Revision and phylogenetic analysis of the spider genus Philisca Simon (Araneae: Anyphaenidae, Amaurobioidinae) 3443 Soto, Eduardo M. Ramírez, Martín J. Zootaxa 2012 2012-08-29 3443 1 1 65 779ZY Soto & Ramírez, 2012 Soto & Ramírez 2012 [151,363,151,177] Arachnida Anyphaenidae Philisca Animalia Araneae 36 37 Arthropoda species robinson sp. nov.   Figs 21–23, 32c, d, 35c   Type material:  Holotypemale and female paratypefrom Chile, Región V( Valparaíso), Archip. Juan Fernández, Is.Robinson Crusoe, área de Plazoleta El Yunque, S33.65276º W78.84403º(GPS, trayecto +-  25m), elev.  299m(barómetro),  16–20.II.2011, bosque de luma, M. Ramírez, E. Soto, J. Pizarro( MHNS, temporary preparations, male: EMS-00156; female: EMS-00155); same locality, 1 maleand 1 female paratypes,  23–28.I.1992, S. Marshall( AMNH; male EMS-00078, female EMS-00068).   Etymology:The specific name is a noun in apposition taken from the typelocality, which is also the name of the main character in the novel Robinson Crusoe, by Daniel Defoe.   Diagnosis:It resembles other species of the  ornatagroup by the modified male chelicerae and the epigyne with smooth, heart shaped median field close to the epigastric furrow ( Figs 35c, 32c, d), and is very similar to  P. pizarroiby the contrasted coloration. It can be distinguished from the latter species by having a pattern of dark spots in the thoracic area ( Figs 21b, 22b) and a darker venter of the opistosoma ( Figs 21c, 22c). The females have the accessory bulbs closer to the spermathecae ( Fig. 32d).   Description: Female( paratypeMHNS EMS-00155): Total length 6.92. Prosoma length 2.67, width 2.07, wider between legs II–III. Tibia/metatarsus length: I, 1.50/1.23; II, 1.33/1.27; III, 1.13/1.40; IV, 1.67/2.03. Chelicerae with three teeth (middle tooth larger) on promargin, three on retromargin (basal tooth larger). Sternum length 1.40, width 1.10. Spines: leg I, femur d 1–1–1, p 2ap; tibia v 2–2–p1; metatarsus v 2bas. II, femur d 1–1–1, p d1ap; tibia v 2–2–2; metatarsus v 2bas, p d1. III, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–p1–r1–2, p and r d1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. IV, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2, p 1–1, r 1–1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. Color in ethanol ( Fig. 21): the entire specimen has a striped appearance. Prosoma and legs reddish brown, almost uniform, both with dark brown spots. Sternum brown, paler in the center. Opisthosoma yellowish brown, venter with dark brown marking with general“w” form ( Fig. 21c). Opisthosoma length 3.99, epigastrium–spiracle 2.17, spiracle–spinnerets 0.37. Epigyne small, sclerotized, translucent ( Fig. 32c, d). MF smooth, heart-shaped, close to the epigastric furrow. Anterior pouch wide, with short lumen. Ducts of AB long, with conspicuous gland ducts.  Male( holotype): Total length 5.59. Prosoma length 2.40, width 1.83. Length of tibia/metatarsus: I, 1.60/1.40; II, 1.47/1.43; III, 1.07/1.30; IV, 1.53/1.83. Chelicerae modified (internal superior margins projecting) ( Fig. 35c), with three teeth (middle tooth larger) on promargin, three on retromargin. Sternum length 1.28, width 1.03. Spines: leg I, femur d 1–1–1, p 2ap; tibia v 2–2–2; metatarsus v 2bas, p d1. II, femur d 1–1–1, p d1ap; tibia v 2–2–2, p 0– v1 or 0; metatarsus v 2–p1–0, p 0–d1–0. III, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2, p and r d1– 1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. IV, femur d 1–1–1, p and r d1ap; patella r d1; tibia v p1–2–2 or 2–2–2, p 1–1, r 1–1–1; metatarsus v 2–2–2, d 0–p1–2, p and r d1–1–1. Opisthosoma length 2.87, spiracle– epigastrium 1.73, spiracle–spinnerets 0.30. Color in ethanol as for female described above ( Fig. 22). Palp ( Fig. 23): Tibia width/length 0.48, Embolus normal, thin and cylindrical ( Fig. 23). MA long, with few small thin branches at base ( Fig. 23f, h). PMA long and wide, pPMA simple. Conductor normal and well developed, canal vestigial, limited to tip, ending in longitudinally ridged peak, on rounded projection directed backward. All adults and immatures have a group of thick setae on the base of ALS.  Variability:Spines in males: I, tibia, v 2–2–p1; metatarsus v 2bas–p1, p 0. III, metatarsus v 2–0–2. Spines in females: I, tibia, v 2–2–0. III, tibia v p1–2–2; metatarsus v 2–p1–2. IV, metatarsus v 2–p1–r1–2, p d1–2–2.   Distribution:Known only from the Juan Fernández Archipelago, Is. Robinson Crusoe, Chile.   Other material examined: CHILE:  Región V( Valparaíso): Archip. Juan Fernández: Is. Robinson Crusoe: same locality as holotype, S33.65276º W78.84403º(GPS, trayecto +-  25m), elev.  299m(barómetro),  16–20.II.2011, bosque de luma, M. Ramírez, E. Soto, J. Pizarro, 1 females, 2 malesand 2 immatures( MHNS),  1 femaleand 3 immatures( MACN-Ar 27960),  1 maleand 2 immatures( MACN-Ar 28460),  1 female( MACN-Ar 28089),  1 female( MACN-Ar 28088),  1 male( MACN-Ar 27961),  pitfall, 3 males( MACN-Ar 27958),  pitfall, 1 femaleand 1 immature( MACN-Ar 27962),  pitfall, 2 femalesand 1 immature( MACN-Ar 27964),  1 maleand 4 immatures( UC),  2 immatures( MACN-Ar 27959),  pitfall, 2 immatures( MACN-Ar 27963);  Sector Salsipuedes, S33.63087° W78.84386°(GPS, área +-  100m), elev.  482m(barómetro),  21.II.2011, bosque de canelo, maqui y luma, M. Ramírez, E. Soto, J. Pizarro, 2 femalesand 31 immatures( UC); same locality as holotype,  23–29.I.1992, S. Marshall, 1 femaleand 1 male( AMNH); same locality,  200–250m Camoteside,  1–28.IV.1962, B. Malkin, 3 females( AMNH); same locality, site 2, pans,  23–29.I.1992, S. Marshall, 1 femaleand 7 males( AMNH); same locality, wet areas nr. Plazoleta, pans,  24– 28.I.1992, S. Marshall, 2 femalesand 6 males( AMNH);  Plazoleta El Yunque, pans in wet areas,  23–29.I.1992, S. Marshall, 14 males( AMNH); same locality, pans nr. Plazoletacampsite,  23–29.I.1992, S. Marshall, 10 males( AMNH); same locality, site 1, open forest, pans,  24–28.I.1992, S. Marshall, 1 femaleand 1 male( AMNH); same locality, upper part; El Yunquetrail, ferns, dung,  23–28.I.1992, S. Marshall, 1 femaleand 1 male( AMNH); same locality, slopes of El Yunque, fern covered, pan traps,  28.I.1992, S. Marshall, 2 femalesand 3 males( AMNH); same locality, site 1, open forest, dung traps,  23–28.I.1992, S. Marshall, 1 femaleand 1 male( AMNH); same locality, dung traps,  23– 29.I.1992, S. Marshall, 2 males( AMNH);  Sendero, luego de Mirador Selkirk, ladera W, S33.63749º W78.85155º(GPS, trayecto +-  50m), elev.  460–539m(barómetro),  18–20.II.2011, bosque de luma con helechos arborescentes, M. Ramírez, E. Soto, J. Pizarro, 1 immature( MACN-Ar 28091),  1 immature( MACN-Ar 28090). 1992-01-23 2011-02-20 1992-01-23 MHNS, AMNH Region V & Archip. Juan Fernandez, Is. & de Plazoleta El Yunque & S. Marshall Chile 25 -33.65276 1 -78.84403 37 38 2 1 1 Valparaiso holotype [199,1432,1539,1564] Region V & Archip. Juan Fernandez & Is. Robinson Crusoe Chile Valparaiso 37 38 1 Valparaiso 2011-02-16 2011-02-20 2011-02-16 MHNS Region V & Archip. Juan Fernandez & Is. Robinson Crusoe Chile 25 -33.65276 Valparaiso 1 -78.84403 37 38 5 1 2 2 Region Chile 37 38 MACN-Ar 27960 4 1 3 Region [503,995,1647,1671] Chile 37 38 MACN-Ar 28460 3 2 1 Region [1007,1329,1647,1671] Chile 37 38 MACN-Ar 28089 1 1 Region Chile 37 38 MACN-Ar 28088 1 1 Region [378,676,1684,1708] Chile 37 38 MACN-Ar 27961 1 1 Region [687,1073,1684,1708] Chile 37 38 MACN-Ar 27958 3 3 Region Chile 37 38 MACN-Ar 27962 2 1 1 Region [376,956,1719,1743] Chile 37 38 MACN-Ar 27964 3 2 1 Region [966,1290,1719,1743] UC Chile 37 38 5 4 1 Region Chile 37 38 MACN-Ar 27959 2 2 Region [377,812,1756,1780] Chile 37 38 MACN-Ar 27963 2 2 Region 2011-02-21 UC Chile 100 -33.63087 Sector Salsipuedes 1 -78.84386 37 38 33 2 31 Region 1992-01-23 1992-01-29 1992-01-23 AMNH Marshall Chile 100 -33.63087 Sector Salsipuedes 1 -78.84386 37 38 2 1 1 Region [267,1224,1864,1888] 1962-04-01 1962-04-28 1962-04-01 AMNH Malkin Chile 225 -33.63087 Camote 1 -78.84386 37 38 3 3 Region 1992-01-23 1992-01-29 1992-01-23 AMNH Marshall Chile 225 -33.63087 Camote 1 -78.84386 37 38 8 1 7 Region 1992-01-24 1992-01-28 1992-01-24 AMNH S. Marshall Chile 225 -33.63087 Plazoleta 1 -78.84386 37 38 8 2 6 Region 1992-01-23 1992-01-29 1992-01-23 AMNH S. Marshall Chile Plazoleta El Yunque 37 38 14 14 Region [477,1433,1972,1996] 1992-01-23 1992-01-29 1992-01-23 AMNH S. Marshall Chile Plazoleta 37 38 10 10 Region [151,1206,2008,2032] 1992-01-24 1992-01-28 1992-01-24 AMNH Marshall Chile Plazoleta 37 38 2 1 1 Region 1992-01-23 1992-01-28 1992-01-23 AMNH El Yunque & S. Marshall Chile 38 39 Plazoleta 37 38 2 1 1 Region 1992-01-28 1992-01-28 1992-01-23 AMNH El Yunque & S. Marshall Chile Plazoleta 38 39 5 2 3 Region 1992-01-23 1992-01-28 1992-01-23 AMNH Marshall Chile 39 40 Plazoleta 38 39 2 1 1 Region 1992-01-23 1992-01-29 1992-01-23 AMNH Marshall Chile Plazoleta 39 40 2 2 Region 2011-02-18 2011-02-20 2011-02-18 M. Ramirez & E. Soto & Pizarro Chile 50 -33.63749 Mirador Selkirk 1 -78.85155 Sendero 39 40 MACN-Ar 28091 1 1 Region [824,1173,252,276] 2011-02-18 2011-02-20 2011-02-18 M. Ramirez & E. Soto & Pizarro Chile 50 true -33.63749 Mirador Selkirk 1 -78.85155 Sendero 39 40 MACN-Ar 28090 1 1 Region