice, 17°3.543’N 103°58.452’E,  312 m, Winlon Kongnaraleg.,  4 ♂  8-14.vii.2006, T197;  1 ♂  27.vii-2.viii.2006, T206; Sailom Tongboonchai leg.,  1 ♂ Northof well, 17°3.543’N 103°58.452’E,  312 m,  11-17.i.2007, T1519;  1 ♂ Westof well, 17°3.521’N 103°58.450’E,  322 m,  11-17.i.2007, leg., T1520; 17°3.488’N 103°58.497’E, 318 m, Sailom Tongboonchai leg.,  1 ♂ Behindoffice,  15- 21.vii.2006, T202;  Behind national parkoffice,  3 ♂  17-23.i.2007, T1524;  1 ♂  23-30.i.2007, T1527;  Phu Pha YonNP, Channel, 16°55.639’N 104°10.748’E,  295 m, Manop Ngoyjansri& Chatree Cheaukamjanleg.,  3 ♂  5- 11.vii.2006, T289;  1 ♂  17-23.vii.2006, T295;  1 ♂ Deciduousdipterocarp forest, 16°55.449’N 104°10.757’E,  295 m,  17-23.vii.2006, T297;  1 ♂ Forestnear Huay Nam Pung Dam, 16°54.713’N 103°54.294’E,  289 m,  19- 25.xi.2006, Sailom Tongboonchaileg., T1101.   Phitsanulok: Thung Salaeng LuangNP, Mixeddeciduous forest ( Gang Sopawatrfall), 16°52.642’N 100°49.441’E,  481 m, Pongpitak Praneeleg.,  3 ♂, Pan trap,  16-17.x.2006, T756; Malaise trap,  1 ♂,  4-12.xi.2006, T771;  1 ♂ 16°52.464’N 100°49.665’E,  501 m,  28.x-4.xi.2006, T769;  2 ♂ Moistevergreen, 16°50.641’N 100°52.894’E,  557 m,  18-25.viii.2006, T569;  1 ♂ Mixeddeciduous forest, 16°50.563’N 100°51.757’E,  481 m,  18-25.ix.2006, T930;  Dry Evergreen forest, 16°50.277’N 100°52.917’E,  486 m,  2 ♂  1-8.iv.2007, Pongpitak& Pranee& Sathitleg., T2394,  1 ♂  25.ix-2.x.2006, Pongpitak Praneeleg., T932;  1 ♂, Deciduous forest, 16°50.563’N 100°51.757’E  481 m,  8-15.iv.2007, Pongpitak& Pranee& Sathitleg. T2398;  Khao Kho NP, Nursery,  520 m, Somchai Chatchumnanand Saink Singtongleg.,  1 ♂ 16°52.581’N 101°8.060’E,  12-19.x.2006, T808;  1 ♂ 16°52.573’N 101°8.077’E,  12-19.x.2006, T806;  1 ♂ 16°52.573’N 101°8.077’E,  26.x- 2.xi.2006, T812;  2 ♂ Savanaat nursery, 16°52.568’N 101°8.104’E,  520 m,  5-12.x.2006, T804;  1 ♂ Checkpoint, 16°43.695’N 101°33.797’E,  921 m,  1-8.iii.2007, T2285;  1 ♂ Mixdeciduous, 16°39.589’N 101°8.185’E,  168 m, 5- 101°8.123’E,  230 m,  1 ♂  26.ii-5.iii.2007, T1610;  1 ♂  19-26.ii.2007, T1607;  1 ♂office, 16°39.550’N 101°8.123’E,  230 m,  12-19.vii.2006, T167;  Mixeddeciduous forest at Ta Phol river, 16°32.561’N 101°2.479’E,  242 m,  1 ♂  5- 12.xi.2006, T968;  1 ♂  12-19.xi.2006, T971;  2 ♂  19-26.xi.2006,. T974;  1 ♂  26.xi-2.xii.2006, T977;  1 ♂ Mixeddeciduous forest, 16°32.546’N 101°2.501’E,  560 m,  26.xii.2006-  2.i.2007, T1189.   Khonkaen: Nam Pong NP, office, Khamphol Jaideeleg.,  2 ♂ 16°37.201’N 102°34.481’E,  218 m,  19-26.vii.2006, T113; 16°37.341’N 102°34.467’E, 324 m,  2 ♂  5-12.vii.2006, T115;  1 ♂  11-18.vii.2006, T105; 16°37.377’N 102°34.454’E, 344 m,  1 ♂  5-12.vii.2006, T106;  1 ♂  12-19.vii.2006, T110;  1 ♂  19-26.vii.2006, T111;  Chaiyaphum: Tat Tone NP, Tawit Jaruphanleg.,  1 ♂ Drydipterocarp forest near swamp at Sabsomboon forest unit, 16°1.059’N 101°58.603’E,  674 m,  5-12.xi.2006, T1137;  Lawnnear Sab Somboon forestunit, 16°0.792’N 101°58.472’E,  648 m,  2 ♂  5-12.xi.2006, T1136;  1 ♂  26.xi-3.xii.2006, T1145; 16°0.738’N 102°1.342’E, 195 m, Tawit Jaruphan&Orawan Budsawong leg.,  4 ♂ Forestfire Protection station,  3-9.vi.2006, T2535;  Chaiyapoom forestfire station,  1 ♂  19-23.xii.2006, T1374;  1 ♂  26.xii.2006-  2.i.2007, T1377;  Dry Dipterocarp Forest, 15°59.037’N 102°2.103’E,  250 m,  1 ♂  19-26.vi.2006, Boonnak Nongpeuleg., T22;  1 ♂  21-28.vi.2006, Leng Chanteableg., T32;  1 ♂  23-26.vi.2006, Lumyai Ittichanleg., T27;  1 ♂  25-27.vi.2006, M. Sharkeyleg., T10;  1 ♂  26-29.vi.2006, Lumyai Ittichanleg., T30; Tawit Jaruphan & Orawan Budsawong leg.,  1 ♂ Nearstream, 15°58.771’N 102°2.397’E,  305 m,  26.vii-7.viii.2006, T227;  Phuhang sing, 15°58.723’N 102°2.231’E,  290 m,  1 ♂  5-12.vii.2006, T220;  1 ♂  12-19.vii.2006, T223;  Nurserynear head water, 15°58.344’N 102°2.169’E,  257 m,  2 ♂  5-12.viii.2006, T547;  1 ♂,  26.ii-2.iii.2007, T1737;  1 ♂ Entranceto Pa Eangwaterfall, 15°57.520’N 101°54.442’E,  297 m,  12-19.x.2006, T681;  2 ♂ Watertank at Tat Fahwaterfall, 15°56.468’N 102°5.855’E,  245 m,  19-26.iii.2007, T2354;  Streamsideat Tat Fahwaterfall, 15°56.463’N 102°5.953’E,  242 m,  1 ♂  19-26.iii.2007, T2352;  2 ♂  26.iii-2.iv.2007, T2355;  1 ♂ Oficerhouse at Tat Fahwaterfall, 15°56.461’N 102°5.955’E,  19-26.iii.2007, T2353; Pa Hin Ngam NP, Katae Sanog & Buakaw Adnafai leg.,  4 ♂, Savana( Thepanawaterfall), 15°38.948’N 101°25.625’E,  604 m,  19-25.x.2006, T666; ( Thepanawaterfall),  2 ♂ Mixeddeciduous Forest, 15°38.920’N 101°25.077’E,  614 m,  1-7.x.2006, T656;  Dryevergreen forest, 15°38.884’N 101°25.840’E,  605 m,  1 ♂  7-13.x.2006, T661;  2 ♂  13-19.x.2006, T664;  1 ♂  19-25.x.2006, T667; 15°38.438’N 101°23.576’E, 780 m,  1 ♂car park at Tung Dok Grajeaw,  6-12.vii.2006, T330;  2 ♂ Thung Dok Kra Jeow in Dryevergreen, Pan trap,  6-7.i.2007, T1451; 15°38.391’N 101°23.609’E, 750 m, Malaise trap,  1 ♂creek at Tung Dok Grajeaw,  6-12.vii.2006, T329;  1 ♂ Carpark at Thung Dok Kra Jeow,  13-19.i.2007, T1459; ecotone between mix deciduous and dipterocarp forest, 15°38.132’N 101°23.922’E, 698 m,  1 ♂ Pan trap,  3- 4.ii.2007, T1636;  1 ♂ Malaise trap,  19-25.ii.2007, T1652;  2 ♂ Dipterocarp forest, 15°38.099’N 101°23.921’E,  698 m,  1-7.ii.2007, T1644;  1 ♂ Naturetrail at Lan Hin Nor, 15°37.615’N 101°23.436’E,  668 m,  9-15.iii.2007, T2328.   Suphanburi: 1 ♂, Pu Toei NP, Phu Toeihill top/road, 14°57.32’N 99°26.972’E,  650 m,  1-7.vii.2008, Saunbua.L.leg., T3138.   Ubon Ratchathani: Pha Taem NP,  1 ♂ Tung Na Meaung Waterfall, 15°30.914’N 105°35.369’E,  120 m,  6-12.iii.2007, Porntip Tonsu& Bunlu Sapsirileg., T2144;  1 ♂ Wildflower field 1, 15°27.336’N 105°34.870’E,  232 m,  18-25.xi.2006, Sorawitand Thongdeeleg., T1060;  1 ♂ Firstlevel-Huay Sa-nhomwaterfall, 15°27.407’N 105°34.867’E,  230 m,  2-9.v.2007, Sorawit Mingmanleg., T2185 14°28.524’N 101°22.928’E, 757 m;  1 ♂ Dong Ngu How area,  5-12.xii.2006, Pong Sandowleg., T1303;  2 ♂ Cobrazone near Fireprotection office,  5-12.vi.2007, Pong Sandaoleg., T2221.   Petchaburi:  1 ♂ Kaeng Krachan NP, Panernthung/km27/water pump, 12°49.151’N 99°22.483’E,  970 m,  5-12.ix.2008, Sirichai& Chusakleg., T4377.   Nakhon Nayok: 1 ♂ Khao Yai NP, Naturetrail in secondary forest, 14°24.522’N 101°22.434’E,  750 m,  12-19.viii.2006, Pong Sandaoleg., T402.   Chanthaburi: Khao Khitchakut NP, NE/ Prabaht Unit, Suthida& Charoenchaileg.,  1 ♂,  50 m, 12°48.779’N 102°9.181’E,  107 m,  7-14.vii.2008, T2976,  2 ♂  100 m, 12°48.842’N 102°9.144’E,  203 m,  7-14.vii.2008, T2973;  Campground/ Prabaht Unit, 12°48.852’N 102°9.204’E,  99 m,  1 ♂  11-18.viii.2008, T2982,  1 ♂  28.vii-3.viii.2008, T2985. Prachuab Khiri Khan:  1 ♂, Khao Sam Roi Yot NP, Nursery, 12°7.58’N 99°57.478’E, Pan trap,  3-4.vii.2008, Amnad& Yaileg., T3039.  Surat Thani: Khao Sok NP, Pongphanleg.,  1 ♂, Klong Mog Unit, 8°53.725’N 98°39.025’E,  87 m,  31.vii-7.viii.2008, T3067;  Bang Huaraed, 8°54.555’N 98°30.522’E,  122 m,  1 ♂,  7-14.iv.2009, T4569,  1 ♂  10- 17.viii.2008, T3361.   Nakhon Si Thammarat: Namtok Yong NP,  1 ♂ Roadto Khao Mhen.  150 m, from Nern499, 8°16.959’N 99°39.149’E  499 m,  15-22.vii.2008, Samnaokan.S.leg., T3096;  1 ♂, Klong Jangwaterfall, 8°16.283’N 99°38.702’E,  154 m,  29.xii.2008-  5.i.2009, Samnaokan;S. leg., T4290;  Behindcampground lavatory, 8°10.434’N 99°44.508’E,  80 m, U.prai.K. leg.,  2 ♂  12-19.viii.2008, T3080,  1 ♂  29.vii-5.viii.2008, T3072,  1 ♂  19- 26.viii.2008, T3074. metasoma varying between dark castaneous with castaneous spots or fully castaneous; frontal with punctures small, shallow and irregularly sparse; vertex crest straight; lower tooth of mandible sub-angulate; last flagellomere with angulate apex; placoid sensillae elliptical; eye gibbous; scutellar groove slightly convex; ventral outer lobe of aedeagal dorsal body straight, varying between very short (apical margin of ventral pair not reaching basal region of ventral pair) and short (apical margin of ventral pair aligned with median region of ventral pair).   Remarks.  Dissomphalus verus  sp. nov.is similar to  D. brownibecause of the following characteristics: hypopygium with long and narrow median stalk, lateral margins slightly convergent posterad, posterior margin with angulate corners, dorsal outer lobe of aedeagal dorsal body with laminar process, ventral outer lobe of dorsal body short and arched; aedeagal ventral ramus as long as dorsal body, apex with triangular cross-section. However,  D. verus  sp. nov.differs from  D. brownias follows: lateral stalks of hypopygium triangular, apical region of median stalk with constriction, posterior margin slightly sinuous; base of paramere wider than basiparamere apex; base of aedeagal dorsal body wide, laminar process sickle-shaped with apical margin strongly concave; outer ventral lobe of dorsal body strongly arched with round apex.   Distribution. Thailand( Chaiyaphum, Mae Hong Son, Chiang Mai, Loei, Sakon Nakhon, Phitsanulok, Chaiyaphum, Suphanburi, Ubon Ratchathani, Petchaburi, Nakhon Nayok, Chanthaburi, Prachuab Khiri Khan, Surat Thaniand Nakhon Si Thammarat). 2006-10-01 2006-10-07 2006-10-01 T Pa Hin Ngam NP & Katae Sa-nog & Buakaw Adnafai 614 15.648666 Mixed 1 101.41795 26 27 T656 1 1 Chaiyaphum holotype [382,1014,763,788] Mae Hong Son 26 27 1 Mae Hong Son paratype 2008-01-27 2008-02-03 2008-01-27 T Kamkoon 19.3436 Huai Fai Kor 87 97.988335 Namtok Mae 26 27 T3496 1 1 Surin paratype 2006-08-30 2006-09-06 2006-08-30 T Doi Inthanon NP & Y. Areeluck 700 18.538517 Vachiratharn Falls 1 98.6008 26 27 T236 1 1 Chiang Mai paratype [151,375,871,896] Phu Ruea NP Loei 26 27 1 Loei paratype 2006-07-05 2006-07-12 2006-07-05 T Patikhom Tamtip 860 17.479534 Subhnonghin 1 101.35513 26 27 T310 1 1 Loei paratype [225,1381,908,932] 2007-06-05 2007-06-08 2007-06-05 T Patikhom Tumtip 691 17.463818 1 101.356 26 27 T2514 1 1 Loei paratype 2006-09-18 2006-09-25 2006-09-18 T 274 16.942816 1 101.81741 26 27 T950 1 1 Loei paratype [224,1411,980,1004] 2006-11-04 2006-11-11 2006-11-04 T 273 16.94315 Pong 1 101.70123 Dry 26 27 T792 1 1 Loei paratype 2006-09-25 2006-10-02 2006-09-25 T Sutin Glong-lasae 273 16.942749 Lam Huay Taad 1 101.81493 Bamboo forest 26 27 T954 1 1 Loei paratype 2006-08-16 2006-08-23 2006-08-16 T Sutin Khonglasae 270 16.863617 Koke Hin Ngam 1 101.84507 26 27 T486 1 1 Loei paratype 2007-02-13 2007-02-19 2007-02-13 T Wuthicahi 406 16.842333 Phakbung 1 101.69438 Forest 26 27 T1499 1 1 Loei paratype 2006-11-26 2006-12-02 2006-11-26 T Suthin Gong-lasae Road 265 16.815216 Na Noy Forest 1 101.7939 Ta Krong 26 27 T1079 1 1 Loei paratype Phu Phan NP Sakon Nakhon 26 27 1 Sakon Nakhon paratype 2006-09-16 2006-09-22 2006-09-16 T Winlon Khongnara 347 17.123583 Haew Sin 1 104.01965 Kam Hom 26 27 T616 1 1 Sakon Nakhon paratype [410,1431,1231,1255] Sailom Tongboonchai 344 17.122334 Sao Hi 1 104.01313 Nam Hom Waterfall 26 27 1 Sakon Nakhon paratype [151,443,1268,1292] 2007-03-04 2007-03-10 2007-03-04 T 26 27 T2368 2 2 Sakon Nakhon paratype [457,762,1268,1292] 2007-03-10 2007-03-17 2007-03-10 T 26 27 T2371 1 1 Sakon Nakhon paratype Winlon Kongnara 312 17.05905 Behind 1 103.9742 26 27 1 Sakon Nakhon paratype [328,612,1303,1327] 2006-07-08 2006-07-14 2006-07-08 T 26 27 T197 4 4 Sakon Nakhon paratype [626,954,1303,1327] 2006-07-27 2006-08-02 2006-07-27 T 26 27 T206 1 1 Sakon Nakhon paratype 2007-01-11 2007-01-17 2007-01-11 T 312 17.05905 North 1 103.9742 26 27 T1519 1 1 Sakon Nakhon paratype 2007-01-11 2007-01-17 2007-01-11 T 322 17.058683 West 1 103.97417 26 27 T1520 1 1 Sakon Nakhon paratype 2006-07-15 2006-07-21 2006-07-15 T Behind 26 27 T202 1 1 Sakon Nakhon paratype [368,681,1412,1436] Behind national park 26 27 1 Sakon Nakhon paratype [693,978,1412,1436] 2007-01-17 2007-01-23 2007-01-17 T 26 27 T1524 3 3 Sakon Nakhon paratype [991,1276,1412,1436] 2007-01-23 2007-01-30 2007-01-23 T 26 27 T1527 1 1 Sakon Nakhon paratype Manop Ngoyjansri & Chatree Cheaukamjan 295 16.927317 Channel 1 104.17913 Phu Pha Yon 26 27 1 Sakon Nakhon paratype 2006-07-05 2006-07-11 2006-07-05 T 26 27 T289 3 3 Sakon Nakhon paratype [368,661,1484,1508] 2006-07-17 2006-07-23 2006-07-17 T 26 27 T295 1 1 Sakon Nakhon paratype 2006-07-17 2006-07-23 2006-07-17 T 295 16.92415 Deciduous 1 104.17928 26 27 T297 1 1 Sakon Nakhon paratype 2006-11-19 2006-11-25 2006-11-19 T Sailom Tongboonchai 289 16.911884 Huay Nam Pung Dam 1 103.9049 Forest 26 27 T1101 1 1 Sakon Nakhon paratype Pongpitak Pranee Thung Salaeng Luang 481 16.877367 Gang Sopa 1 100.82402 Mixed 26 27 1 Phitsanulok paratype 2006-10-16 2006-10-17 2006-10-16 T 26 27 T756 3 3 Phitsanulok paratype [381,661,1628,1652] 2006-11-04 2006-11-12 2006-11-04 T 26 27 T771 1 1 Phitsanulok paratype [674,1379,1628,1652] 2006-10-28 2006-11-04 2006-10-28 T 501 16.8744 1 100.82775 26 27 T769 1 1 Phitsanulok paratype 2006-08-18 2006-08-25 2006-08-18 T 557 16.844017 Moist 1 100.88157 26 27 T569 2 2 Phitsanulok paratype 2006-09-18 2006-09-25 2006-09-18 T 481 16.842716 Mixed 1 100.86262 26 27 T930 1 1 Phitsanulok paratype 486 16.83795 Dry Evergreen forest 1 100.88195 26 27 1 Phitsanulok paratype [187,846,1735,1759] 2007-04-01 2007-04-08 2007-04-01 T Pongpitak & Pranee & Sathit 26 27 T2394 2 2 Phitsanulok paratype [858,1410,1735,1759] 2006-09-25 2006-10-02 2006-09-25 T Pongpitak Pranee 26 27 T932 1 1 Phitsanulok paratype 2007-04-08 2007-04-15 2007-04-08 T Pongpitak & Pranee & Sathit 481 16.842716 Deciduous forest 1 100.86262 26 27 T2398 1 1 Phitsanulok paratype [151,1065,1807,1832] Khao Kho NP & Somchai Chatchumnan & Saink Singtong 520 Nursery 26 27 1 Phitsanulok paratype 2006-10-12 2006-10-19 2006-10-12 T 16.87635 1 101.13433 26 27 T808 1 1 Phitsanulok paratype [394,993,1844,1868] 2006-10-12 2006-10-19 2006-10-12 T 16.876217 1 101.13461 26 27 T806 1 1 Phitsanulok paratype 2006-10-26 2006-11-02 2006-10-26 T 16.876217 1 101.13461 26 27 T812 1 1 Phitsanulok paratype [348,1233,1879,1904] 2006-10-05 2006-10-12 2006-10-05 T 520 16.876133 Savana 1 101.13506 26 27 T804 2 2 Phitsanulok paratype 2007-03-01 2007-03-08 2007-03-01 T 921 16.72825 Checkpoint 1 101.563286 26 27 T2285 1 1 Phitsanulok paratype 168 27 28 16.659817 Mix 1 101.13538 26 27 1 1 Phitsanulok paratype [391,696,151,175] 2007-02-26 2007-03-05 2007-02-26 T 27 28 T1610 1 1 Phitsanulok paratype [707,996,151,175] 2007-02-19 2007-02-26 2007-02-19 T 27 28 T1607 1 1 Phitsanulok paratype 2006-07-12 2006-07-19 2006-07-12 T 230 16.659166 1 101.13538 27 28 T167 1 1 Phitsanulok paratype [494,1348,188,212] 242 16.542683 Ta Phol river 1 101.04131 Mixed 27 28 1 Phitsanulok paratype 2006-11-05 2006-11-12 2006-11-05 T 27 28 T968 1 1 Phitsanulok paratype [364,655,223,247] 2006-11-12 2006-11-19 2006-11-12 T 27 28 T971 1 1 Phitsanulok paratype [669,967,223,247] 2006-11-19 2006-11-26 2006-11-19 T 27 28 T974 2 2 Phitsanulok paratype [981,1294,223,247] 2006-11-26 2006-12-02 2006-11-26 T 27 28 T977 1 1 Phitsanulok paratype 2006-12-26 2007-01-02 2006-12-26 T 560 16.542433 Mixed 1 101.04169 27 28 T1189 1 1 Phitsanulok paratype Nam Pong NP & Khamphol Jaidee Khonkaen 27 28 1 Phitsanulok paratype [519,1270,295,319] 2006-07-19 2006-07-26 2006-07-19 T 218 16.620016 1 102.574684 27 28 T113 2 2 Phitsanulok paratype [405,679,332,356] 2006-07-05 2006-07-12 2006-07-05 T 27 28 T115 2 2 Phitsanulok paratype [690,977,332,356] 2006-07-11 2006-07-18 2006-07-11 T 27 28 T105 1 1 Phitsanulok paratype 2006-07-05 2006-07-12 2006-07-05 T 27 28 T106 1 1 Phitsanulok paratype [398,700,367,391] 2006-07-12 2006-07-19 2006-07-12 T 27 28 T110 1 1 Phitsanulok paratype [716,1016,367,391] 2006-07-19 2006-07-26 2006-07-19 T 27 28 T111 1 1 Phitsanulok paratype 2006-11-05 2006-11-12 2006-11-05 T 674 16.01765 Sab 1 101.976715 Dry 27 28 T1137 1 1 Phitsanulok paratype [429,1226,439,463] 648 16.0132 Sab Somboon forest 1 101.97453 Lawn 27 28 1 Phitsanulok paratype 2006-11-05 2006-11-12 2006-11-05 T 27 28 T1136 2 2 Phitsanulok paratype [237,553,476,500] 2006-11-26 2006-12-03 2006-11-26 T 27 28 T1145 1 1 Phitsanulok paratype 2006-06-03 2006-06-09 2006-06-03 T Protection station Forest 27 28 T2535 4 4 Phitsanulok paratype [747,1092,511,535] Chaiyapoom forest 27 28 1 Phitsanulok paratype [1104,1410,511,535] 2006-12-19 2006-12-23 2006-12-19 T 27 28 T1374 1 1 Phitsanulok paratype 2006-12-26 2007-01-02 2006-12-26 T 27 28 T1377 1 1 Phitsanulok paratype [520,1200,548,572] 250 15.98395 Dry Dipterocarp Forest 1 102.03505 27 28 1 Phitsanulok paratype 2006-06-19 2006-06-26 2006-06-19 T Boonnak Nongpeu 27 28 T22 1 1 Phitsanulok paratype [494,1011,583,608] 2006-06-21 2006-06-28 2006-06-21 T Leng Chanteab 27 28 T32 1 1 Phitsanulok paratype 2006-06-23 2006-06-26 2006-06-23 T Lumyai Ittichan 27 28 T27 1 1 Phitsanulok paratype [266,729,620,644] 2006-06-25 2006-06-27 2006-06-25 T M. Sharkey 27 28 T10 1 1 Phitsanulok paratype [741,1253,620,644] 2006-06-26 2006-06-29 2006-06-26 T Lumyai Ittichan 27 28 T30 1 1 Phitsanulok paratype [483,1377,655,679] 2006-07-26 2006-08-07 2006-07-26 T 305 15.979517 Near 1 102.03995 27 28 T227 1 1 Phitsanulok paratype 290 15.978717 Phu 1 102.037186 27 28 1 Phitsanulok paratype [681,964,692,716] 2006-07-05 2006-07-12 2006-07-05 T 27 28 T220 1 1 Phitsanulok paratype [977,1274,692,716] 2006-07-12 2006-07-19 2006-07-12 T 27 28 T223 1 1 Phitsanulok paratype 257 15.9724 Nursery 1 102.03615 27 28 1 Phitsanulok paratype [717,1020,727,751] 2006-08-05 2006-08-12 2006-08-05 T 27 28 T547 2 2 Phitsanulok paratype [1038,1370,727,751] 2007-02-26 2007-03-02 2007-02-26 T 27 28 T1737 1 1 Phitsanulok paratype 2006-10-12 2006-10-19 2006-10-12 T 297 15.958667 Pa Eang 1 101.907364 Entrance 27 28 T681 1 1 Phitsanulok paratype 2007-03-19 2007-03-26 2007-03-19 T 245 15.9411335 Tat Fah 1 102.09758 Water 27 28 T2354 2 2 Phitsanulok paratype 242 15.94105 Tat Fah 1 102.09922 Streamside 27 28 1 Phitsanulok paratype [410,726,836,860] 2007-03-19 2007-03-26 2007-03-19 T 27 28 T2352 1 1 Phitsanulok paratype [743,1072,836,860] 2007-03-26 2007-04-02 2007-03-26 T 27 28 T2355 2 2 Phitsanulok paratype 2007-03-19 2007-03-26 2007-03-19 T 15.941016 Tat Fah 1 102.09925 Oficer 27 28 T2353 1 1 Phitsanulok paratype 2006-10-19 2006-10-25 2006-10-19 T Thepana 604 15.649134 Thepana 1 101.427086 Savana 27 28 T666 4 4 Phitsanulok paratype [278,1256,943,967] 2006-10-01 2006-10-07 2006-10-01 T 614 15.648666 Forest 1 101.41795 Mixed 27 28 T656 2 2 Phitsanulok paratype 605 15.6480665 Dry 1 101.430664 27 28 1 Phitsanulok paratype [649,916,980,1004] 2006-10-07 2006-10-13 2006-10-07 T 27 28 T661 1 1 Phitsanulok paratype [930,1211,980,1004] 2006-10-13 2006-10-19 2006-10-13 T 27 28 T664 2 2 Phitsanulok paratype 2006-10-19 2006-10-25 2006-10-19 T 27 28 T667 1 1 Phitsanulok paratype [645,1295,1015,1040] 2006-07-06 2006-07-12 2006-07-06 T Tung Dok Grajeaw 27 28 T330 1 1 Phitsanulok paratype 2007-01-06 2007-01-07 2007-01-06 T Thung Dok Kra Jeow in Dry 27 28 T1451 2 2 Phitsanulok paratype 2006-07-06 2006-07-12 2006-07-06 T Tung Dok Grajeaw 27 28 T329 1 1 Phitsanulok paratype [750,1431,1087,1111] 2007-01-13 2007-01-19 2007-01-13 T Thung Dok Kra Jeow Car 27 28 T1459 1 1 Phitsanulok paratype 2007-02-03 2007-02-04 2007-02-03 T 27 28 T1636 1 1 Phitsanulok paratype [352,793,1159,1183] 2007-02-19 2007-02-25 2007-02-19 T 27 28 T1652 1 1 Phitsanulok paratype 2007-02-01 2007-02-07 2007-02-01 T 698 15.634983 Dipterocarp forest 1 101.39868 27 28 T1644 2 2 Phitsanulok paratype [413,1433,1196,1220] 2007-03-09 2007-03-15 2007-03-09 T 668 15.626917 Lan Hin Nor 1 101.3906 Nature 27 28 T2328 1 1 Phitsanulok paratype 2008-07-01 2008-07-07 2008-07-01 T Pu Toei NP & Saunbua. L. 650 14.955334 Phu Toei 9 99.44953 27 28 T3138 1 1 Suphan Buri paratype [294,686,1267,1292] Pha Taem NP Ubon Ratchathani 27 28 1 Ubon Ratchathani paratype 2007-03-06 2007-03-12 2007-03-06 T Porntip Tonsu & Bunlu Sapsiri 120 15.515233 Tung Na Meaung Waterfall 1 105.589485 27 28 T2144 1 1 Ubon Ratchathani paratype 2006-11-18 2006-11-25 2006-11-18 T Sorawit & Thongdee 232 15.4556 Wild 1 105.58117 27 28 T1060 1 1 Ubon Ratchathani paratype 2007-05-02 2007-05-09 2007-05-02 T Sorawit Mingman 230 15.456783 Sa-nhom 1 105.581116 First 27 28 T2185 1 1 Ubon Ratchathani paratype 2006-12-05 2006-12-12 2006-12-05 T Pong Sandow Dong Ngu How area 27 28 T1303 1 1 Ubon Ratchathani paratype 2007-06-05 2007-06-12 2007-06-05 T Pong Sandao Fire Cobra 27 28 T2221 2 2 Ubon Ratchathani paratype [463,599,1447,1472] Petchaburi 27 28 1 Ubon Ratchathani paratype 2008-09-05 2008-09-12 2008-09-05 T Kaeng Krachan NP & Sirichai & Chusak 970 12.819183 Panernthung 1 99.37472 27 28 T4377 1 1 Ubon Ratchathani paratype 2006-08-12 2006-08-19 2006-08-12 T Khao Yai NP & Pong Sandao 750 14.4087 Nature 1 101.3739 27 28 T402 1 1 Nakhon Nayok paratype Khao Khitchakut NP & Prabaht Unit & Suthida & Charoenchai Chanthaburi 27 28 1 Chanthaburi paratype 2008-07-07 2008-07-14 2008-07-07 T 50 12.8129835 1 102.153015 27 28 T2976 1 1 Chanthaburi paratype [405,1183,1591,1615] 2008-07-07 2008-07-14 2008-07-07 T 100 12.8140335 1 102.1524 27 28 T2973 2 2 Chanthaburi paratype 99 12.8142 Prabaht Unit 1 102.1534 Campground 27 28 1 Chanthaburi paratype [617,940,1628,1652] 2008-08-11 2008-08-18 2008-08-11 T 27 28 T2982 1 1 Chanthaburi paratype [956,1305,1628,1652] 2008-07-28 2008-08-03 2008-07-28 T 27 28 T2985 1 1 Chanthaburi paratype 2008-07-03 2008-07-04 2008-07-03 T Khao Sam Roi Yot NP & Amnad & Yai 12.126333 Nursery 9 99.95797 27 28 T3039 1 1 Chanthaburi paratype 2008-07-31 2008-08-07 2008-07-31 T 87 8.895416 Klong Mog Unit 1 98.65041 27 28 T3067 1 1 Chanthaburi paratype [455,1015,1735,1760] 122 8.90925 Bang Huaraed 1 98.5087 27 28 1 Chanthaburi paratype [1030,1327,1735,1759] 2009-04-07 2009-04-14 2009-04-07 T 27 28 T4569 1 1 Chanthaburi paratype 2008-08-10 2008-08-17 2008-08-10 T 27 28 T3361 1 1 Chanthaburi paratype [388,884,1771,1796] Namtok Yong NP Nakhon Si Thammarat 27 28 1 Nakhon Si Thammarat paratype 2008-07-15 2008-07-22 2008-07-15 T Samnaokan. S. Road 150 8.28265 Nern 1 99.65248 Khao Mhen. 27 28 T3096 1 1 Nakhon Si Thammarat paratype 2008-12-29 2009-01-05 2008-12-29 T Samnaokan 154 8.271383 Klong Jang 1 99.645035 27 28 T4290 1 1 Nakhon Si Thammarat paratype 80 8.1739 Behind 1 99.7418 27 28 1 Nakhon Si Thammarat paratype [683,994,1879,1903] 2008-08-12 2008-08-19 2008-08-12 T 27 28 T3080 2 2 Nakhon Si Thammarat paratype [1005,1338,1879,1903] 2008-07-29 2008-08-05 2008-07-29 T 27 28 T3072 1 1 Nakhon Si Thammarat paratype 2008-08-19 2008-08-26 2008-08-19 T 27 28 T3074 1 1 Nakhon Si Thammarat paratype