Descriptions of two new genera and six new species of ghost-moths (Lepidoptera Hepialoidea: Hepialidae) from south-eastern and southern Brazil Mielke, Carlos G. C. Grehan, John R. Koike, Ricardo M. Zootaxa 2021 2021-08-13 5020 3 561 580 Mielke & Grehan & Koike, 2021 Mielke & Grehan & Koike 2021 [151,398,1446,1473] Insecta Hepialidae Agripialus Animalia Lepidoptera 3 564 Arthropoda species variabilis sp. nov.  ( Figs 2–4, 21, 29, 32, 37, 52)   Type material.   Holotype ♀( Figs 2a–b): Brazil– SC [ Santa Catarina], Urubici, Santa Bárbara,  1360 m,  25.- 27.XII.2019, 28° 8’30.70’’ S, 49° 38’6.84’’ W, C. Mielke& E. Joerkeleg./ 41.065 Col. C. Mielke/  HOLOTYPUS,  Agripialus variabilisC. Mielke, Grehan & Koike, 2021/ ( DZUP).    Paratypes( 7♀).  BRAZIL.  Paraná:  2♀, Curitiba:  4.II.1975( CGCM 15.404( MGCL));   18.II.1975, V. O. Becker leg. ( CGCM 15.746( CGCM)).   Santa Catarina: 1♀, São Bento do Sul, Rio Natal,  4.IX.1999, A. Rank leg. ( CGCM 7261( CGCM));  2♀, same locality and collector as the holotype,  1.–3.II.2019( CGCM 38.744( CGCM), CGCM 38.764( DZUP));  1♀, São Joaquim,  22–24.I.1983,  1400 m, V. Becker leg. ( CGCM 15.036( CGCM)).   Rio Grande do Sul: 1♀,  28–31.I.2000, São José dos Ausentes, Silveira,  1200 m, A. Moser leg. ( CGCM 3.652( CGCM)).   Diagnosis. Distinguished from its most similar species by differences in the shape of the subrectangular subanal plate, particularly the narrow lobate dorso-posterior corner, and the narrower (by about half) ductus bursae ( Fig. 52).   Description. Male. Unknown.  Female( Figs 2–4, 21, 29, 32, 37, 52).  Head. Antenna with ~42 antennomeres. Basal and distal labial palpomeres equal in length, second segment one and a half to two times longer.  Thorax. Forewing length: 15–22 mm, wingspan: 35–45 mm. Wing ornamentation as shown in Figs 2–4.  Genitalia( Fig. 52). Dorsal plates connected by thickened membranous cuticle or by a narrow sclerotised bar. Dorsal plate subrectangular with shallow convex lateral edge; shallow dorsal, ventral, and medial edges with lobe shaped dorsal corner. Lamella antevaginalis weakly sclerotized, in some cases with a ventrally oriented anterior tip. Subanal plate slightly sclerotised, subrectangular. Ductus bursae length subequal to corpus bursae.  Geographical distribution.  Knownfrom southern Brazilin eastern Paraná, Santa Catarinaand northeastern Rio Grande do Sulat altitude ranging from  800 to 1400 m( Figs 58, 62).   Host plants. Unknown.   Etymology. The proposed specific name alludes to variation in the ornamentation of the forewing dorsally ( Figs 2a, 3–4). It is treated as an adjective in the nominative singular. 2019-12-25 2019-12-27 2019-12-25 C. Mielke & E. Joerke & Col. C. Mielke Brazil 1360 -28.141861 Santa Barbara 1 -49.635235 Urubici 3 564 1 1 Santa Catarina holotype DZUP Brazil 3 564 1 Santa Catarina holotype [199,596,1662,1689] Brazil Parana 3 564 7 7 Parana paratype [609,1159,1662,1689] 1975-02-04 MGCL Brazil Curitiba 3 564 CGCM 15.404 2 2 Parana paratype 1975-02-18 CGCM V. O. Becker Brazil 3 564 CGCM 15.746 1 Parana paratype 1999-09-04 CGCM A. Rank Brazil Rio Natal Sao Bento do Sul 3 564 CGCM 7261 1 1 Santa Catarina paratype 2019-02-01 2019-02-03 2019-02-01 CGCM, DZUP A. Rank Brazil Rio Natal Sao Bento do Sul 3 564 CGCM 38.744, CGCM 38.764 2 2 Santa Catarina holotype [352,1285,1770,1797] 1983-01-22 1983-01-24 1983-01-22 CGCM V. Becker Brazil 1400 Sao Joaquim 3 564 CGCM 15.036 1 1 Santa Catarina paratype 2000-01-28 2000-01-31 2000-01-28 CGCM Sao Jose dos Ausentes & Silveira & A. Moser Brazil 1200 Rio Grande do Sul 3 564 CGCM 3.652 1 1 Rio Grande do Sul paratype Brazil 1100 Known 4 565 1 Parana