The Carabidae (Coleoptera) of Shada Al-A'Ala Nature Reserve, Southwestern Saudi Arabia, with description of a new species of Paussinae Abdel-Dayem, Mahmoud S. Elgharbawy, Ali A. Rasool, Iftekhar Nagel, Peter Aldhafer, Hathal M. ZooKeys 2019 812 93 131 8LJHB Dejean, 1831 Dejean 1831 Insecta Carabidae Bembidion CoL Animalia Bembidion niloticum Coleoptera 26 119 Arthropoda subSpecies niloticum niloticum  Material examined. 825 m, 15.XI.2015, LT, 2♂, 3♀. 851 m: 15.XI.2015, LT, 2♂, 2♀. 892 m: 16.II.2014, LT, 2♂; 15.XI.2015, LT, 1♀. 1,225 m: 03.VI.2014, LT, 1♀; 14.XI.2015, LT, 1♂, 2♀.  General distribution and zoogeography. AE, AF, AM, AZ, BG, CN, CY, EG, GE, GR, IL, IN, IQ, IR, JO, JP, KG, KH, KP, KR, KZ, MM, NP, OM, PH, PK, RU (South European Territory), SA, SY, TM, TR, TW, UZ, VN. ORR_PAL_SAR_SJP species.  Published records. Asir ( Abdel-Dayem et al. 2018), Makkah (Britton). New record for Baha Province.  Remarks. A frequent species that was sporadically collected during autumn, winter, and spring in Acaciathorn woodlands. Paolo Neri and Mahmoud Abdel-Dayem identified this subspecies.