Crusbatus spinosus Opitz

Opitz, Weston, 2014, Taxonomy of the New World Checkered Beetle GeneraAkonesisOpitz, New Genus, AntennosusOpitz, New Genus, CrusbatusOpitz, New Genus, andDiutiusOpitz, New Genus (Coleoptera: Cleridae: Peloniinae), The Coleopterists Bulletin 68 (3), pp. 411-428 : 412-413

KEY TO AKONESIS , ANTENNOSUS , CRUSBATUS , DIUTUS, AND THEIR SPECIES

1. Protibial margin with spines........................2

1′. Protibial margin without spines................... 3

2. Pronotum oblong, disc without broad indentations ( Bolivia) ( Fig. 19 ) .............................. ...... Crusbatus spinosus Opitz , new species

2′. Pronotum transverse, disc with 2 broad indentations ( Panama to Argentina) ( Fig. 33 ).......... .......... Antennosus impressocollis (Gorham)

3. Tarsal spur formula 1-2-2 (genus Akonesis ).................................................................4

Character 0- Epipleural fold: in lateral position (0); not in lateral position (1)

Character 1- Capitulum: not longer than length of funicle (0); longer than length of funicle (1)

Character 2- Anterior margin of protibiae: not spinous (0); spinous (1)

Character 3- Epipleuron: not in ventral position (0); in ventral position (1)

Character 4- Pronotal disc: not flat (0); flat (1)

Character 5- Phallobasic struts: not abbreviated (0); abbreviated (1)

Character 6- Phallobasic rod: present (0); absent (1)

Character 7- Funicular antennomeres: not expanded (0); expanded (1)

Character 8- Pronotum: not infuscated (0); infuscated (1)

Character 9- Elytral vittae: absent (0); present (1)

Character 10- Elytral brown maculae: absent (0); present (1)

Character 11- Humeral margin: not infuscated (0); infuscated (1)

Character 12- Pronotal shape: quadrate (0); elongate (1)

Character 13- Pygidial margin: not concave (0); concave (1)

Character 14- Pronotal indentation: absent (0); present (1)

Character 15- Tarsal spur formula: 1-2-2 (0); 0-2-2 (1)

Character 16- Spicular fork apodeme: not bifid (0); bifid (1)

Character 17- Pronotal tubercle: present (0); absent (1)

3′. Tarsal spur formula 0-2-2 (genus Diutius ).....5

4. Elytral disc with brown vitta; humeral margin not infuscated ( Bolivia to Brazil) ( Fig. 63 ) .. ........................... Akonesis vittatis (Spinola)

4′. Elytral disc without brown vitta, disc with small black markings ( Brazil) ( Fig. 64 ) ........ ................... Akonesis pictipennis (Gorham)

5. Elytral disc with brown maculae ( Guyana, French Guiana, Ecuador, Brazil) ( Fig. 61 ).... ................... Diutius gallerucoides (Spinola)

5′. Elytral disc unicolorous, light brown, without brown maculae ( Bolivia) ( Fig. 62 )................ ............ Diutius pallidus Opitz , new species

Figures

Figs. 16–29. Antennosus impressocollis. 16) Head, frontal view; 17) Head, ventral view; 18) Head, dorsal view; 19) Prothorax, ventral view; 20) Antenna, male; 21) Antenna, female; 22) Metathoracic wing; 23) Mandible; 24) Maxilla; 25) Metendosternite; 26) Maxilla; 27) Spicular fork; 28) Tegmen; 29) Phallus.

Figs. 30–42. Morphological structures. 30) Akonesis vittatis, pronotum; 31) Akonesis pictipennis, pronotum; 32) Diutus pallidus, pronotum; 33) Crusbatus spinosus, pronotum; 34) D. pallidus, antenna; 35) Diutus gallerucoides, antenna; 36) C. spinosus, antenna; 37) D. pallidus, tegmen; 38) D. pallidus, phallus; 39) C. spinosus, spicular fork; 40) C. spinosus, male, pygidium; 41) C. spinosus, tegmen; 42) C. spinosus, phallus.

Figs. 61–64. Habitus. 61) Diutius gallerucoides; 62) Diutius pallidus; 63) Akonesis vittatis; 64) Akonesis pictipennis.