Anomalophylla stoetzneri, Ahrens, 2005

Ahrens, Dirk, 2005, Taxonomic revision of the genus Anomalophylla Reitter, 1887 (Coleoptera: Scarabaeidae: Melolonthinae: Sericini), Zootaxa 1076 (1), pp. 1-62 : 29-31

Anomalophylla stoetzneri sp. n.

( Fig. 2K–M , 5 )

Type material. Holotype: ♂ ”Szetschwan Tatsienlu Exp. Stoetzner” ( CF).

Holotype description. Length: 4.9 mm, length of elytra: 3.0 mm, width: 2.6 mm. Body oblong, black; elytra reddish brown with broadly black borders; dorsal surface dull with long, dense, erect setae; pilosity on head and anterior pronotum black, dark yellow on posterior pronotum and elytra. Head: Labroclypeus transverse, widest medially with lateral margins strongly convex and convergent anteriorly and toward base, anterior angles strongly rounded; lateral border and ocular canthus producing distinct blunt angle; anterior and lateral margin strongly reflexed, anterior margin distinctly sinuate medially; surface flat, moderately shiny with double punctation; coarse, dense punctures each bearing a long, erect seta; coarse punctures mixed with fine glabrous punctures. Frontoclypeal suture distinct, weakly curved, slightly elevated; smooth area anterior to eye as wide as long. Ocular canthus moderately long, slender; with fine, dense punctures; densely setose. Frons with double punctation; coarse, dense punctures each bearing a long, erect seta, mixed with fine glabrous punctures; basal punctation less dense. Eyes small, ratio of diameter / interocular width: 0.51. Antenna with ten antennomeres, dark yellow; club brown with five equal in length antennomeres; club 2.5 times as long as the remaining antennomeres combined, reflexed. Prementum flat. Pronotum: widest at base; lateral margins in posterior half almost straight and subparallel, in anterior third strongly convex, convergent; anterior angles not produced and moderately rounded; posterior angles blunt, weakly rounded; anterior margin medially weakly convex with distinct, fine marginal line; basal margin with fine marginal line. Pronotal surface with dense, double punctation; punctures fine, glabrous, mixed with large punctures bearing a long erect seta; pilosity moderately dense. Anterior and lateral borders setose; basal margin of hypomeron not produced ventrally, not transversely sulcate anterior to base. Scutellum : moderately long; apex sharp; with fine, irregularly dense punctures; posterior third almost smooth, minute setae present in the punctures. Elytra: oblong, widest medially; striae indistinctly impressed with fine, dense punctures; intervals flat with fine, moderately dense punctures; punctures with long, erect, evenly dense setae; even intervals with setae near base; sutural interval with robust, long setae. Epipleural edge fine, ending at the strongly convex external apical angle of elytra; epipleura densely setose; elytra strongly curved in anterior third; apical border chitinous, without short microtrichomes. Venter: Ventral surface dull with fine, dense punctures. Metasternum with dense, long setae, setae partially appressed, partially erect. Metacoxa glabrous medially; laterally with moderately dense, fine, long appressed setae. Abdominal sternites with indistinct transverse row of coarse punctures bearing thick setae between fine, dense punctation; all sternites with fine, long setae; tegument of abdominal sternites (60x magnification) with fine polygonal mesh pattern formed by microtrichomes; penultimate sternite at midline with longitudinally impressed line. Mesosternum between mesocoxae narrow, narrower than mesofemur, with irregularly scattered, strong setae. Ratio of length of metepisternum / metacoxa: 1 / 1.6. Pygidium moderately convex; dull; with fine, dense punctures bearing fine, long setae; pilosity denser along apical margin; without smooth midline. Legs: slender with shiny surface. Femora with two longitudinal rows of setae, finely densely punctate and setose; anterior edge of metafemur acute, lacking adjacent serrated line; posterior margin weakly convex with a few fine setae medially, vental posterior margin weakly widened in apical half; posterior margin smooth ventrally and dorsally. Metatibia moderately slender and long, widest at apex; ratio width / length: 1 / 3.5; dorsal margin sharply carinate and smooth with two groups of spines, basal group at one third, apical one at two thirds of metatibial length, basally with a few single spines in punctures; lateral face longitudinally convex with moderately dense, fine, glabrous punctures; ventral edge serrated, with four strong, long, equally spaced spines; medial face finely punctate, apex sharply truncate interiorly near tarsal articulation. Tarsomeres dorsally glabrous, finely densely punctate; ventrally with sparse, short setae. Metatarsomeres dorsally without longitudinal impressions, ventrally with strongly serrated ridge adjacent to strong longitudinal carina, laterally without strong longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined, slightly longer than the upper tibial spur. Protibia short, bidentate. Protarsal claws asymmetrical, basal tooth of inner protarsal claw widened, truncate apically. Aedeagus: Fig. 2K–M .

Diagnosis. Anomalophylla stoetzneri is similar to A. subfastuosa in external shape and genital morphology of male. It differs from A. subfastuosa by the long, slender, but slightly curved right paramere; by the narrow, long basal lobe and similarly long, slender left paramere, which is medially slightly curved interiorly; and the slender, sharply pointed lateral apophysis of the phallobase.

Etymology. The new species is named after Walter Stoetzner, who led the expedition in course of which the holotype has been collected.

Figures

FIGURE 2. (A–C) Anomalophylla bicolor sp. n. (Lectotype: Tatsienlu ­ Kiulung); (D–F) A. liciata sp. n. (holotype); (G–J) A. subfastuosa sp. n. (holotype); (K–M) A. stoetzneri sp. n. (holotype); (N– P) A. hispidulosa sp. n. (holotype); (Q–S) A. majori sp. n. (holotype). (A, D, G, K, N, Q) aedeagus, left side lateral view; (B, E, H, L, O, R) parameres, dorsal view; (C, F, J, M, P, S) aedeagus, right side lateral view. Scale bar = 0.5 mm.

FIGURE 5. Present known distribution of Anomalophylla stoetzneri, A. tristicula, and A. tsangpoana as well as their intermediate form.