The Geometridae of Ethiopia III: genus Zamarada (Lepidoptera: Geometridae Ennominae, Cassymini)
Sciarretta, Andrea
Hausmann, Axel
Zootaxa
2020
2020-12-10
4894
3
301
328
5CVJ5
(Guenee, 1858)
Guenee
1858
[151,589,294,321]
Insecta
Geometridae
Zamarada
Animalia
Lepidoptera
7
308
Arthropoda
species
secutaria
( Figs 14, 15, 41) Stegania secutaria Guenée, [1858]: 45. Locustypicus: Abyssinie [ Ethiopia] ( Holotype ♀in MNHN). Literature records.Takusa: 6 milesS.W. of Lake Tana( Fletcher 1974).
Examined material. 1♀: Ethiopie, Sidamo Wondo Genet, 2400 m, 18.11.1978, P.C. Rougeot( Coll. Herbulotpr. n. [1]5828); 1Ƌ: Ethiopia, West Shewa, Ambo, 2200 ma.s.l., v.2007, leg. V. Kravchenko& G. Müller( ZSMG 20201 / Ƌ / BC ZSM Lep09909); 1Ƌ: S. ETHIOPIA, Oromia, 13kmS Agere Maryam, 1960 m(lux), 5.5149°N 38.2529°E, 7.XI.2010, leg. De Freina, Hacker, Peks, Schreier( ZSMG 20211 / Ƌ / LEP-SS-00521 failure); 1♀: S. ETHIOPIA, Sidamo, 16kmSW Kibre Mengist, 1700m, 5.8107°N 38.8880°E, 25.–26.III.2009, lg. R. Beck, M. Dietl( ZSMG 20355 / ♀/ BC ZSM Lep45490); 1Ƌ: ÄTHIOPIEN, ( EIP) Äthiopisches Hochland, Bedele 3kmN, 2050m, 08°28.571′N, 36°28.497′O, 10.vii.2015, Beck R. & R. Wanningerleg. ( ZSMG 20316 / Ƌ / BC ZSM Lep101616); 1Ƌ: S. Ethiopia, Oromia, 6kmESE Jimma, 1850m(lux), 07°39‘34“N 36°53‘20“E, 30.10.2010, leg. de Freina, Hacker, Peks, Schreier( ZSMG 20532/ Ƌ); 1♀: S. Ethiopia, Oromia, Abiyata –Shala-HayakNP, 1646m(lux), 07°31‘27“N 38°39‘44“E, 16.II.2012, leg. de Freina, Hacker, Peks, Schreier( ZSMG20533/ ♀); 1♀: S. Ethiopia SN, 2,65km ESE Wondo Genet, 1950m(lux), 07°04‘36“N 38°38‘30“E, 5.XI.2010, leg. de Freina, Hacker, Peks, Schreier; 1♀: ETHIOPIA, Oromia, 7kmS Kibre Mengist, 1500m, 21.IV.2009, 5°48′46″N 38°57′41″E, S. Naumann, H. Schnitzler( ZSMG 20804 / ♀); 1Ƌ: S. ETHIOPIA—SN Arba Minch, Nechisar NP, 2.75 kmsw headquarter, 1170 m(lux) 06°00’13,6’’N 37°33’23,4’’E, 22.II.2012, leg. Hacker& Schreier( ZSMG 20813 / Ƌ). Description. Adult Ƌ ( Fig. 14): forewing length: 10–12 mm. Hyaline area of wings light yellow, irrorated with brown and black; basal and medial fasciae in fore- and hindwing poorly defined or absent; costa of forewing light brown, irrorated with brown or black, especially in the medial part; discal spot on forewing grey or light brown, black-ringed; discal spot on hindwings punctiform and black. Non-hyaline terminal area of fore- and hindwings light brown, variably irrorated with brown or black. Termen yellowish-brown, interneurally black-spotted, better marked between R4 and M3 on forewing. Male genitalia ( Fig. 41). Uncus digitate, pointed apically. Ventral plate of gnathus as long as uncus, tapered and scobinate with rounded tip. Dorsal process of valva with a narrow, strongly sclerotized projection, apically spined. Dorsal process stout from the projection toward the base, thin and curved from the projection to the apex, which is pointed. Fulcrum extending beyond aedeagus, curved, with the apex bearing 2–4 stout thorn-shaped spines. Aedeagus with a scobinate ridge just below apex and a single cornutus-like lateral projection. Differential diagnosis. The shape of dorsal process of valva with its projection and fulcrum are the diagnostic characters in male genitalia to discriminate Z. secutariafrom Z. chrysothyraand Z. amicta. Genetic data. BIN: BOLD:AAH8489 (n=3 from Ethiopia). Genetically heterogeneous, one specimen (BC ZSMLep 45490) diverging by 1.3 %. Nearest species: Z. metallicataWarren, 1914(4.4%)
Distribution. Algeria, Egypt, Ethiopia, Kenya, Niger, Saudi Arabia, Somalia, Sudan( Guenée 1858; Warren & Rothschild 1905; Rothschild 1915, 1921; Fawcett 1916; Prout, 1916; Wiltshire 1947).
Remarks.So far, only the female was known, unambiguous attribution of males to females was possible by DNA barcodes.
2995113335
[347,508,399,423]
MNHN
Ethiopia
Locus
7
308
1
1
holotype
2995113491
1978-11-18
Rougeot & Coll. Herbulot
Ethiopia
2400
Sidamo Wondo Genet
7
308
1
2995113525
G 20201
2007-05
V, ZSM
Kravchenko & G. Muller
Ethiopia
West Shewa
2200
Lep
Ambo
7
308
1
2995113395
G 20211
2010-11-07
ZSM
De Freina & Hacker & Peks & Schreier
Ethiopia
1960
5.5149
13 km S Agere Maryam
7
38.2529
7
308
1
Oromiya
2995113444
G 20355
2009-03-25
2009-03-26
2009-03-25
R, ZSM
Beck, M & Dietl
Ethiopia
1700
5.8107
Lep
7
38.888
Sidamo
7
308
1
1
2995113396
[758,983,675,703]
EIP
Ethiopia
ATHIOPIEN
7
308
1
2995113526
G 20316
2015-07-10
R, ZSM
Beck & Wanninger
Ethiopia
Athiopisches Hochland
2050
8.476183
Lep
1
-36.47495
Bedele
7
308
1
2995113380
G 20532
2010-10-30
ZSM
de Freina & Hacker & Peks & Schreier
Ethiopia
1850
7.6594443
6 km ESE Jimma
21
36.88889
7
308
1
Oromiya
2995113458
G20533
2012-02-16
ZSM
de Freina & Hacker & Peks & Schreier
Ethiopia
1646
7.5241666
Abiyata - Shala-Hayak
21
38.662224
7
308
2
2
Oromiya
2995113510
2010-11-05
de Freina & Hacker & Peks & Schreier
Ethiopia
1950
7.076667
Wondo Genet
21
38.64167
7
308
1
1
Oromiya
2995113344
G 20804, G 20813
2009-04-21
2012-02-22
2009-04-21
ZSM
Naumann & Schnitzler
Ethiopia
1500
6.003778
7 km S Kibre Mengist
1
37.5565
7
308
1
1
Oromia
2995113443
G 20804, G 20813
2009-04-21
2012-02-22
2009-04-21
ZSM
Naumann & Schnitzler & Arba Minch & Nechisar NP & Hacker & Schreier
Ethiopia
1500
6.003778
7 km S Kibre Mengist
1
37.5565
7
308
1
1
Oromiya