Anopinella araguana Brown and Adamski

Brown, John W. & Adamski, David, 2003, Systematic revision of Anopinella Powell (Lepidoptera: Tortricidae: Euliini) and phylogenetic analysis of the Apolychrosis group of genera, Zootaxa 200, pp. 1-94 : 17-19

Anopinella araguana Brown and Adamski , new species

Figs. 5 , 39 , 61

Anopinella ophiodes ; Powell 1986 (not Walsingham): 390 [illustration of male genitalia].

Diagnosis. Anopinella araguana is most similar to A. razowksii both in facies and genitalia but has considerably larger distal lobes of the gnathos and shorter socii. Putative autapomorphies for A. araguana include the elongate spines along the posterior margin of the distal lobes of the gnathos, the elongate, rod­like sclerite of the vesica, and the quadrate sclerotized mesal patch of the lamella postvaginalis in the female genitalia.

Description. Head: Frontoclyeus and vertex pale brown intermixed with dark brown; labial palpus with outer surface of segments I­II brown intermixed with pale brown, segment III pale brown, inner surface pale brown. Antenna with scape brown intermixed with pale brown; basal 6­10 flagellomeres brown; distal flagellomeres gray.

Thorax: Tegula and mesonotum brown intermixed with pale brown and reddish brown. Forewing (Fig. 61) length 7.0­9.0 mm (mean = 9.5; n = 10); incomplete basal fascia brown intermixed with reddish brown and dark brown; costal blotch dark brown intermixed with reddish brown and brown; basal fascia and costal blotch separated by an oblique band of pale brown intermixed with pale reddish brown and few dark brown scales demarcating basal fascia and costal blotch to near cubitus; band recurved from distal costa through subapical area to tornus, encircling crescent­shaped ocellus, except for posterior part; costal blotch with small, subcircular, white spot near posterior end; area between CuP and posterior margin pale reddish brown intermixed with brown and dark brown; apical area dark brown; submarginal area with gray spots near outer margin of ocellus. Fringe gray intermixed with reddish brown. Hindwing brown.

Abdomen: Male genitalia ( Fig. 5 ; drawn from DA slide 4160; n = 1) with uncus curved throughout. Socius digitate, ca. 0.85 length of basal portion of gnathos arms. Gnathos expanded distally, forming two large, free, laterally flattened, crescent­shaped lobes, each with elongate spines on outer margin. Valva densely setose from inner margin of cucullus to basal ridge; sacculus broadly rounded distally, confluent with postsacculus; postsacculus broadly concave, slightly recurved towards apicoventral angle; apicoventral angle protuberant; cucullus densely setose, with a broadly rounded outer margin from an acuminate apex. Phallus simple; vesica densely microspinulate, with an elongate, slightly curved cornutus. Female genitalia ( Fig. 39 ; drawn from DA slide 4161; n = 6) with weakly sclerotized, quadrate mesal patch at lamella postvaginalis; eighth tergum not modified; ostium broadly U­shaped, ventral rim narrowly sclerotized, ca. 0.5 width of seventh sternum at widest point. Antrum parallel­sided, cup­shaped; ductus bursae long, wide; accessory bursae originating ca. 0.8 distance from ostium to corpus bursae. Corpus bursae large, subspherical, with microspicules on inner ca. 0.17 of side bearing ductus seminalis.

Holotype, ɗ, Venezuela, Aragua, Rancho Grande, 1100 m, 24­31 Oct 1966, S. S. & W. D. Duckworth. Deposited in USNM.

Paratypes (2ɗ, 7Ψ). VENEZUELA: Aragua: 5 km West of Tovar, 6300', 24 Jan 1978 (1ɗ), J. B. Heppner (USNM). Rancho Grande, 1100 m, cloud forest, 17­20 Jan 1978 (1ɗ), J. B. Heppner (USNM), 16­23 Oct 1966 (5Ψ), 11­15 Jan 1966 (1Ψ), S. S. Duckworth & W. D. Duckworth (USNM), 1­7 Aug 1967 (1Ψ), R. Poole (USNM).

Additional Specimen Examined: VENEZUELA: Merida: Mucuy Fish Hatchery, 7 km E Tabay, 6600' [2300 m], 10­13 Feb (1ɗ), J. B. Heppner (USNM).

Etymology. The specific epithet is derived from the province, Aragua, Venezuela. Remarks. The male genitalia of the specimen from Merida are nearly identical to those of other males, but they lack the characteristic rod­like sclerite of the vesica; hence this specimen is excluded from the type series. The difference in elevation of the collecting locations (1100 m vs. 2300 m) also suggests that the two may not be conspecific. The genitalia illustrated by Powell (1986) as A. ophiodes is a USNM specimen from Rancho Grande, Venezuela.

Figures

FIGURES 5 ­ 6. Male genitalia of Anopinella, valvae spread, aedeagus removed. 5, A. araguana; 6, A. nr. razowksii.

FIGURES 38 ­ 39. Female genitalia of Anopinella, posterior end up. 38, A. powelli, with insert of tergum eight; 39, A. araguana.