Ampullaceana relicta pinteri ( Schuett , 1974)

Vinarski, Maxim V., Aksenova, Olga V. & Bolotov, Ivan N., 2020, Taxonomic assessment of genetically-delineated species of radicine snails (Mollusca, Gastropoda, Lymnaeidae), Zoosystematics and Evolution 96 (2), pp. 577-608 : 577

16b. Ampullaceana relicta pinteri ( Schuett, 1974) Fig. 9B.

Radix pinteri Schütt 1974: 471, pl. 1.

Radix pinteri - Albrecht et al. 2008: 160, fig. 2; Welter-Schultes 2012: 54, textfig.; Glöer 2019: 244, fig. 304.

Ampullaceana relicta pinteri - Aksenova et al. 2018a: 4.

TL.

North Macedonia, Lake Prespa near village of Perovo.

Types.

Holotype - SMF; paratypes were placed in NHMW and in a series of private collections in Germany and Hungary ( Schütt 1974); currently, specimens from these private collections are kept in the Hungarian Natural History Museum (Budapest) and in the Natural History Museum of Bern (Switzerland). We examined the paratypes kept in NHMW and in the Budapest museum (see Fig. 9B ).

This subspecies has been characterised both morphologically and genetically in a series of papers ( Schütt 1974; Albrecht et al. 2008; Welter-Schultes 2012). Most authors regard this taxon as a separate species. For example, Albrecht et al. (2008) considered A. r. relicta and A. r. pinteri as two sister species from two ‘sister’ lakes, Ohrid and Prespa (but see Aksenova et al. 2018a). According to the anatomical data presented in Albrecht et al. (2008), the morphology of the copulatory apparatus in A. r. pinteri is virtually identical with that of the nominative subspecies.

A. r. pinteri is endemic to Prespa Lake (Albania, Greece and Macedonia).

Figures

Figure 9. Shells of species of the genera Ampullaceana, Kamtschaticana, Myxas, Tibetoradix, Racesina, Orientogalba and Bullastra. A. Ampullaceana relicta relicta (05.1975, Macedonia, a spring near St. Naum monastery, Ohrid Lake region; NHMW). B. A. r. pinteri, a paratype (05.06.1973, Macedonia, Prespa Lake near Perovo; MHMW). C, D. Kamtschaticana kamtschatica (Russia, Kamchatka Peninsula, Valley of Geysers; RMBH). E. Myxas glutinosa (14.09.1963, Kazakhstan, floodplain of the Tobol River near Kustanay City; ZISP). F, G. Tibetoradix hookeri, two syntypes (NHMUK). H. T. kozlovi, the holotype (ZISP). I. Racesina luteola (India, Bengal, Paksay, Pabna; NHMUK). J. R. oxiana (04.07.2016, Tajikistan, a roadside ditch near Kurban-Shakhid village; LMBI). K. R. siamensis (" Thailand "; ZMB). L. Orientogalba viridis, a syntype (Guam Island; NMNH). M. O. ollula (1883, Japan, Yokohama; NHMUK). N. Limnaea bowelli, a type (after Preston 1909). O. Orientogalba cf. bowelli (30.06.1924, China, Tibet, Lingka, 14,500 feet a. s. l.; NHMUK). P. Bullastra cumingiana (Philippine, Cebu Island; NHMUK). Scale bars: 2 mm (C, D, F-H, L, M, O), 5 mm (A, B, E, J-K, N, P).