Revision of the genus Draconarius Ovtchinnikov 1999 (Agelenidae: Coelotinae) in Yunnan, China, with an analysis of the Coelotinae diversity in the Gaoligongshan Mountains Wang, XIN-PING Griswold, CHARLES E. Miller, JEREMY A. Zootaxa 2010 2593 1 127 92ZRN Wang Wang Arachnida Agelenidae Draconarius CoL Animalia Araneae 81 85 Arthropoda species pseudobrunneus  (Figs 361-406, 548)    Draconarius pseudobrunneus Wang2003: 544, figs 52A-B ( female holotypeand  paratypesfrom  Gongshan County, Yunnan, China, in HNUand  CAS, examined).  Additional material examined: CHINA: Yunnan: Lushui County: 1♀, Feng Xue Yakou, E side on Pianma Road, N25.97288°, E98.68336°,  3150 m, night collectingon buildings, cliffs, roadcut,  May 17, 2005, C. Griswold( HNU, CASENT9022263);  2♀, Pianma Township, Feng Xue Yakou, N26.00949°, E98.61704°,  3142 m, along the road,  May17, 2005, K.J. Guo( CAS, CASENT9022073);  1♀, Pianma Township, Yaojiaping Forest Station, N25.96911°, E98.70713°,  2586 m, roadside,  May 18, 2005, K.J. Guo( CAS, CASENT9022057);  1♀, Pianma Township, Feng Xue Yakou, N25.97814°, E98.67508°,  3088 m, roadside,  May 19, 2005, K.J. Guo( HNU, CASENT9022040).  Gongshan County: 2♀, Cikai Township, 41 kmW of Gongshan on Dulong Valley Roadat Dabadi both sides, 27°27'N, 98°37'E,  3100 m,  September 23, 2002, H.M. Yan( CAS, CASENT9025836);  1♀, Cikai Township, 41 kmW of Gongshan on Dulong Valley Road at Dabadi, N27.79655°, E098.50562°,  3000 m,  September 27 to October 6, 2002, D. Kavanaugh, P.E. Marek, D.Z. Dong, & X.C. Liang( HNU, CASENT9025844);  3♀, Cikai Township, Dabadi troops placealong river banks, 3030m N27°47', E98°30',  3045 m,  October 2-3, 2002, night,  October 8-10, 2002, H.M. Yan( CAS, CASENT9025843);  1♀, Cikai Township, 41 kmW of Gongshan on Dulong Valley Road at Dabadi, N27.79655°, E098.50562°,  3000 m,  September 27 - October 6, 2002, D. Kavanaugh, P.E. Marek, D.Z. Dong, & X.C. Liang( HNU, CASENT9025834);  1♀, Cikai Township, 41 kmW of Gongshan on Dulong Valley Road at Dabadi both sides, N27°47', E98°37',  3100 m,  September 30, 2002, H.M. Yan( CAS, CASENT9025841).  Fugong County: 1♀, Lishadi subdistrict, Gaoligongshan, Shibali, N27.16784°, E98.77782°,  2582 m,  May 2, 2004, H.M. Yan, G.X. Peng( CAS, CASENT9020627);  1♀, Gaoligongshan, Shibali Forest Station, in moist stream gully, N27.16894°, E98.77205°,  2650 m,  May 9, 2004, C. Griswold& H.M. Yan( CAS, CASENT9019906);  1♂, Gaoligongshan, Shibali forest station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020460);  1♂, Gaoligongshan, Shibali Forest Station, in moist stream gully, N27.16894°, E98.77205°,  2650 m,  May 9, 2004, C. Griswold& H.M. Yan( HNU, CASENT9019905);  1♀, Gaoligongshan, 0.5 kmradius of Shibali Forest Station, N27.16519°, E98.77891°,  2525 m, searching at night in forest,  May 2-5, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020026);  1♀, Lishadi subdistrict, Gaoligongshan, Shibali, N27.16784°, E98.77782°,  2582 m,  May 2, 2004, H.M. Yan, G.X. Peng( HNU, CASENT9020628);  1♀, Yaping subdistrict, Gaoligongshan, Maxidi, 7.2 kmabove Shibali, N27.16784°, E98.77782°,  3059 m,  May 4, 2004, H.M. Yan, G.X. Peng( CAS, CASENT9020583);  1♀, Gaoligongshan, Shibali Forest Station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( HNU, CASENT9020459);  1♀, Gaoligongshan, 0.4 kmSSE of Shibali Forest Station, beneath wood in good forest, N27.16337°, E98.78208°,  2475 m,  May 5, 2004, C. Griswold& D. Kavanaugh( HNU, CASENT9019963);  1♀, Gaoligongshan, 0.4 kmSSE of Shibali Forest Station, beneath wood in good forest, N27.16337°, E98.78208°,  2475 m,  May 5, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9019964);  10♂, Gaoligongshan, Shibali Forest Station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020464);  1♂, Shibali Forest Station, N27.16636°, E98.77667°,  2563 m, sifting leaf litterin good forest,  May 4, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020064);  10♂, Gaoligongshan, Shibali Forest Station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( HNU, CASENT9020465);  1♂, Gaoligongshan, Shibali Forest Station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( HNU, CASENT9020462);  1♂, Gaoligongshan, 4 kmW of Shibali Forest Station, N27.17746°, E98.75527°,  2820 m, night collectingalong roadcut,  May 3, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9019925);  1♂, Gaoligongshan, 10.1-11.5 road km from Shibali on Yaping Road, N27.20049°, E98.71354°- N27.20676°, E98.71763,  3225-3290m, night collectingalong road,  May 8, 2004, C. Griswold& D. Kavanaugh( HNU, CASENT9020034);  4♂, Gaoligongshan, Shibali Forest Station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020466);  3♂, Gaoligongshan, Shibali forest station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( HNU, CASENT9020467);  1♂, Gaoligongshan, 0.5 kmradius of Shibali Forest Station, N27.16519°, E98.77891°,  2525m, general collectionin disturbed areas and forest,  May 1-11, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020670);  1♂, Gaoligongshan, 6.18 kmW of Shibali, turning rocksin open meadow along stream, N27.18413°, E98.72024°,  3100 m,  May 7, 2004, C. Griswold& D. Kavanaugh( HNU, CASENT9020706);  2♂, Gaoligongshan, Shibali Forest Station, N27.16636°, E98.77667°,  2563 m, pitfall trapsin good forest,  May 3-11, 2004, C. Griswold& D. Kavanaugh( CAS, CASENT9020463);  1♀, Gaoligongshan (label missing)( CAS, CASENT9025845).    FIGURES355-360. Draconarius pseudoagrestis sp. nov., male holotypefrom Tengchong County, Bawan village ( HNU, CASENT9025789). 355-357. Palp (prolateral, ventral, retrolateral); 358. Habitus, dorsal; 359. Eyes, fronto-dorsal, with outlines redrawn with black lines; 360. Labium, ventral ( L/ W=1.25).  Diagnosis: D. pseudobrunneusis similar to D. incertus Wang2003 and D. latusincertus sp. nov.in having similar epigynum and palp, but can be distinguished from D. latusincertus sp. nov.by the small body size, the small, anteriorly extending conductor in the male (Figs 363-364, 367-369), and the slightly anteriorly extending spermathecae in the female; from D. incertusby the more or less long patellar apophysis and the smooth posterior margin of embolus in the male, and anteriorly extending spermathecae in the female (Figs 361-362, 365, 375-406).    FIGURE361-364. Draconarius pseudobrunneus Wang2003, female holotype(361-362) from Gongshan County, Danzhu He ( HNU) and male (363-364) from Fugong County, Shibali Forest Station ( CAS, CASENT9020464). 361-362. Epigynum (ventral, dorsal); 363-364. Palp (ventral, retrolateral).  Description: Female. See Wang (2003). Photos of habitus, eyes, labium and epigynum are provided in this study (Figs 365-366, 373-374, 375-406). Male (one male of CASENT9020464 measured). The male is here described for the first time. Medium sized Coelotinae, total length 6.28 (Fig. 370). Dorsal shield of prosoma 3.28 long, 2.00 wide; opisthosoma 3.00 long, 2.15 wide. AMEsmallest, half the size of ALE; ALE largest, PLE slightly smaller than ALE, PME slightly smaller than PLE ( AME0.06, ALE0.14, PME0.13, PLE0.11); anterior eyes close together; AMEseparated from each other by half of their diameter, from ALE by of AMEdiameter; PME separated from each other by Ά of their diameter, from PLE by half of PME diameter (AME-AME 0.03, AME-ALE 0.02, PME-PME 0.04, PME-PLE 0.07, AME-PME 0.06) (Fig. 371). Labium length and width subequal ( L/ W=1.0) (Fig. 372). Chelicerae with 3 promarginal and 2 retromarginal teeth. Palp with patellar apophysis moderately long, slender; RTA slightly less than or about half of tibial length, with moderately protruding distal end; lateral tibial apophysis large, distinctly separated from RTA; cymbial furrow small, less than half of cymbial length; conductor moderately long, with a slightly bifurcate apex, a small basal lamella, and a dorsal apophysis; median apophysis simple, not spoon-shaped; embolus short, filiform, prolateral in origin (Figs 363-364, 367-369).  Distribution: China (Yunnan: Lushui, Fugong, Gongshan) (Fig. 548). 923903159 HNU China Gongshan County 1 1 Yunnan holotype 923903177 HNU China Gongshan County Yunnan paratype 923903163 CAS China Gongshan County Yunnan paratype 923903158 2005-05-17 night collecting HNU C. Griswold China Lushui County 3150 m 25.97288 Feng Xue Yakou, E side on Pianma Road 0 98.68336 CASENT 9022263 1 1 Yunnan Other material 923903140 2005-05-17 CAS K. J. Guo China Lushui County 3142 m 26.00949 Pianma Township, Feng Xue Yakou 0 98.61704 CASENT 9022073 2 2 Yunnan Other material 923903165 2005-05-18 CAS K. J. Guo China Lushui County 2586 m 25.96911 Pianma Township, Yaojiaping Forest Station 0 98.70713 CASENT 9022057 1 1 Yunnan Other material 923903155 2005-05-19 HNU K. J. Guo China Lushui County 3088 m 25.97814 Pianma Township, Feng Xue Yakou 0 98.67508 CASENT 9022040 1 1 Yunnan Other material 923903162 2002-09-23 CAS H. M. Yan China Gongshan County 3100 m 27.45 Cikai Township, 41 km W of Gongshan on Dulong Valley Road at Dabadi both sides 98.62 CASENT 9025836 2 2 Yunnan Other material 923903169 2002-09-27 2002-10-06 2002-09-27 HNU D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Gongshan County 3000 m 27.79655 Cikai Township, 41 km W of Gongshan on Dulong Valley Road at Dabadi 0 98.50562 CASENT 9025844 1 1 Yunnan Other material 923903150 2002-10-02 2002-10-10 2002-10-02 CAS H. M. Yan China Gongshan County 3030 - 3045 m 27.783333 Cikai Township, Dabadi troops place 1 98.5 CASENT 9025843 3 3 Yunnan Other material 923903168 2002-09-27 2002-10-06 2002-09-27 HNU D. Kavanaugh, P. E. Marek, D. Z. Dong, X. C. Liang China Gongshan County 3000 m 27.79655 Cikai Township, 41 km W of Gongshan on Dulong Valley Road at Dabadi 0 98.50562 CASENT 9025834 1 1 Yunnan Other material 923903154 2002-09-30 CAS H. M. Yan China Gongshan County 3100 m 27.783333 Cikai Township, 41 km W of Gongshan on Dulong Valley Road at Dabadi both sides 1 98.61667 CASENT 9025841 1 1 Yunnan Other material 923903186 2004-05-02 CAS H. M. Yan, G. X. Peng China Fugong County 2582 m 27.16784 Lishadi subdistrict, Gaoligongshan, Shibali 0 98.77782 CASENT 9020627 1 1 Yunnan Other material 923903182 2004-05-09 CAS C. Griswold, H. M. Yan China Fugong County 2650 m 27.16894 Gaoligongshan, Shibali Forest Station 0 98.77205 CASENT 9019906 1 1 Yunnan Other material 923903157 2004-05-03 2004-05-11 2004-05-03 pitfall traps CAS C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali forest station 0 98.77667 CASENT 9020460 1 1 Yunnan Other material 923903152 2004-05-09 HNU C. Griswold, H. M. Yan China Fugong County 2650 m 27.16894 Gaoligongshan, Shibali Forest Station 0 98.77205 CASENT 9019905 1 1 Yunnan Other material 923903172 2004-05-02 2004-05-05 2004-05-02 searching at night in forest CAS C. Griswold, D. Kavanaugh China Fugong County 2525 m 27.16519 Gaoligongshan, 0.5 km radius of Shibali Forest Station 0 98.77891 CASENT 9020026 1 1 Yunnan Other material 923903185 2004-05-02 HNU H. M. Yan, G. X. Peng China Fugong County 2582 m 27.16784 Lishadi subdistrict, Gaoligongshan, Shibali 0 98.77782 CASENT 9020628 1 1 Yunnan Other material 923903171 2004-05-04 CAS H. M. Yan, G. X. Peng China Fugong County 3059 m 27.16784 Yaping subdistrict, Gaoligongshan, Maxidi, 7.2 km above Shibali 0 98.77782 CASENT 9020583 1 1 Yunnan Other material 923903170 2004-05-03 2004-05-11 2004-05-03 pitfall traps HNU C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali Forest Station 0 98.77667 CASENT 9020459 1 1 Yunnan Other material 923903175 2004-05-05 HNU C. Griswold, D. Kavanaugh China Fugong County 2475 m 27.16337 Gaoligongshan, 0.4 km SSE of Shibali Forest Station 0 98.78208 CASENT 9019963 1 1 Yunnan Other material 923903166 2004-05-05 CAS C. Griswold, D. Kavanaugh China Fugong County 2475 m 27.16337 Gaoligongshan, 0.4 km SSE of Shibali Forest Station 0 98.78208 CASENT 9019964 1 1 Yunnan Other material 923903183 2004-05-03 2004-05-11 2004-05-03 pitfall traps CAS C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali Forest Station 0 98.77667 CASENT 9020464 10 10 Yunnan Other material 923903167 2004-05-04 sifting leaf litter CAS C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Shibali Forest Station 0 98.77667 CASENT 9020064 1 1 Yunnan Other material 923903180 2004-05-03 2004-05-11 2004-05-03 pitfall traps HNU C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali Forest Station 0 98.77667 CASENT 9020465 10 10 Yunnan Other material 923903184 2004-05-03 2004-05-11 2004-05-03 pitfall traps HNU C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali Forest Station 0 98.77667 CASENT 9020462 1 1 Yunnan Other material 923903201 2004-05-03 night collecting CAS C. Griswold, D. Kavanaugh China Fugong County 2820 m 27.17746 Gaoligongshan, 4 km W of Shibali Forest Station 0 98.75527 CASENT 9019925 1 1 Yunnan Other material 923903188 2004-05-08 night collecting HNU C. Griswold, D. Kavanaugh China Fugong County 33258 m 27.20 Gaoligongshan, 10.1 - 11.5 road km from Shibali on Yaping Road 98.71 CASENT 9020034 1 1 Yunnan Other material 923903178 2004-05-03 2004-05-11 2004-05-03 pitfall traps CAS C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali Forest Station 0 98.77667 CASENT 9020466 4 4 Yunnan Other material 923903164 2004-05-03 2004-05-11 2004-05-03 pitfall traps HNU C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali forest station 0 98.77667 CASENT 9020467 3 3 Yunnan Other material 923903189 2004-05-01 2004-05-11 2004-05-01 general collection CAS C. Griswold, D. Kavanaugh China Fugong County 2525 m 27.16519 Gaoligongshan, 0.5 km radius of Shibali Forest Station 0 98.77891 CASENT 9020670 1 1 Yunnan Other material 923903176 2004-05-07 turning rocks HNU C. Griswold, D. Kavanaugh China Fugong County 3100 m 27.18413 Gaoligongshan, 6.18 km W of Shibali 0 98.72024 CASENT 9020706 1 1 Yunnan Other material 923903187 2004-05-03 2004-05-11 2004-05-03 pitfall traps CAS C. Griswold, D. Kavanaugh China Fugong County 2563 m 27.16636 Gaoligongshan, Shibali Forest Station 0 98.77667 CASENT 9020463 2 2 Yunnan Other material 923903204 CAS China Fugong County Gaoligongshan (label missing) CASENT 9025845 1 1 Yunnan Other material