The Geometridae of Ethiopia III: genus Zamarada (Lepidoptera: Geometridae Ennominae, Cassymini) Sciarretta, Andrea Hausmann, Axel Zootaxa 2020 2020-12-10 4894 3 301 328 5CVJ5 (Guenee, 1858) Guenee 1858 [151,589,294,321] Insecta Geometridae Zamarada Animalia Lepidoptera 7 308 Arthropoda species secutaria  ( Figs 14, 15, 41)   Stegania secutaria Guenée, [1858]: 45. Locustypicus: Abyssinie [ Ethiopia] ( Holotype ♀in MNHN).  Literature records.Takusa: 6 milesS.W. of Lake Tana( Fletcher 1974).   Examined material. 1♀:  Ethiopie, Sidamo Wondo Genet,  2400 m,  18.11.1978, P.C. Rougeot( Coll. Herbulotpr. n. [1]5828); 1Ƌ:  Ethiopia, West Shewa, Ambo,  2200 ma.s.l.,  v.2007, leg. V. Kravchenko& G. Müller( ZSMG 20201 / Ƌ / BC ZSM Lep09909); 1Ƌ: S.  ETHIOPIA, Oromia,  13kmS Agere Maryam,  1960 m(lux), 5.5149°N 38.2529°E,  7.XI.2010, leg. De Freina, Hacker, Peks, Schreier( ZSMG 20211 / Ƌ / LEP-SS-00521 failure); 1♀: S.  ETHIOPIA, Sidamo,  16kmSW Kibre Mengist,  1700m, 5.8107°N 38.8880°E,  25.–26.III.2009, lg. R. Beck, M. Dietl( ZSMG 20355 / ♀/ BC ZSM Lep45490); 1Ƌ:  ÄTHIOPIEN, ( EIP)  Äthiopisches Hochland, Bedele 3kmN,  2050m, 08°28.571′N, 36°28.497′O,  10.vii.2015, Beck R. & R. Wanningerleg. ( ZSMG 20316 / Ƌ / BC ZSM Lep101616); 1Ƌ: S.  Ethiopia, Oromia,  6kmESE Jimma,  1850m(lux), 07°39‘34“N 36°53‘20“E,  30.10.2010, leg. de Freina, Hacker, Peks, Schreier( ZSMG 20532/ Ƌ);  1♀: S. Ethiopia, Oromia, Abiyata –Shala-HayakNP,  1646m(lux), 07°31‘27“N 38°39‘44“E,  16.II.2012, leg. de Freina, Hacker, Peks, Schreier( ZSMG20533/ ♀);  1♀: S. Ethiopia SN, 2,65km ESE Wondo Genet,  1950m(lux), 07°04‘36“N 38°38‘30“E,  5.XI.2010, leg. de Freina, Hacker, Peks, Schreier; 1♀:  ETHIOPIA, Oromia,  7kmS Kibre Mengist,  1500m,  21.IV.2009, 5°48′46″N 38°57′41″E, S. Naumann, H. Schnitzler( ZSMG 20804 / ♀); 1Ƌ: S. ETHIOPIA—SN Arba Minch, Nechisar NP, 2.75 kmsw headquarter,  1170 m(lux) 06°00’13,6’’N 37°33’23,4’’E,  22.II.2012, leg. Hacker& Schreier( ZSMG 20813 / Ƌ). Description. Adult Ƌ ( Fig. 14): forewing length: 10–12 mm. Hyaline area of wings light yellow, irrorated with brown and black; basal and medial fasciae in fore- and hindwing poorly defined or absent; costa of forewing light brown, irrorated with brown or black, especially in the medial part; discal spot on forewing grey or light brown, black-ringed; discal spot on hindwings punctiform and black. Non-hyaline terminal area of fore- and hindwings light brown, variably irrorated with brown or black. Termen yellowish-brown, interneurally black-spotted, better marked between R4 and M3 on forewing. Male genitalia ( Fig. 41). Uncus digitate, pointed apically. Ventral plate of gnathus as long as uncus, tapered and scobinate with rounded tip. Dorsal process of valva with a narrow, strongly sclerotized projection, apically spined. Dorsal process stout from the projection toward the base, thin and curved from the projection to the apex, which is pointed. Fulcrum extending beyond aedeagus, curved, with the apex bearing 2–4 stout thorn-shaped spines. Aedeagus with a scobinate ridge just below apex and a single cornutus-like lateral projection. Differential diagnosis. The shape of dorsal process of valva with its projection and fulcrum are the diagnostic characters in male genitalia to discriminate  Z. secutariafrom  Z. chrysothyraand  Z. amicta.  Genetic data. BIN: BOLD:AAH8489 (n=3 from Ethiopia). Genetically heterogeneous, one specimen (BC ZSMLep 45490) diverging by 1.3 %. Nearest species:  Z. metallicataWarren, 1914(4.4%)   Distribution. Algeria, Egypt, Ethiopia, Kenya, Niger, Saudi Arabia, Somalia, Sudan( Guenée 1858; Warren & Rothschild 1905; Rothschild 1915, 1921; Fawcett 1916; Prout, 1916; Wiltshire 1947).   Remarks.So far, only the female was known, unambiguous attribution of males to females was possible by DNA barcodes. 2995113335 [347,508,399,423] MNHN Ethiopia Locus 7 308 1 1 holotype 2995113491 1978-11-18 Rougeot & Coll. Herbulot Ethiopia 2400 Sidamo Wondo Genet 7 308 1 2995113525 G 20201 2007-05 V, ZSM Kravchenko & G. Muller Ethiopia West Shewa 2200 Lep Ambo 7 308 1 2995113395 G 20211 2010-11-07 ZSM De Freina & Hacker & Peks & Schreier Ethiopia 1960 5.5149 13 km S Agere Maryam 7 38.2529 7 308 1 Oromiya 2995113444 G 20355 2009-03-25 2009-03-26 2009-03-25 R, ZSM Beck, M & Dietl Ethiopia 1700 5.8107 Lep 7 38.888 Sidamo 7 308 1 1 2995113396 [758,983,675,703] EIP Ethiopia ATHIOPIEN 7 308 1 2995113526 G 20316 2015-07-10 R, ZSM Beck & Wanninger Ethiopia Athiopisches Hochland 2050 8.476183 Lep 1 -36.47495 Bedele 7 308 1 2995113380 G 20532 2010-10-30 ZSM de Freina & Hacker & Peks & Schreier Ethiopia 1850 7.6594443 6 km ESE Jimma 21 36.88889 7 308 1 Oromiya 2995113458 G20533 2012-02-16 ZSM de Freina & Hacker & Peks & Schreier Ethiopia 1646 7.5241666 Abiyata - Shala-Hayak 21 38.662224 7 308 2 2 Oromiya 2995113510 2010-11-05 de Freina & Hacker & Peks & Schreier Ethiopia 1950 7.076667 Wondo Genet 21 38.64167 7 308 1 1 Oromiya 2995113344 G 20804, G 20813 2009-04-21 2012-02-22 2009-04-21 ZSM Naumann & Schnitzler Ethiopia 1500 6.003778 7 km S Kibre Mengist 1 37.5565 7 308 1 1 Oromia 2995113443 G 20804, G 20813 2009-04-21 2012-02-22 2009-04-21 ZSM Naumann & Schnitzler & Arba Minch & Nechisar NP & Hacker & Schreier Ethiopia 1500 6.003778 7 km S Kibre Mengist 1 37.5565 7 308 1 1 Oromiya