Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae)
Edward, Karen L.
Harvey, Mark S.
Bulletin of the American Museum of Natural History
2014
2014-06-06
2014
389
1
144
3Q64V
Edward & Harvey, 2014
Edward & Harvey
2014
[159,471,1293,1316]
Arachnida
Oonopidae
Ischnothyreus
Animalia
Araneae
77
77
Arthropoda
species
corniculatum
sp. nov.
ETYMOLOGY: The specific epithet is the Latin corniculatummeaning ‘‘little horn’’ ( Brown, 1956), and refers to the small hornshaped process on the distal tip of the male palp.
DIAGNOSIS: Like I. cornuatus, sp. nov., the male of this species possesses a hornlike modification on the embolic region of the male pedipalp. However, I. corniculatum, sp. nov., has two small lateral processes and only one dorsal hornlike process, instead of two (fig. 46 A, B). The epigynal atrium of the female epigynal region is not paired, and exists as a wide, but short depression that is heavily sclerotized around the posterior edge (figs. 45J, 46F).
MALE (PBI_OON 5611, figs. 45A–C, G–I, 46A–E). Total length 1.53. CEPHALOTHO- RAX: Carapaceyellow-brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. Clypeusmargin unmodified, curved downward in front view, high, ALE separated from edge of carapace by their radius or more; setae dark. Eyes:ALE largest, ALE oval, PME circular, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternumas long as wide, pale orange, darker anteriorly near labium forming unique pattern (fig. 45H, I); setae dark, evenly scattered. Chelicerae, endites, and labium dark redbrown. Chelicerae straight, anterior face unmodified; promargin with one larger denticle; fang shape normal, without prominent basal process; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, much more heavily sclerotized than sternum, with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. ABDOMEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum yellow-brown, covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface finely reticulate, sides finely reticulate. Epigastric scutum small lateral sclerites present. Postepigastric scutum pale orange, covering about 1/2 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter; patella about as long as femur; cymbium dark redbrown; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region with one posteriorly directed hornlike process and two small lateral processes (fig. 46A, B). Fig. 45. Ischnothyreus corniculatum, sp. nov.Holotype male (PBI_OON 00005611): A.habitus, dorsal view; B.habitus, ventral view; C.habitus, lateral view; G.carapace, anterior view; H.sternum, ventral view; I.sternum, oblique view. Allotype female (PBI_OON 00026309): D.habitus, dorsal view; E.habitus, ventral view; F.habitus, lateral view; J.epigynum, ventral view. Fig. 46. Ischnothyreus corniculatum, sp. nov.Holotype male (PBI_OON 00005611): A.left palp, prolateral view; B.left palp, retrolateral view; C.left chelicerae, anterior view; D.left chelicerae, posterior view; E.left palp, dorsal view. Allotype female (PBI_OON 00026309): F.epigynum, ventral view. Scale lines 5 0.1 mm. FEMALE (PBI_OON 26309, figs. 45D–F, J, 46F). Total length 1.50. CEPHALOTHO- RAX: Carapacepars cephalica slightly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less. Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum between 1/4 and 1/2 abdomen width. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Leg spination: femora: I p0-2-0; r0-1-1; II p0-1-0; r0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1- 1-0; v1-1-0; spines on retrolateral side much smaller than prolateral side. GENITALIA: Ventral view: epigynal atrium wider than high, sclerotization heavier around curved posterior edge; convoluted duct thicker than apodemes (figs. 45J, 46F). OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: 2 kmE of South Johnstone Forestry Camp, 17.60000 °S, 146.00000 °E, Dec. 1, 1993, to Feb. 25, 1994(J. Hasenpusch, QM S63216, PBI_OON 22037), 1 ♀; Boulder Creek, via Tully, rainforest, leaf litter, 17.83333 °S, 145.90000 °E, Oct. 27, 1983(G. Monteith, D. Yeates, G. Thompson, QM S16024, PBI_OON 25798), 1 ³; Downey Creek, 25 kmSE Millaa Millaa, 400 m, 17.65000 °S, 145.78330 °E, Dec. 7, 1988( G. Monteith, G. Thompson, QM S31962, PBI_OON 21746), 1 ♀; Herberton Range State Forest, Forestrytrack, rainforest, leaf litter, 17.27111 °S, 145.42277 °E, Apr. 23, 2009( K. Edwardand P. Cullen, QM S95947, PBI_ OON 5535), 1 ³; Kenny Road, 850 m, 17.46667 °S, 145.53330 °E, Nov. 25, 1994, to Jan. 10, 1995(G. Monteith, J. Hasenpusch, QM S27980, PBI_OON 21789), 1 ³; Longlands GapSF, forest, 17.45500 °S, 145.47420 °E, July 23–Nov. 26, 1992(R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S19940, PBI_OON 25927), 1 ³, 1 ♀; MalaanSF, rainforest, 17.58834 °S, 145.60750 °E, Nov. 26, 1992, to Apr. 15, 1993(R. Raven, P. Lawless, QM S24383, PBI_OON 21735), 1 ³; Malaan SF, rainforest, 1000 m, 17.58333 °S, 145.58330 °E, Apr. 20–24, 1978(R. Raven, V. Davies, QM S16126, PBI_OON 25796), 4 ³; Malaan SF, rainforest, 1000 m, 17.58333 °S, 145.58330 °E, Apr. 20–24, 1978(R. Raven, V. Davies, QM S16126, PBI_OON 25915), 3 ³; McNamee Creek, rainforest, leaf litter, 400 m, 17.63333 °S, 145.8500 °E, July 8, 1971( Taylorand Feehan, ANIC, PBI_OON 25848), 2 ³; Mount Fisher, 7 kmSW Millaa Millaa, Kjellberg Road, 1000 m, 17.55000 °S, 145.56670 °E, May 3, 1983( G. Monteith, D. Yeates, QM S16097, PBI_OON 25800), 1 ³; Mount Fisher, 7 kmSW Millaa Millaa, Kjellberg Road, 1000 m, 17.55000 °S, 145.56670 °E, May 3, 1983( G. Monteith, D. Yeates, QM S16091, PBI_OON 26241), 2 ♀; Mount Fisher, Kjellberg Road, 1000 m, 17.53333 °S, 145.55000 °E, Dec. 1, 1993, to Feb. 25, 1994(J. Hasenpusch, QM S34940, PBI_OON 21947), 1 ³; Palmerston National Park, rainforest, 670 m, 17.58834 °S, 145.70000 °E, Nov. 30, 1992, to Apr. 15, 1993(R.J. and S. Raven, P. and E. Lawless, QM S22948, PBI_OON 25919), 1 ³; Palmerston National Park, rainforest, 670 m, 17.58834 °S, 145.70000 °E, July 25–Nov. 30, 1992(R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S21922, PBI_OON 25931), 1 ³; Palmerston National Park, Downey CreekRoad, rainforest, 17.60500 °S, 145.76670 °E, July 25–Nov. 30, 1992(R. Raven, P. Lawless, M. Shaw, QM S24238, PBI_OON 21705), 2 ³; Palmerston National Park, Downey CreekRoad, rainforest, 17.60500 °S, 145.76670 °E, Oct. 30, 1991, to July 24, 1992(R. Raven, P. Lawless, M. Shaw, QM S24577, PBI_OON 21708), 2 ♀; the Millstream, 10 kmNNE Ravenshoe, 1040 m, 17.53333 °S, 145.51670 °E, Dec. 6, 1998, to Feb. 5, 1999(G. Monteith, D. Cook, QM S49225, PBI_OON 22202), 1 ³, 1 ♀; Tully Gorge National Park, rainforest, leaf litter, 750 m, 17.7000 °S, 145.5500 °E, July 2, 1971( Taylorand Feehan, ANIC, PBI_ OON 5848), 1 ♀; Upper Boulder Creek, 8 kmN of Tully, 250 m, Dec. 4–7, 1989(G. Monteith, G. Thompson, H. Janetzki, QMS46824, PBI_OON 22027), 1 ♀; Upper Boulder Creek, Walter Hill Range, 950 m, 17.83333 °S, 145.90000 °E, Nov. 17–18, 1984(V. Davies, G. Monteith, J. Gallon, D. Cook, G. Thompson, QM S12923, PBI_ OON 6434), 1 ♀; Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E, Apr. 27, 2009( K. Edwardand P. Cullen, QM S95948, PBI_ OON 5566), 1 ♀; Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E, Apr. 27, 2009( K. Edwardand P. Cullen, QM S95949, PBI_OON 26306), 1 ♀; Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E, May 17, 2007( K. Edwardand K. Pitz, QM S95950, PBI_OON 26312), 1 ♀.
DISTRIBUTION: This species is only known from the Atherton Tablelands (AU) and the Cairns-Cardwell Lowlands (CC), of the Wet Tropics Bioregion, in northeastern Queensland(map 6).
S95945, S95946
2009-04-23
QM, PBI
K. Edward & P. Cullen
Australia
Male
-17.27111
Forestry
1
145.42278
Herberton Range State Forest
77
77
OON 00005611, OON 00026309
5
2
Queensland
holotype
S63216, OON 22037
1993-12-01
1994-02-25
1993-12-01
QM, PBI
Hasenpusch
-17.6
2 km E of South Johnstone Forestry Camp
1
146.0
80
80
1
1
Queensland
S16024, OON 25798
1983-10-27
QM, PBI
Monteith & D. Yeates & G. Thompson
-17.83333
Tully
1
145.9
Boulder Creek
80
80
1
Queensland
S31962, OON 21746
1988-12-07
QM, PBI
G. Monteith & G. Thompson
400
-17.65
Downey Creek
1
145.7833
80
80
1
1
Queensland
S95947
2009-04-23
QM, PBI
K. Edward & P. Cullen
-17.27111
Forestry
1
145.42278
Herberton Range State Forest
80
80
OON 5535
1
Queensland
S27980, OON 21789
1994-11-25
1995-01-10
1994-11-25
QM, PBI
Monteith & Hasenpusch
850
-17.46667
Kenny Road
1
145.5333
80
80
1
Queensland
S19940, OON 25927
1992-07-23
1992-11-26
1992-07-23
QM, PBI
Raven & P. Lawless & E. Lawless & Shaw
-17.455
Longlands Gap
1
145.4742
80
80
1
1
Queensland
S24383, OON 21735
1992-11-26
1993-04-15
1992-11-26
QM, PBI
Raven & Lawless
-17.58834
Malaan
1
145.6075
80
80
1
Queensland
S16126, OON 25796
1978-04-20
1978-04-24
1978-04-20
QM, PBI
Malaan SF & Raven & V. Davies
1000
-17.58333
1
145.5833
80
80
1
Queensland
S16126, OON 25915
1978-04-20
1978-04-24
1978-04-20
QM, PBI
Malaan SF & Raven & V. Davies
1000
-17.58333
1
145.5833
80
80
1
Queensland
OON 25848
1971-07-08
ANIC, PBI
Taylor & Feehan
400
-17.63333
McNamee Creek
5
145.85
80
80
1
Queensland
S16097, OON 25800
1983-05-03
QM, PBI
G. Monteith & D. Yeates
1000
-17.55
Kjellberg Road
1
145.5667
Mount Fisher
80
80
1
Queensland
S16091, OON 26241
1983-05-03
QM, PBI
G. Monteith & D. Yeates
1000
-17.55
Kjellberg Road
1
145.5667
Mount Fisher
80
80
2
2
Queensland
S34940, OON 21947
1993-12-01
1994-02-25
1993-12-01
QM, PBI
Hasenpusch
1000
-17.53333
Kjellberg Road
1
145.55
Mount Fisher
80
80
1
Queensland
S22948, OON 25919
1992-11-30
1993-04-15
1992-11-30
QM, PBI
Raven & P. and E. Lawless
670
-17.58834
Palmerston National Park
1
145.7
80
80
1
Queensland
S21922, OON 25931
1992-07-25
1992-11-30
1992-07-25
QM, PBI
Raven & P. Lawless & E. Lawless & M. Shaw
670
-17.58834
Palmerston National Park
1
145.7
80
80
1
Queensland
S24238, OON 21705
1992-07-25
1992-11-30
1992-07-25
QM, PBI
Raven & P. Lawless & M. Shaw
-17.605
Downey Creek
1
145.7667
Palmerston National Park
80
80
1
Queensland
S24577, OON 21708
1991-10-30
1992-07-24
1991-10-30
QM, PBI
Raven & P. Lawless & M. Shaw
-17.605
Downey Creek
1
145.7667
Palmerston National Park
80
80
2
2
Queensland
S49225, OON 22202
1998-12-06
1999-02-05
1998-12-06
QM, PBI
Monteith & D. Cook
1040
-17.53333
Millstream
1
145.5167
80
80
1
1
Queensland
1971-07-02
ANIC, PBI
Taylor & Feehan
750
-17.7
Tully Gorge National Park
7
145.55
80
80
OON 5848
1
1
Queensland
S46824, OON 22027
1989-12-04
1989-12-07
1989-12-04
QM, PBI
Monteith & Thompson & Janetzki
250
Upper Boulder Creek
80
80
1
1
Queensland
S12923
1984-11-17
1984-11-18
1984-11-17
QM, PBI
Davies & G. Monteith & J. Gallon & D. Cook & G. Thompson
950
-17.83333
Walter Hill Range
1
145.9
Upper Boulder Creek
80
80
OON 6434
1
1
Queensland
S95948
2009-04-27
QM, PBI
K. Edward & P. Cullen
81
81
-17.61
Tchupala Falls
1
145.77722
Wooroonooran National Park
80
80
OON 5566
1
1
Queensland
S95949, OON 26306
2009-04-27
QM, PBI
K. Edward & P. Cullen
-17.61
Tchupala Falls
1
145.77722
Wooroonooran National Park
81
81
1
1
Queensland
S95950, OON 26312
2007-05-17
QM, PBI
K. Edward & K. Pitz
-17.61
Tchupala Falls
1
145.77722
Wooroonooran National Park
81
81
1
1
Queensland