Liparotoma doilu Ramírez, 1993 Philisca doilu : Ramírez 2003: 191 Revision and phylogenetic analysis of the spider genus Philisca Simon (Araneae: Anyphaenidae, Amaurobioidinae) 3443 Soto, Eduardo M. Ramírez, Martín J. Zootaxa 2012 2012-08-29 3443 1 1 65 76XZV (Ramirez) Ramirez 1993 [151,451,671,697] Arachnida Anyphaenidae Philisca Animalia Araneae 13 14 Arthropoda species doilu   Figs 6–8, 34e, 38e    Liparotoma doilu  Ramírez, 1993(female holotype, from Argentina: Neuquén: Lanín Natl. Park: Tromen Lake,  I.1991, V. Fiorani, MACN, reexamined).     Philisca doilu: Ramírez 2003: 191.   Diagnosis:Resembles  P. hyadesiby the low cephalic area ( Ramírez 1993: fig. 30), the transversely wrinkled median field of the epigyne advanced from the epigastric furrow ( Fig. 6b), and the elongate paramedian apophysis ( Fig. 8). It can be distinguished by the narrower sternum ( Fig. 7c; width less than 63% of the sternum length).   Description: Female: described in Ramírez (1993).   FIGURE 6.Female of  Philisca doiluRamírez(MACN-Ar 26762). ( afemale dorsal habitus; bepigyne, ventral view; ccleared epigyne, dorsal view). (AB: accessory bulbs, CD: copulatory ducts, MF: median field, S: spermathecae).   FIGURE 7.  Philisca doiluRamírez(MACN-Ar 15097). a–dmale ( adorsal habitus; bprosoma, dorsal view; cventral habitus; dprosoma, ventral view).   FIGURE 8.  Philisca doiluRamírez(MACN-Ar 15097). Male, a–cleft palp ( aprolateral view, bventral view, cretrolateral view); d–hcopulatory bulb ( dprolateral view, eventral view, fretrolateral view, gapical view, hapical–retrolateral detail view). (C: conductor, E: embolus, MA: median apophysis, PMA: paramedian apophysis, rPMA: retrolateral cusp of the paramedian apophysis).  Male(MACN-Ar 15097): Total length 11.97. Prosoma length 5.45, width 4.26. Length of tibia/metatarsus: I, 6.12/4.79; II, 5.05/4.26; III, 2.39/2.53; IV, 3.72/3.46. Chelicerae very strong ( Fig. 34e), with two teeth on retromargin (basal tooth larger) and three on promargin (apical one very large). Sternum length 2.93, width 1.87. Spines: leg I, femur d 1–1–1, p 2ap. II, femur d 1–1–1, p d2ap. III, femur d 1–1–1, p and r d1ap or 0; patella r d1; tibia v 1–2ap or 0–2ap, p 0–1, r d1–1; metatarsus v 2–0–2, p and r d1–2ap. IV, femur d 3–2–1 or 2–2–1, r d1ap or 0; patella r d1; tibia v 2ap or p1ap, r d1–1; metatarsus v 2–p1–p1–2 or 2–0–0–2, d 2ap, p 1ap, r d1–1. Opisthosoma length 6.38, spiracle–epigastrium 2.80, spiracle–spinnerets 0.53. Color in ethanol ( Fig. 7): Prosoma and chelicerae reddish brown with white pubescence. Sternum brown. Legs brown with black spots, leg I darker than the others. Opistosoma light brown, with dorsal and ventral brown pattern. Femur IV in dorsal view with 2–4 spines at base. Palp ( Fig. 8): Tibia width/length 0.28. Embolus short, anterior margin longitudinally flattened, slightly projecting in a ridge; basal process flattened, rounded ( Fig. 8). MA long, with one big branch at base ( Fig. 8h). PMA long and sinuous. Sperm duct without apical or retrolateral loop. pPMA simple, wide. Conductor complex, with canal, dentate prolateral ridge absent, prolateral portion with triangular, flattened projection; retrolateral portion thin, projecting in acute peak.  Variability:Spines: Females: III, tibia v 0–p1–2; IV, femur d 3–1–1, p 0, r d1ap; tibia r d1–d1–0–1. Males: IV, femur d 4–1–1 or 3–1–1, p d1ap.   Distribution:Southern forests in Chile, from Arauco(Region VIII) to Valdivia provinces (Region XIV), and in Argentina, in Neuquén province( Fig. 38e).  New records:   ARGENTINA:  Neuquén: Cerro La Mona, S40,5678º W71,69727º,  1500m,  I.2006, V. Werenkraut, Pitfall, 1 male( MACN-Ar 15097),  2 males( MACN-Ar 15098).   CHILE:  RegiónIX ( Araucanía): Malleco Prov.: Nahuelbuta Natl. Park, S37º47’ W73º00’,  1200m,  12.II.2005, J. BarrigaT., fogging  Nothofagus dombeyi, 1 female( MACN-Ar 26762),  1 immature( MACN-Ar 26761);  Malalcahuello Way Lonquimay, fogging,  1236m, 15ºC S38º26.377’ W71º30.540’,  N. dombeyi,  35m, 9:47AM,  19.I.2007, Ariaset al. Berkeley, 1 immature( CASENT 9039004);  Malalcahuello Way Lonquimay, fogging,  1221m, 29°C, S38°26.388’ W71°30.613’,  N. dombeyi,  50m, 6:55PM,  18.I.2007, Ariaset al. Berkeley, 1 immature( CASENT 9039011);  Malalcahuello Way Lonquimay, fogging,  1236m, 15ºC, S38º26.377’ W71º30.540’,  N. dombeyi  35m, 9:47AM,  19.I.2007, Ariaset al. Berkeley, 3 immatures( CASENT 9039005).   RegiónXIV (de los Ríos): Valdivia Prov.: Valdivia, Chaihuin, Reserva Costera Valdiviana, Las Garzas, S39°59.640’ W73°35.197’,  360m, 20.9° C Fogging150cc/l  Nothofagus nitida, 11:00AM,  27.II.2008, Ariaset al. UCB, 1 immature( CASENT 9039015);  Valdivia, Chaihuin, Reserva Costera Valdiviana, Las Garzas, S39°49.687’ W73°15.227, 360m, 17.3°C, fogging 150cc/l  N. nitida,11:00AM,  27.II.2008, Ariaset al., 1 femaleand 9 immatures( CASENT 9039063),  1 female( CASENT 9039067).   Material re-examined:   ARGENTINA:  Neuquén:Lanín Natl. Park, Lago Huechulaufquen, Coloradas,  1.I.1985, M. Ramírez, 1 female( MACN-Ar 14800). [342,495,778,800] 1991-01 V. Fiorani Argentina Tromen Lake 13 14 1 Neuquen holotype 2006-01 V. Werenkraut Argentina 1500 -40.5678 Cerro La Mona 5 -71.69727 16 17 MACN-Ar 15097 1 1 Neuquen [701,1027,836,860] 2006-01 V. Werenkraut Argentina 1500 true -40.5678 Cerro La Mona 5 -71.69727 16 17 MACN-Ar 15098 2 2 Neuquen 2005-02-12 J. Barriga Chile Region 1200 -37.783333 Nahuelbuta Natl. Park 1179 -73.0 Malleco Prov. 16 17 MACN-Ar 26762 1 1 Araucania [595,958,908,932] 2005-02-12 J. Barriga Chile Region 1200 true -37.783333 Nahuelbuta Natl. Park 1179 -73.0 Malleco Prov. 16 17 MACN-Ar 26761 1 1 Araucania 2007-01-19 CASENT Arias & Berkeley Chile 1236 -38.439617 Malalcahuello Way Lonquimay 1 -71.509 16 17 CASENT 9039004 1 1 2007-01-18 CASENT Arias & Berkeley Chile 1221 -38.4398 Malalcahuello Way Lonquimay 1 -71.510216 16 17 CASENT 9039011 1 1 2007-01-19 CASENT Arias & Berkeley Chile 1236 -38.439617 Malalcahuello Way Lonquimay 1 -71.509 16 17 CASENT 9039005 3 3 2008-02-27 CASENT C Fogging & Arias Chile Region 360 -39.994 Valdivia 1 -73.58662 Valdivia Prov. 16 17 CASENT 9039015 1 1 Los Rios 2008-02-27 CASENT Arias Chile Valdivia -39.828117 Reserva Costera Valdiviana 1 -73.253784 Chaihuin 16 17 CASENT 9039063 10 1 9 [962,1302,1232,1256] 2008-02-27 CASENT Arias Chile Valdivia true -39.828117 Reserva Costera Valdiviana 1 -73.253784 Chaihuin 16 17 CASENT 9039067 1 1 1985-01-01 M. Ramirez Argentina Coloradas Lago Huechulaufquen 16 17 MACN-Ar 14800 1 1 Neuquen