Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae) Edward, Karen L. Harvey, Mark S. Bulletin of the American Museum of Natural History 2014 2014-06-06 2014 389 1 144 3Q66S Edward & Harvey, 2014 Edward & Harvey 2014 [168,462,1630,1653] Arachnida Oonopidae Ischnothyreus Animalia Araneae 129 129 Arthropoda species pterodactyl sp. nov.    Superbus, false summit, 28.23333 °S, 152.46670 °E,  1200 m(  12 Mar–13 June 1990 T.B. Churchill, R.J. Raven), deposited in QM( ³ holotype: QM S16568, PBI_OON 21959; ♀ allotype: QM S16568, PBI_ OON 5609).  ETYMOLOGY: The specific epithet is a noun in apposition.  DIAGNOSIS: This species can be easily recognized by the extremely elongate, obtusely bent embolic region of the male palp (fig. 73A, B) and the basal process of the fang consisting of numerous heavily sclerotized lobes (fig. 73C, D). The female possesses a heavily sclerotized circular epigynal atrium, which appears to be fused to the overhanging process and the horizontal sclerotization anterior to the epigynal atrium is less sclerotized than  I. rixi, sp. nov.(figs. 72I, 73E).  MALE (PBI_OON 21959, fig. 72 A–C, G, H; paratype(PBI_OON 05631), fig. 73A–D). Total length 1.73. CEPHALOTHORAX: Carapacebrown, broadly oval in dorsal view, pars cephalica slightly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. Clypeusmargin unmodified, curved downward in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes:ALE largest, ALE circular, PME oval, PLE circular; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternumas long as wide, pale orange, uniform; setae dark, evenly scattered. Chelicerae, endites, and labium pale orange. Chelicerae slightly divergent, anterior face unmodified; promargin with one larger denticle; fang shape normal, basal process consisting of numerous heavily sclerotized lobes, anterior lobe with distal ridges; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, same as sternum in sclerotization; with six or more setae on anterior margin. Endites distally not excavated, anteromedian tip with one strong, toothlike projection, same as sternum in sclerotization. ABDOMEN: Ovoid; dorsum soft portions pale orange. Book lung covers elliptical. Dorsal scutum brown, covering more than 3/4 of abdomen, no soft tissue visible from above, middle surface finely reticulate, sides finely reticulate. Epigastric scutum small lateral sclerites absent. Postepigastric scutum orange-brown, covering about 2/3 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Orange-brown, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter; patella about as long as femur; cymbium dark redbrown; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region extremely narrow and elongate, obtusely bent, tip rounded, transparent membranous tissues visible medially (fig. 73A, B).  Fig. 72.  Ischnothyreus pterodactyl, sp. nov.Holotype male (PBI_OON 00021959): A.habitus, dorsal view; B.habitus, ventral view; C.habitus, lateral view; G.carapace, anterior view; H.sternum, ventral view. Allotype female (PBI_OON 00005609): D.habitus, dorsal view; E.habitus, ventral view; F.habitus, lateral view; I.Epigynum, ventral view.  Fig. 73.  Ischnothyreus pterodactyl, sp. nov.Paratype male (PBI_OON 00005631): A.left palp, prolateral view; B.left palp, retrolateral view; C.left chelicerae, anterior view; D.left chelicerae, posterior view. Paratype female (PBI_OON 00005639): E.epigynum, ventral view. Scale lines 5 0.1 mm. FEMALE (PBI_OON 5609, fig. 72D–E, I; paratype(PBI_OON 5639), fig. 73E). Total length 1.98. CEPHALOTHORAX: Carapaceanteriorly narrowed to 0.49 times its maximum width or less. ABDOMEN: Dorsal scutum covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Leg spination: femora: I p0-2-2; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: strong horizontal scleriotization anterior to rounded epigynal atrium clearly joins posteriorly directed lateral apodemes; heavy sclerotization around epigynal atrium circular, thick; overhanging process slightly triangular, sides appear fused to postepigastric scutum; convoluted duct thicker than apodemes (figs. 72I, 73E).  OTHER MATERIAL EXAMINED: AUS- TRALIA:  New South Wales: Beech Picnic Area, Wiangaree,  1000 m, 28.37300 °S, 153.10070 °E,  Apr. 22, 2002(G. Monteith, QM S69613, PBI_OON 22217), 3 ♀;  Border Ranges National Park, Brindle Creek, rainforest alongside Creek, leaf litter, 28.37277 °S, 153.06916 °E,  May 9, 2007(K. Edwardand K. Pitz, QM S96010, PBI_OON 25729), 1 ♀;  Bruxner Pk, via Coffs Harbour, rainforest, leaf litter,  150 m, 30.23333 °S, 153.10000 °E,  Mar. 22, 1980(G. Monteith, QM S16054), 1 ♀;  Bruxner Park,  150 m, 30.23333 °S, 153.10000 °E,  Mar. 22–Nov. 13, 1980(G. Monteith, QM S16045, S16151), 7 ³, 9 ♀;  Bruxner Park,  150 m, 30.23333 °S, 153.10000 °E,  Nov. 13–Mar. 17, 1980(G. Monteith, QM S16114), 1 ♀;  Bruxner Pk, via Coffs Harbour, rainforest, leaf litter,  150 m, 30.23333 °S, 153.10000 °E,  May 14, 1969(R.W. Taylor, ANIC, PBI_OON 25871), 2 ³;  Bruxner Pk, via Coffs Harbour, rainforest, leaf mold, rotten wood,  150 m, 30.23333 °S, 153.10000 °E,  June 25, 1967(R.W. Taylor, ANIC, PBI_OON 26297), 1 ³, 1 ♀;  Gibraltar Range National Park, rainforest,  450 m, 29.58333 °S, 152.21670 °E,  Mar. 30, 1980(G. Monteith, QM S16112), 1 ³;  Washpool State Forest, Coombadjah Creek, leaf litter, 29.26666 °S, 152.36666 °E,  Feb. 12, 1982(C. Horseman, AM KS9354, PBI_OON 26293), 2 ♀.  Queensland:  5.5 kmE of One Tree Hill, semievergreen vine thicket,  120 m, 25.33583 °S, 151.90970 °E,  Dec. 14, 1999, to  Mar. 19, 2000(G. and S. Monteith, QMS57432, PBI_OON 22134), 1 ³;   5.5 kmE of One Tree Hill, semievergreen vine thicket,  120 m, 25.33583 °S, 151.90970 °E,  Dec. 14, 1999, to  Mar. 19, 2000(G. and S. Monteith, QM S47072, PBI_OON 22246), 1 ³, 1 ♀;  Kroombit Tops, 45 kmSSW. of Calliope, rainforest, leaf litter,  890 m, 24.41666 °S, 151.01666 °E,  Dec. 10, 1983(G. Monteith, V. Davies, J. Gallon, G. Thomp- son, QM S16096, PBI_OON 25890), 1 ³;  Lamington National Park, Binnaburra Lodge, 28.18333 °S, 153.18333 °E(  July 2, 1986, M.S. Harvey, P.J. Vaughan, WAM T90/912, PBI_ OON 00004311–2), 1 ³, 1 ♀;  Lamington National Park, O’Reillys Guest House, rainforest, leaf litter, 28.23333 °S, 153.13333 °E,  Dec. 14, 1981(G. Monteith, D. Yeates, QM S12990, S16064, S16185, S12985), 3 ³, 4 ♀;  Lamington National Park, O’Reilly’s, rainforest, 28.23333 °S, 153.13333 °E,  Mar. 2–Apr. 26, 1980(G. Monteith, D. Yeates, QM S78262, PBI_ OON 7218), 4 ³;  Lamington National Park, O’Reilly’s, rainforest, 28.23333 °S, 153.13333 °E,  Mar. 2–Apr. 26, 1980(G. Monteith, D. Yeates, QMS78249, PBI_ OON 7230), 1 ³, 1 ♀;  Lamington National Park, O’Reilly’s, rainforest, leaf litter, 28.23333 °S, 153.13333 °E,  Nov. 22–27, 1978( Lawrence, Weir, ANIC, PBI_OON 26295), 1 ³;  Lamington National Park, O’Reilly’s, rainforest, leaf litter, 28.23333 °S, 153.13333 °E,  Nov. 22–27, 1978( Lawrence, Weir, ANIC, PBI_OON 26296), 1 ³, 1 ♀;  Lamington National Park, rainforest, leaf litter,  960 m, 28.20000 °S, 153.16670 °E,  July 9–10, 1977(R. Raven, QM, S16198, S86457, S86440, S86437, S86465– S86466, S86448, S86459), 8 ³, 3 ♀;  Lamington National Park, Binna Burra, rainforest, leaf litter, 28.18333 °S, 153.18333 °E,  Mar. 21–23, 2006(M. Ramírez, QMS72917, PBI_OON 21974), 1 ♀;  Lamington National Park, IBISCA Plot #IQ–700–CKA, rainforest, leaf litter,  720 m, 28.23694 °S, 153.15194 °E,  Jan. 22, 2008(C. Burwell, QM, S86441, S86443, S86455, S16018– S16021), 11 ³, 4 ♀;  Mount Superbus, Site  7, 850 m, 28.23333 °S, 152.48330 °E,  Mar. 12–June 13, 1990(T.B. Churchill, R.J. Raven, QM S15910, PBI_OON 21994), 1 ♀;  Mount Superbus, summit,  1350 m, 28.22183 °S, 152.45280 °E,  Oct. 24, 1998(G. Monteith, QM S54213, PBI_OON 21973), 1 ♀;  Mount Superbus, Trap1, 1350 m, 28.23333 °S, 152.48330 °E,  Mar. 12–June 13, 1990(T.B. Churchill, R.J. Raven, QM S15911, PBI_OON 22004), 1 ♀;  Tamborine National Park, Curtis Falls Track, rainforest, leaf litter, 27.93194 °S, 153.19388 °E,  May 11, 2007(K. Edwardand K. Pitz, QM S96011, PBI_OON 25728), 1 ♀;  Upper Tallebudgera Creek,  550 m, 28.25000 °S, 153.26670 °E,  Jan. 8, 1984(G. Monteith, G. Thompson, QMS69632, PBI_OON 22041), 1 ³;  Upper Tallebudgera valley, below Springbrook, rainforest, leaf litter,  550 m, 28.25000 °S, 153.26666 °E,  Jan. 8–Mar. 17, 1985(G. Monteith, D. Cook, G. Thompson, QM S16473, PBI_ OON 5910), 2 ♀;  Lamington National Park, Binna Burra, along Border Track,  900 m, 28.19861 °S, 153.1875 °E,  Apr 29–30, 2009(H. Wood, CASENT 9035029, PBI_ OON 00005631), 1 ³;  Lamington National Park, Binna Burra, along Border Track,  900 m, 28.19861 °S, 153.1875 °E,  Apr 29–30, 2009(H. Wood, CASENT 9035032, PBI_ OON 00005639, 1 ♀.  DISTRIBUTION: This species is relatively widespread from Coffs Harbour, New South- Walesto Bundaberg, northeastern Queensland (map 14). S16568, OON 21959 1990-03-12 1990-06-13 1990-03-12 QM, PBI Superbus & T. B. Churchill & R. J. Raven 1200 -28.23333 1 152.4667 129 129 OON 5609 4 1 allotype S69613, OON 22217 2002-04-22 QM, PBI Monteith 1000 -28.373 Wiangaree 1 153.1007 Beech Picnic Area 131 131 3 3 New South Wales S96010, OON 25729 2007-05-09 QM, PBI Edward & Pitz Border Ranges National Park -28.37277 Creek 1 153.06915 Brindle Creek 131 131 1 1 New South Wales S16054 1980-03-22 QM Monteith 150 132 132 -30.23333 Coffs Harbour 1 153.1 Bruxner Pk 131 131 1 1 New South Wales S16045, S16151 1980-03-22 1980-11-13 1980-03-22 QM Monteith 150 -30.23333 Bruxner Park 1 153.1 132 132 9 9 New South Wales S16114 1980-03-17 1980-11-13 1980-03-17 QM Monteith 150 -30.23333 Bruxner Park 1 153.1 132 132 1 1 New South Wales OON 25871 1969-05-14 ANIC, PBI Taylor 150 -30.23333 Coffs Harbour 1 153.1 Bruxner Pk 132 132 1 New South Wales OON 26297 1967-06-25 ANIC, PBI Taylor 150 -30.23333 Coffs Harbour 1 153.1 Bruxner Pk 132 132 1 1 New South Wales S16112 1980-03-30 QM Monteith 450 -29.58333 Gibraltar Range National Park 1 152.2167 132 132 1 New South Wales OON 26293, S57432, OON 22134 1982-02-12 2000-03-19 1982-02-12 AM, PBI, QM Horseman & Monteith 120 -25.33583 Coombadjah Creek 1 151.9097 Washpool State Forest 132 132 KS9354 2 2 Queensland S47072, OON 22246 1999-12-14 2000-03-19 1999-12-14 QM, PBI Monteith 120 -25.33583 5.5 km E of One Tree Hill 1 151.9097 132 132 1 1 Queensland S16096, OON 25890 1983-12-10 QM, PBI Monteith & Davies & Gallon 890 133 133 -24.41666 Calliope 1 151.01666 Kroombit Tops 132 132 1 Queensland 1986-07-02 WAM, PBI Harvey & Vaughan -28.18333 Binnaburra Lodge 1 153.18333 Lamington National Park 133 133 T90, OON 00004311-2 1 1 Queensland S12990, S16064, S16185, S12985 1981-12-14 QM Monteith & Yeates -28.23333 O'Reillys Guest House 1 153.13333 Lamington National Park 133 133 4 4 Queensland S78262 1980-03-02 1980-04-26 1980-03-02 QM, PBI Monteith & Yeates -28.23333 O'Reilly's 1 153.13333 Lamington National Park 133 133 OON 7218 1 Queensland S78249 1980-03-02 1980-04-26 1980-03-02 QM, PBI Monteith & Yeates -28.23333 O'Reilly's 1 153.13333 Lamington National Park 133 133 OON 7230 1 1 Queensland OON 26295 1978-11-22 1978-11-27 1978-11-22 ANIC, PBI Lawrence & Weir -28.23333 O'Reilly's 1 153.13333 Lamington National Park 133 133 1 Queensland OON 26296 1978-11-22 1978-11-27 1978-11-22 ANIC, PBI Lawrence & Weir -28.23333 O'Reilly's 1 153.13333 Lamington National Park 133 133 1 1 Queensland S16198, S86457, S86440, S86437, S86465, S86466, S86448, S86459 1977-07-09 1977-07-10 1977-07-09 QM Raven 960 -28.2 Lamington National Park 1 153.1667 133 133 3 3 Queensland S72917, OON 21974 2006-03-21 2006-03-23 2006-03-21 QM, PBI Ramirez -28.18333 Binna Burra 1 153.18333 Lamington National Park 133 133 1 1 Queensland S86441, S86443, S86455, S16018, S16021 2008-01-22 QM Burwell 720 -28.23694 Plot 1 153.15195 Lamington National Park 133 133 4 4 Queensland S15910, OON 21994 1990-03-12 1990-06-13 1990-03-12 QM, PBI Churchill & Raven 7850 -28.23333 Site 1 152.4833 Mount Superbus 133 133 1 1 Queensland S54213, OON 21973 1998-10-24 QM, PBI Monteith 1350 -28.22183 Mount Superbus 1 152.4528 133 133 1 1 Queensland S15911, OON 22004 1990-03-12 1990-06-13 1990-03-12 QM, PBI Churchill & Raven -28.23333 Trap 1 152.4833 Mount Superbus 133 133 1 1 Queensland S96011, OON 25728 2007-05-11 QM, PBI Edward & Pitz -27.93194 Curtis Falls Track 1 153.19388 Tamborine National Park 133 133 1 1 Queensland S69632, OON 22041 1984-01-08 QM, PBI Monteith & Thompson 550 -28.25 Upper Tallebudgera Creek 1 153.2667 133 133 1 Queensland S16473 1985-01-08 1985-03-17 1985-01-08 QM, PBI Monteith & Cook & Thompson 550 -28.25 Springbrook 1 153.26666 Upper Tallebudgera valley 133 133 OON 5910 2 2 Queensland 2009-04-29 2009-04-30 2009-04-29 PBI Wood Lamington National Park 900 -28.19861 Border Track 4 153.1875 Binna Burra 133 133 OON 00005631 1 Queensland 2009-04-29 2009-04-30 2009-04-29 PBI Wood Lamington National Park 900 -28.19861 Border Track 4 153.1875 Binna Burra 133 133 OON 00005639 1 1 Queensland