Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae) Edward, Karen L. Harvey, Mark S. Bulletin of the American Museum of Natural History 2014 2014-06-06 2014 389 1 144 3Q64V Edward & Harvey, 2014 Edward & Harvey 2014 [159,471,1293,1316] Arachnida Oonopidae Ischnothyreus Animalia Araneae 77 77 Arthropoda species corniculatum sp. nov.  ETYMOLOGY: The specific epithet is the Latin  corniculatummeaning ‘‘little horn’’ ( Brown, 1956), and refers to the small hornshaped process on the distal tip of the male palp.  DIAGNOSIS: Like  I. cornuatus, sp. nov., the male of this species possesses a hornlike modification on the embolic region of the male pedipalp. However,  I. corniculatum, sp. nov., has two small lateral processes and only one dorsal hornlike process, instead of two (fig. 46 A, B). The epigynal atrium of the female epigynal region is not paired, and exists as a wide, but short depression that is heavily sclerotized around the posterior edge (figs. 45J, 46F).  MALE (PBI_OON 5611, figs. 45A–C, G–I, 46A–E). Total length 1.53. CEPHALOTHO- RAX: Carapaceyellow-brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. Clypeusmargin unmodified, curved downward in front view, high, ALE separated from edge of carapace by their radius or more; setae dark. Eyes:ALE largest, ALE oval, PME circular, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternumas long as wide, pale orange, darker anteriorly near labium forming unique pattern (fig. 45H, I); setae dark, evenly scattered. Chelicerae, endites, and labium dark redbrown. Chelicerae straight, anterior face unmodified; promargin with one larger denticle; fang shape normal, without prominent basal process; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, much more heavily sclerotized than sternum, with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. ABDOMEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum yellow-brown, covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface finely reticulate, sides finely reticulate. Epigastric scutum small lateral sclerites present. Postepigastric scutum pale orange, covering about 1/2 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-2-0; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter; patella about as long as femur; cymbium dark redbrown; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region with one posteriorly directed hornlike process and two small lateral processes (fig. 46A, B).  Fig. 45.  Ischnothyreus corniculatum, sp. nov.Holotype male (PBI_OON 00005611): A.habitus, dorsal view; B.habitus, ventral view; C.habitus, lateral view; G.carapace, anterior view; H.sternum, ventral view; I.sternum, oblique view. Allotype female (PBI_OON 00026309): D.habitus, dorsal view; E.habitus, ventral view; F.habitus, lateral view; J.epigynum, ventral view.  Fig. 46.  Ischnothyreus corniculatum, sp. nov.Holotype male (PBI_OON 00005611): A.left palp, prolateral view; B.left palp, retrolateral view; C.left chelicerae, anterior view; D.left chelicerae, posterior view; E.left palp, dorsal view. Allotype female (PBI_OON 00026309): F.epigynum, ventral view. Scale lines 5 0.1 mm. FEMALE (PBI_OON 26309, figs. 45D–F, J, 46F). Total length 1.50. CEPHALOTHO- RAX: Carapacepars cephalica slightly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less. Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum between 1/4 and 1/2 abdomen width. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Leg spination: femora: I p0-2-0; r0-1-1; II p0-1-0; r0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1- 1-0; v1-1-0; spines on retrolateral side much smaller than prolateral side. GENITALIA: Ventral view: epigynal atrium wider than high, sclerotization heavier around curved posterior edge; convoluted duct thicker than apodemes (figs. 45J, 46F).  OTHER MATERIAL EXAMINED: AUS- TRALIA:  Queensland:  2 kmE of South Johnstone Forestry Camp, 17.60000 °S, 146.00000 °E,  Dec. 1, 1993, to  Feb. 25, 1994(J. Hasenpusch, QM S63216, PBI_OON 22037), 1 ♀;  Boulder Creek, via Tully, rainforest, leaf litter, 17.83333 °S, 145.90000 °E,  Oct. 27, 1983(G. Monteith, D. Yeates, G. Thompson, QM S16024, PBI_OON 25798), 1 ³;  Downey Creek,  25 kmSE Millaa Millaa,  400 m, 17.65000 °S, 145.78330 °E,  Dec. 7, 1988( G. Monteith, G. Thompson, QM S31962, PBI_OON 21746), 1 ♀;  Herberton Range State Forest, Forestrytrack, rainforest, leaf litter, 17.27111 °S, 145.42277 °E,  Apr. 23, 2009( K. Edwardand P. Cullen, QM S95947, PBI_ OON 5535), 1 ³;  Kenny Road,  850 m, 17.46667 °S, 145.53330 °E,  Nov. 25, 1994, to  Jan. 10, 1995(G. Monteith, J. Hasenpusch, QM S27980, PBI_OON 21789), 1 ³;  Longlands GapSF, forest, 17.45500 °S, 145.47420 °E,  July 23–Nov. 26, 1992(R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S19940, PBI_OON 25927), 1 ³, 1 ♀;  MalaanSF, rainforest, 17.58834 °S, 145.60750 °E,  Nov. 26, 1992, to  Apr. 15, 1993(R. Raven, P. Lawless, QM S24383, PBI_OON 21735), 1 ³;  Malaan SF, rainforest,  1000 m, 17.58333 °S, 145.58330 °E,  Apr. 20–24, 1978(R. Raven, V. Davies, QM S16126, PBI_OON 25796), 4 ³;  Malaan SF, rainforest,  1000 m, 17.58333 °S, 145.58330 °E,  Apr. 20–24, 1978(R. Raven, V. Davies, QM S16126, PBI_OON 25915), 3 ³;  McNamee Creek, rainforest, leaf litter,  400 m, 17.63333 °S, 145.8500 °E,  July 8, 1971( Taylorand Feehan, ANIC, PBI_OON 25848), 2 ³;  Mount Fisher,  7 kmSW Millaa Millaa, Kjellberg Road,  1000 m, 17.55000 °S, 145.56670 °E,  May 3, 1983( G. Monteith, D. Yeates, QM S16097, PBI_OON 25800), 1 ³;  Mount Fisher,  7 kmSW Millaa Millaa, Kjellberg Road,  1000 m, 17.55000 °S, 145.56670 °E,  May 3, 1983( G. Monteith, D. Yeates, QM S16091, PBI_OON 26241), 2 ♀;  Mount Fisher, Kjellberg Road,  1000 m, 17.53333 °S, 145.55000 °E,  Dec. 1, 1993, to  Feb. 25, 1994(J. Hasenpusch, QM S34940, PBI_OON 21947), 1 ³;  Palmerston National Park, rainforest,  670 m, 17.58834 °S, 145.70000 °E,  Nov. 30, 1992, to  Apr. 15, 1993(R.J. and S. Raven, P. and E. Lawless, QM S22948, PBI_OON 25919), 1 ³;  Palmerston National Park, rainforest,  670 m, 17.58834 °S, 145.70000 °E,  July 25–Nov. 30, 1992(R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S21922, PBI_OON 25931), 1 ³;  Palmerston National Park, Downey CreekRoad, rainforest, 17.60500 °S, 145.76670 °E,  July 25–Nov. 30, 1992(R. Raven, P. Lawless, M. Shaw, QM S24238, PBI_OON 21705), 2 ³;  Palmerston National Park, Downey CreekRoad, rainforest, 17.60500 °S, 145.76670 °E,  Oct. 30, 1991, to  July 24, 1992(R. Raven, P. Lawless, M. Shaw, QM S24577, PBI_OON 21708), 2 ♀; the Millstream,  10 kmNNE Ravenshoe,  1040 m, 17.53333 °S, 145.51670 °E,  Dec. 6, 1998, to  Feb. 5, 1999(G. Monteith, D. Cook, QM S49225, PBI_OON 22202), 1 ³, 1 ♀;  Tully Gorge National Park, rainforest, leaf litter,  750 m, 17.7000 °S, 145.5500 °E,  July 2, 1971( Taylorand Feehan, ANIC, PBI_ OON 5848), 1 ♀;  Upper Boulder Creek,  8 kmN of Tully,  250 m,  Dec. 4–7, 1989(G. Monteith, G. Thompson, H. Janetzki, QMS46824, PBI_OON 22027), 1 ♀;  Upper Boulder Creek, Walter Hill Range,  950 m, 17.83333 °S, 145.90000 °E,  Nov. 17–18, 1984(V. Davies, G. Monteith, J. Gallon, D. Cook, G. Thompson, QM S12923, PBI_ OON 6434), 1 ♀;  Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E,  Apr. 27, 2009( K. Edwardand P. Cullen, QM S95948, PBI_ OON 5566), 1 ♀;  Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E,  Apr. 27, 2009( K. Edwardand P. Cullen, QM S95949, PBI_OON 26306), 1 ♀;  Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E,  May 17, 2007( K. Edwardand K. Pitz, QM S95950, PBI_OON 26312), 1 ♀.  DISTRIBUTION: This species is only known from the Atherton Tablelands (AU) and the Cairns-Cardwell Lowlands (CC), of the Wet Tropics Bioregion, in northeastern Queensland(map 6). S95945, S95946 2009-04-23 QM, PBI K. Edward & P. Cullen Australia Male -17.27111 Forestry 1 145.42278 Herberton Range State Forest 77 77 OON 00005611, OON 00026309 5 2 Queensland holotype S63216, OON 22037 1993-12-01 1994-02-25 1993-12-01 QM, PBI Hasenpusch -17.6 2 km E of South Johnstone Forestry Camp 1 146.0 80 80 1 1 Queensland S16024, OON 25798 1983-10-27 QM, PBI Monteith & D. Yeates & G. Thompson -17.83333 Tully 1 145.9 Boulder Creek 80 80 1 Queensland S31962, OON 21746 1988-12-07 QM, PBI G. Monteith & G. Thompson 400 -17.65 Downey Creek 1 145.7833 80 80 1 1 Queensland S95947 2009-04-23 QM, PBI K. Edward & P. Cullen -17.27111 Forestry 1 145.42278 Herberton Range State Forest 80 80 OON 5535 1 Queensland S27980, OON 21789 1994-11-25 1995-01-10 1994-11-25 QM, PBI Monteith & Hasenpusch 850 -17.46667 Kenny Road 1 145.5333 80 80 1 Queensland S19940, OON 25927 1992-07-23 1992-11-26 1992-07-23 QM, PBI Raven & P. Lawless & E. Lawless & Shaw -17.455 Longlands Gap 1 145.4742 80 80 1 1 Queensland S24383, OON 21735 1992-11-26 1993-04-15 1992-11-26 QM, PBI Raven & Lawless -17.58834 Malaan 1 145.6075 80 80 1 Queensland S16126, OON 25796 1978-04-20 1978-04-24 1978-04-20 QM, PBI Malaan SF & Raven & V. Davies 1000 -17.58333 1 145.5833 80 80 1 Queensland S16126, OON 25915 1978-04-20 1978-04-24 1978-04-20 QM, PBI Malaan SF & Raven & V. Davies 1000 -17.58333 1 145.5833 80 80 1 Queensland OON 25848 1971-07-08 ANIC, PBI Taylor & Feehan 400 -17.63333 McNamee Creek 5 145.85 80 80 1 Queensland S16097, OON 25800 1983-05-03 QM, PBI G. Monteith & D. Yeates 1000 -17.55 Kjellberg Road 1 145.5667 Mount Fisher 80 80 1 Queensland S16091, OON 26241 1983-05-03 QM, PBI G. Monteith & D. Yeates 1000 -17.55 Kjellberg Road 1 145.5667 Mount Fisher 80 80 2 2 Queensland S34940, OON 21947 1993-12-01 1994-02-25 1993-12-01 QM, PBI Hasenpusch 1000 -17.53333 Kjellberg Road 1 145.55 Mount Fisher 80 80 1 Queensland S22948, OON 25919 1992-11-30 1993-04-15 1992-11-30 QM, PBI Raven & P. and E. Lawless 670 -17.58834 Palmerston National Park 1 145.7 80 80 1 Queensland S21922, OON 25931 1992-07-25 1992-11-30 1992-07-25 QM, PBI Raven & P. Lawless & E. Lawless & M. Shaw 670 -17.58834 Palmerston National Park 1 145.7 80 80 1 Queensland S24238, OON 21705 1992-07-25 1992-11-30 1992-07-25 QM, PBI Raven & P. Lawless & M. Shaw -17.605 Downey Creek 1 145.7667 Palmerston National Park 80 80 1 Queensland S24577, OON 21708 1991-10-30 1992-07-24 1991-10-30 QM, PBI Raven & P. Lawless & M. Shaw -17.605 Downey Creek 1 145.7667 Palmerston National Park 80 80 2 2 Queensland S49225, OON 22202 1998-12-06 1999-02-05 1998-12-06 QM, PBI Monteith & D. Cook 1040 -17.53333 Millstream 1 145.5167 80 80 1 1 Queensland 1971-07-02 ANIC, PBI Taylor & Feehan 750 -17.7 Tully Gorge National Park 7 145.55 80 80 OON 5848 1 1 Queensland S46824, OON 22027 1989-12-04 1989-12-07 1989-12-04 QM, PBI Monteith & Thompson & Janetzki 250 Upper Boulder Creek 80 80 1 1 Queensland S12923 1984-11-17 1984-11-18 1984-11-17 QM, PBI Davies & G. Monteith & J. Gallon & D. Cook & G. Thompson 950 -17.83333 Walter Hill Range 1 145.9 Upper Boulder Creek 80 80 OON 6434 1 1 Queensland S95948 2009-04-27 QM, PBI K. Edward & P. Cullen 81 81 -17.61 Tchupala Falls 1 145.77722 Wooroonooran National Park 80 80 OON 5566 1 1 Queensland S95949, OON 26306 2009-04-27 QM, PBI K. Edward & P. Cullen -17.61 Tchupala Falls 1 145.77722 Wooroonooran National Park 81 81 1 1 Queensland S95950, OON 26312 2007-05-17 QM, PBI K. Edward & K. Pitz -17.61 Tchupala Falls 1 145.77722 Wooroonooran National Park 81 81 1 1 Queensland