Anthobium nigropiceum, Motschulsky, 1860

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1-650 : 1-650

publication ID

https://doi.org/ 10.1206/0003-0090.265.1.1

persistent identifier

https://treatment.plazi.org/id/BF684949-422A-B501-00BC-2C52FDB47CB2

treatment provided by

Felipe

scientific name

Anthobium nigropiceum
status

 

nigropiceum Motschulsky, 1860 View in CoL , see: boreale Paykull, 1792 .

obtectum Erichson, 1840: 865 ( Olophrum ; Type locality: America septentrionalis ; [Note: See lectotype designation by Campbell, 1983]).

— Fauvel, 1869: 493 ( Olophrum ; synonym of emarginatum ).

— Fauvel, 1878d: 242 [= 1878e: 78] ( Olophrum ; characters; Michigan; Massachusetts; Missouri).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Blatchley, 1910: 479 ( Olophrum ; characters; Indiana).

— Scheerpeltz, 1929: 45 ( Olophrum ; characters; New Jersey, New York, Pennsylvania, Maine, New Hampshire, Massachusetts).

Campbell, 1983: 608 ( Olophrum ; lectotype designation; characters; Ontario; Quebec; Connecticut; Illinois; Indiana; Kansas; Kentucky; Maryland; Massachusetts; Michigan; New Jersey; New York; North Carolina; Ohio; Pennsylvania; Tennessee; Virginia; West Virginia; Wisconsin) .

— Levesque and Levesque, 1996: 290 ( Olophrum ; seasonal abundance; Quebec).

— Downie and Arnett, 1996: 432 ( Olophrum ; characters; USA).

- DISTRIBUTION: Canada, USA.

rotundicolle Say, 1832: 50 [= 1834: 464] ( Omalium ; [preoccupied]; Type locality: Pennsylvania).

— Erichson, 1840: 868 ( Olophrum ; synonym of emarginatum ).

— Melsheimer, 1844: 43 ( Olophrum ; synonym of emarginatum Erichson ).

— Fauvel, 1878d: 242 [= 1878e: 78] ( Olophrum ; synonym of obtectum ).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; synonym of obtectum ).

— Campbell, 1983: 608 ( Olophrum ; synonym of obtectum ).

emarginatum Erichson, 1840: 868 ( Olophrum ; [Note: Not so stated but presumably a replacement name for rotundicolle Say ]).

— Melsheimer, 1844: 43 ( Olophrum ; characters; found on blossoms of Red Maple).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; synonym of obtectum ).

— Campbell, 1983: 608 ( Olophrum ; synonym of obtectum ).

paphlagonicum Coiffait, 1978d: 172 ( Olophrum ; Type locality: Col de l’Ilgaz Dag, 1800 m).

- DISTRIBUTION: Turkey.

parvipenne Scheerpeltz, 1929 , see: consimile Gyllenhal, 1810 .

parvulum Mäklin, 1853 , see: consimile Gyllenhal, 1810 .

parvum Cameron, 1941 , see: kashmiricum Cameron, 1941 .

piceum Gyllenhal, 1810: 200 ( Omalium ; Type locality: Westmannia).

— Gyllenhal, 1827: 461 ( Omalium ; notes).

— Zetterstedt, 1828: 49 ( Omalium ; characters; Lapland).

— Curtis, 1829: 28 ( Anthobium ; catalog; Britain).

— Mannerheim, 1830: 51 ( Omalium ; Sweden).

— Mannerheim, 1831: 465 ( Omalium ; Sweden).

— Runde, 1835: 21 ( Omalium ; characters; Germany).

— Lacordaire, 1835: 485 ( Omalium ; characters; France).

— Erichson, 1839a: 623 ( Olophrum ; characters; Germany).

— Heer, 1839: 189 ( Olophrum ; characters; Switzerland).

— Stephens, 1839: 426 ( Olophrum ; characters; England).

— Erichson, 1840: 864 ( Olophrum ; characters; Germany; Hungary; England; Sweden).

— Laporte, 1840: 191 ( Omalium ; characters; Sweden; France).

— Hochhuth, 1849: 203 ( Olophrum ; characters; Caucasus).

— Redtenbacher, 1849: 749 ( Olophrum ; characters; Austria).

— Hardy, 1851: 57 ( Olophrum ; notes; England).

— Fairmaire and Laboulbène, 1856: 635 ( Olophrum ; characters; France).

— Redtenbacher, 1857: 247 ( Olophrum ; characters; Austria).

— Thomson, 1861: 193 ( Olophrum ; characters; Scandinavia).

— Hochhuth, 1862: 103 ( Olophrum ; Russia).

— Fauvel, 1871a: 123 [= 1872: 97] ( Olophrum ; characters; France; Scandinavia; Britain; Switzerland; Germany; Austria).

— Fauvel, 1873b: 118 [= 1873c: 11] ( Olophrum ; Sardinia; Russia).

— Redtenbacher, 1874: 268 ( Olophrum ; characters; Austria).

— Seidlitz, 1875: 240 ( Olophrum ; characters; Baltic region).

— Mulsant and Rey, 1880: 135 ( Olophrum ; characters; France).

— Fowler, 1888: 404 ( Olophrum ; characters; Britain).

— Ganglbauer, 1895: 720 ( Olophrum ; characters; north and middle Europe).

— Luze, 1905: 38 ( Olophrum ; characters; central and corthern Europe).

— Reitter, 1909: 186 ( Olophrum ; characters; Germany).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Sharp and Muir, 1912: 501 ( Olophrum ; characters of male genitalia).

— Johansen, 1914: 592 ( Olophrum ; characters; Denmark).

— Scheerpeltz, 1929: 29 ( Olophrum ; characters; Czech Republic, Germany, Holland, Belgium, France, Denmark, England, Sweden, Spain).

— Portevin, 1929: 440 ( Olophrum ; characters; France).

— Munster, 1935: 25 ( Olophrum ; characters; notes; Sweden).

— Paulian, 1941: 152 ( Olophrum ; larval characters; France).

— Palm, 1948: 105 ( Olophrum ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 109 ( Olophrum ; Britain; Norway; Sweden; Finland; Denmark; Holland; Belgium; France; Switzerland; Italy; Baltic region; Czech Republic; Slovakia; Spain; Germany; Austria).

— Lohse, 1964: 53 ( Olophrum ; characters; central Europe).

— Smetana, 1964c: 170 ( Olophrum ; characters).

— Smetana, 1964d: 54 ( Olophrum ; Czechoslovakia).

— Pototskaia, 1967: 20 ( Olophrum ; larval characters).

— Kasule, 1968a: 72 ( Olophrum ; life history).

— Hammond, 1971: 167 ( Olophrum ; notes; characters; Britain).

— Pope, 1977: 22 ( Olophrum ; Britain).

— Hammond, 1980: 136 ( Olophrum ; Ireland).

— Tóth, 1982: 84 ( Olophrum ; characters; Hungary).

— Zanetti, 1987: 301 ( Olophrum ; characters; Italy).

— Lucht, 1987: 88 ( Olophrum ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Welch, 1993: 227 ( Olophrum ; ovariole number and ovary structure).

— Boháč, 1993: 41 ( Olophrum ; Slovakia; Czech Republic).

— Zanetti, 1995: 8 ( Olophrum ; Italy).

— Owen, 1997a: 148 ( Olophrum ; collecting sites; Ireland).

— Cibuļskis, 1999: 35 ( Olophrum ; list; Latvia).

- DISTRIBUTION: Europe, Russia.

blattoides Ahrens, 1812: 20 ( Silpha , cited as Sylpha ; Type locality: im Harze).

— Erichson, 1839a: 623 ( Olophrum ; synonym of piceum ).

— Heer, 1839: 189 ( Olophrum ; synonym of piceum ).

— Redtenbacher, 1849: 749 ( Olophrum ; synonym of piceum ).

— Erichson, 1840: 864 ( Olophrum ; synonym of piceum ).

— Kraatz, 1857d: 940 ( Olophrum ; synonym of piceum ).

— Mulsant and Rey, 1880: 136 ( Olophrum ; synonym of piceum ).

— Ganglbauer, 1895: 720 ( Olophrum ; synonym of piceum ).

— Luze, 1905: 38 ( Olophrum ; synonym of piceum ).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; synonym of piceum ).

puncticolle Eppelsheim, 1880 a: 212 ( Olophrum ; Type locality: Südungarn).

— Ganglbauer, 1895: 720 ( Olophrum ; characters; Hungary; Bosnia).

— Scheerpeltz, 1929: 97 ( Olophrum ; characters; Croatia; Siebenbürgen; Bosnia; Herzegowina).

— Tóth, 1982: 86 ( Olophrum ; characters; Hungary).

- DISTRIBUTION: Austria, Hungary, Bosnia Hercegovina, Croatia, Romania, Turkey, Syria.

quesneli Hatch, 1957 , see: boreale Paykull, 1792 .

recticolle Scheerpeltz, 1929 , see: consimile Gyllenhal, 1810 .

reitteri Scheerpeltz, 1929: 83 ( Olophrum ; Type locality: Russia merid.).

- DISTRIBUTION: Russia.

rotundicolle Say, 1832 , see: obtectum Erichson, 1840 .

rotundicolle C. Sahlberg, 1830: 281 ( Omalium ; Type locality: Lapponia).

— Mannerheim, 1830: 51 ( Omalium ; Lapland).

— Mannerheim, 1831: 465 ( Omalium ; Lapland).

— Runde, 1835: 21 ( Omalium ; characters; Germany).

— Zetterstedt, 1838: 49 ( Omalium ; characters; habitat; Lapland).

— Erichson, 1840: 866 ( Olophrum ; characters; Finland).

— Thomson, 1861: 194 ( Olophrum ; characters; Scandinavia).

— Seidlitz, 1875: 240 ( Olophrum ; characters; Baltic region).

— J. Sahlberg, 1876: 213 ( Olophrum ; Finland).

— Fauvel, 1878d: 243 [= 1878e: 79] ( Olophrum ; characters; Lake Superior).

— Ganglbauer, 1895: 721 ( Olophrum ; characters; Sudeten; Finland; Lapland; North America ).

— Luze, 1905: 46 ( Olophrum ; characters; Lithuania; Russia; Finland; Lapland; Norway; North America ).

— Reitter, 1909: 186 ( Olophrum ; characters).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Blatchley, 1910: 479 ( Olophrum ; characters; Ohio).

— Scheerpeltz, 1929: 163 ( Olophrum ; characters; Sudeten; Norway; Sweden; Finland; Russia).

— Munster, 1936: 101 ( Olophrum ; characters; notes).

— Palm, 1948: 108 ( Olophrum ; characters; Sweden; Norway; Finland).

— Hatch, 1957: 63 ( Olophrum ; characters; British Columbia ).

— Horion, 1963: 116 ( Olophrum ; Russia; Finland; Sweden; Norway; Germany; Poland; Czech Republic).

— Lohse, 1964: 53 ( Olophrum ; characters; central Europe).

— Smetana, 1964c: 164 ( Olophrum ; Czechoslovakia).

— Smetana, 1964c: 172 ( Olophrum ; characters).

— Smetana, 1966: 325 ( Olophrum ; notes; Czechoslovakia).

— Ashworth, 1972: 214 ( Olophrum ; late-glacial subfossil; Britain).

— Schwert and Morgan, 1980: 101 ( Olophrum ; late glacial fossil; New York).

— Miller and Morgan, 1982: 264 ( Olophrum ; Postglacial fossils; New York).

— Campbell, 1983: 615 ( Olophrum ; characters; Alberta; British Columbia ; Manitoba; Newfoundland; Northwest Territories; Ontario; Quebec; Saskatchewan; Yukon Territory; Alaska; Minnesota; New York).

— Lucht, 1987: 89 ( Olophrum ; checklist; Germany; Poland; Czechoslovakia).

— Ryabukhin, 1991b: 23 ( Olophrum ; characters; Russia).

— Elias, 1991: 556 ( Olophrum ; Quaternary fossil collection; Colorado).

— Boháč, 1993: 41 ( Olophrum ; Czech Republic).

— Downie and Arnett, 1996: 432 ( Olophrum ; characters; Canada; USA).

— Lavoie, Elias, and Payette, 1997: 231 ( Olophrum ; Holocene fossil; subarctic Quebec).

— Ryabukhin, 1999: 23 ( Olophrum ; catalog; habitat; distribution; northeastern Russia).

- DISTRIBUTION: Lithuania, Finland, Norway, Sweden, Germany, Poland, Czech Republic, Russia, Canada, USA.

convexicolle LeConte, 1850: 221 ( Lathrium ; Type locality: Eagle Harbor, Lake Superior; [Note: See lectotype designation by Campbell, 1983]).

— Fauvel, 1878d: 242 [= 1878e: 79] ( Olophrum ; synonym of rotundicolle ).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; synonym of rotundicolle ).

— Campbell, 1983: 615 ( Olophrum ; lectotype designation; synonym of rotundicolle ).

convexum Mäklin, 1853: 195 ( Olophrum ; Type locality: peninsula Kenai ; [Note: See lectotype designation by Campbell, 1983]).

— Fauvel, 1878d: 244 [= 1878e: 80] ( Olophrum ; characters; Alaska).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 168 ( Olophrum ; characters; Alaska).

— Steel, 1964a: 248 ( Olophrum ; synonym of rotundicolle ).

— Campbell, 1983: 615 ( Olophrum ; lectotype designation; synonym of rotundicolle ).

roubali Scheerpeltz, 1929: 47 ( Olophrum ; Type locality: Nordkaukasus, Talsystem des Teberda-Flusses; Gebiet des Kluchor-Passes).

- DISTRIBUTION: Russia.

roubalianum Scheerpeltz, 1929: 53 ( Olophrum ; Type locality: Nordkaukasus, Talsystem des Teberda).

- DISTRIBUTION: Russia.

scheerpeltzi Bernhauer, 1938: 18 ( Olophrum ; Type locality: China: Provinz Fukien , Foochow). - DISTRIBUTION: China.

schusteri Scheerpeltz, 1929 , see: fuscum Gravenhorst, 1806 .

simplex Sharp, 1874: 97 ( Olophrum ; Type locality: Nagasaki).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 108 ( Olophrum ; characters; Japan).

— Watanabe, 1990: 139 ( Olophrum ; characters; Japan).

— J. Li, 1993: 19 ( Olophrum ; China).

- DISTRIBUTION: Japan, China.

sinense Scheerpeltz, 1929: 28 , 64 ( Olophrum ; Type locality: Umgebung von Shanghai).

- DISTRIBUTION: China.

stouti Hatch, 1957: 63 ( Olophrum ; Type locality: Washington: Chehalis. Also cited from Oregon and British Columbia ).

- DISTRIBUTION: Canada, USA.

subsolanum Watanabe, 1990: 132 ( Olophrum ; Type locality: Japan: Honshu: Shizuoka Pref.: Odaru-onsen, Izu ).

- DISTRIBUTION: Japan.

tadashii Watanabe, 1990: 135 ( Olophrum ; Type locality: Japan: Honshu: Tochigi Pref.: near Sainoko Lake, Oku-Nikkô ).

- DISTRIBUTION: Japan.

transversicolle Luze, 1905: 41 ( Olophrum ; Type locality: Osttirol, Taufers, 850 m).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; synonym of fuscum ).

— Scheerpeltz, 1929: 40 ( Olophrum ; characters; Alps).

— Munster, 1935: 28 ( Olophrum ; variety of fuscum ; notes).

— Horion, 1935: 170 ( Olophrum ; characters; Austria).

— Horion, 1963: 110 ( Olophrum ; Austria; Italy).

— Lohse, 1964: 54 ( Olophrum ; characters; central Europe).

— Smetana, 1964c: 164, 171 ( Olophrum ; notes; characters; Czechoslovakia).

— Zanetti, 1987: 302 ( Olophrum ; characters; Italy).

— Lucht, 1987: 89 ( Olophrum ; checklist; Austria).

— Boháč, 1993: 41 ( Olophrum ; Slovakia; Czech Republic).

— Zanetti, 1995: 8 ( Olophrum ; Italy).

- DISTRIBUTION: Italy, Austria, Czechoslovakia.

vicinum Sharp, 1889: 472 ( Olophrum ; Type locality: Iwakisan).

— Bernhauer and Schubert, 1910: 67 ( Olophrum ; catalog).

— Scheerpeltz, 1929: 103 ( Olophrum ; characters; Japan).

— Watanabe, 1990: 133 ( Olophrum ; characters; Japan).

viennense Scheerpeltz, 1929: 90 ( Olophrum ; Type locality: Mariabrunn b. Wien. Also cited from Slovakia, Hungary).

— Horion, 1963: 112 ( Olophrum ; Austria; Czech Republic; Slovakia; Hungary).

— Lohse, 1964: 54 ( Olophrum ; characters; central Europe).

— Smetana, 1964c: 171 ( Olophrum ; characters).

— Tóth, 1982: 86 ( Olophrum ; characters; Hungary).

— Lucht, 1987: 89 ( Olophrum ; checklist; Czechoslovakia; Austria).

— Boháč, 1993: 41 ( Olophrum ; Slovakia; Czech Republic).

- DISTRIBUTION: Austria, Hungary, Slovakia.

yasudai Watanabe, 1990: 139 ( Olophrum ; Type locality: Japan: Hokkaido: Mt. Kurodake on Daisetsu Mts. ).

- DISTRIBUTION: Japan.

Omalorphanus

[1 species; Nearctic Region]

Omalorphanus Campbell and Chandler, 1987: 315 (species included: aenigma ). TYPE SPECIES: Omalorphanus aenigma Campbell and Chandler , fixed by original designation and monotypy.

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (1 Nearctic species; notes; characters in key). aenigma Campbell and Chandler, 1987: 325 ( Omalorphanus ; Type locality: Oregon: Lane Co., H. J. Andrews Exp. For., Road 1506, 4000 feet).

- DISTRIBUTION: USA.

Orobanus

[7 species; Nearctic Region]

Orobanus LeConte, 1878: 453 (species included: simulator ). TYPE SPECIES: Orobanus simulator LeConte , fixed by monotypy.

— Fauvel, 1878d: 249 [= 1878e: 83] (characters).

— Duvivier, 1883: 194 (catalog).

— Casey, 1886: 247 (key to species).

— Casey, 1894: 405 (notes; key to North American species).

— Bernhauer and Schubert, 1910: 74 (world catalog; 3 species).

— Leng, 1920: 95 (catalog of North American species).

— Mank, 1934: 123 (key to species).

— Blackwelder, 1952: 277 (type species: simulator ).

— Hatch, 1957: 51, 72 (characters; key to species of Pacific Northwest).

— Moore, 1966: 48 (characters; notes).

— Moore and Legner, 1975: 203 (catalog; North America north of Mexico).

— Campbell, 1978: 1 (moved to Anthophagini ).

— Moore and Legner, 1979: 187 (characters; notes).

— Campbell and Davies, 1991: 90 (checklist of species; Canada).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (7 Nearctic species; notes; characters in key). densus Casey, 1886: 246 ( Orobanus ; Type locality: California: San Diego).

— Casey, 1894: 406 ( Orobanus ; characters; Sonoran region).

— Bernhauer and Schubert, 1910: 74 ( Orobanus ; catalog).

— Mank, 1934: 123 ( Orobanus ; characters in key).

- DISTRIBUTION: USA.

falli Mank, 1934: 122 ( Orobanus ; Type locality: Redwood Park, Contra Costa Co., California). — Hatch, 1957: 73 ( Orobanus ; characters; British Columbia ; Washington; Oregon).

- DISTRIBUTION: Canada, USA.

montanus Mank, 1934: 121 ( Orobanus ; Type locality: Lake St. Mary near Sun Camp, Glacier National Park , Montana).

- DISTRIBUTION: USA.

mormonus Mank, 1934: 122 ( Orobanus ; Type locality: Ogden, Ut[ah]).

- DISTRIBUTION: USA.

rufipes Casey, 1886: 245 ( Orobanus ; Type locality: California: Hoopa Val., Humboldt Co.). — Casey, 1894: 406 ( Orobanus ; characters; Pacific region).

— Bernhauer and Schubert, 1910: 74 ( Orobanus ; catalog).

— Mank, 1934: 123 ( Orobanus ; characters in key).

— Hatch, 1957: 72 ( Orobanus ; characters; British Columbia ; Washington; Oregon; Idaho).

- DISTRIBUTION: Canada, USA.

simulator LeConte, 1878: 453 ( Orobanus ; Type locality: Leavenworth Valley, 9000 to 10,000 feet; Vancouver Island; Gilroy and Holcomb Valley , California).

Fauvel, 1878d: 249 [= 1878e: 83] ( Orobanus ; characters; Colorado; California; Vancouver Island ) .

— Casey, 1894: 405 ( Orobanus ; characters; Rocky Mts.).

— Bernhauer and Schubert, 1910: 74 ( Orobanus ; catalog).

— Mank, 1934: 123 ( Orobanus ; characters in key).

— Hatch, 1957: 73 ( Orobanus ; characters; British Columbia ; Washington; Oregon).

- DISTRIBUTION: Canada, USA.

tarsalis Hatch, 1957: 73 ( Orobanus ; Type locality: Oregon: McMinnville. Also cited from British Columbia and Washington).

- DISTRIBUTION: Canada, USA.

Orochares

[6 species; Nearctic and Palaearctic Regions]

Orochares Kraatz, 1857d: 955 (species included: angustatus ). TYPE SPECIES: Deliphrum angustatum Erichson , fixed by monotypy.

— Redtenbacher, 1857: 993 (characters).

— Schaum, 1859: 31 (catalog; European species).

— Wencker and Silbermann, 1866: 36 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 662 (catalog).

— Fauvel, 1871a: 126 [= 1872: 100] (characters).

— Redtenbacher, 1874: 270 (characters).

— Fauvel, 1875a:VII [= 1875b: 209] (catalog).

— Mulsant and Rey, 1880: 89 (characters).

— Seidlitz, 1891: 342 (key to some European species).

— Seidlitz, 1891a: 364 (key to some European species).

— Heyden, Reitter, and Weise, 1891: 118 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 725 (characters).

— Heyden, Reitter, and Weise, 1906: 133 (list of species of Europe and Caucasus).

— Reitter, 1909: 184 (characters).

— Bernhauer and Schubert, 1910: 62 (world catalog; 1 species).

— Johansen, 1914: 591 (characters).

— Leng, 1920: 94 (catalog of North American species).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Porta, 1926: 24 (characters; species of Italy).

— Joy, 1932: 95 (key to British species).

— Scheerpeltz, 1933: 1053 (world catalog supplement).

— Kloet and Hincks, 1945: 161 (list of British species).

— Palm, 1948: 93 (characters).

— Tottenham, 1949: 356, 407 (type species: angustatum Erichson ; list of British species).

— Blackwelder, 1952: 277 (type species: angustata ).

— Tottenham, 1954: 28 (characters; British species).

— Horion, 1963: 99 (list of central European species).

— Székessy, 1963: 16 (characters).

— Lohse, 1964: 50 (characters; key to central European species).

— Scheerpeltz, 1968a: 12 (catalog of Austrian species).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Moore and Legner, 1975: 204 (catalog; North America north of Mexico).

— Shibata, 1976: 108 (checklist of species of Japan).

— Pope, 1977: 23 (list of British species).

— Burakowski, Mroczkowski, and Stefańska, 1979: 52 (catalog; Poland).

— Moore and Legner, 1979: 188 (characters; notes).

— Muona, 1979: 18 (list of Scandinavian species).

— Tóth, 1982: 70 (characters; habitus; species of Hungary).

— Campbell, 1984a: 1234 (characters; review of and key to North American species).

— Zanetti, 1987: 268 (characters; habitus; species of Italy).

— Watanabe, 1990: 260 (characters; species of Japan).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Thayer, 1993: 286 (checklist; synonymy; notes; discussion).

— Zanetti, 1995: 8 (list of species of Italy).

— M. Hansen, 1996: 95 (list of Danish species).

— Downie and Arnett, 1996: 433 (characters; key to species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (2 Nearctic species; notes; characters in key). Paradeliphrum Hatch, 1957: 50 , 67 (species included: tumidum , inflatum ). TYPE SPECIES:

Paradeliphrum tumidum Hatch , fixed by original designation.

— Moore, 1966: 48, 51 (characters; notes).

— Moore and Legner, 1975: 205 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 189 (characters; notes).

— Campbell, 1984a: 1234 (synonym of Orochares ).

— Thayer, 1993: 286 (synonym of Orochares ). Osellia Zanetti, 1983: 423 (species included: calabra ). TYPE SPECIES: Osellia calabra

Zanetti, fixed by original designation.

— Zanetti, 1987: 278 (characters; habitus; species of Italy).

— Thayer, 1993: 286 (synonym of Orochares ). angustatus Erichson, 1840: 874 ( Deliphrum ; Type locality: Thuringia, Austria).

— Redtenbacher, 1849: 750 ( Deliphrum ; characters; Austria).

— Fairmaire and Laboulbène, 1856: 638 ( Deliphrum ; characters; France).

— Redtenbacher, 1857: 249 ( Deliphrum ; characters; Austria).

— Kraatz, 1857d: 956 ( Orochares ; characters; Germany).

— Fauvel, 1871a: 126 [= 1872: 100] ( Orochares ; characters; France; Germany; Austria).

— Redtenbacher, 1874: 270 ( Orochares ; characters; Austria).

— Seidlitz, 1875: 236 ( Orochares ; characters; Baltic region).

— Mulsant and Rey, 1880: 91 ( Orochares ; characters; France).

— Ganglbauer, 1895: 726 ( Orochares ; characters; Germany; France; North America ).

— Sainte-Claire Deville, 1907: 41 ( Orochares ; characters; France).

— Reitter, 1909: 185 ( Orochares ; characters; Germany).

— Bernhauer and Schubert, 1910: 62 ( Orochares ; catalog).

— Johansen, 1914: 591 ( Orochares ; characters; Denmark).

— Portevin, 1929: 438 ( Orochares ; characters; France).

— Scheerpeltz, 1933: 1054 ( Orochares ; catalog).

— Palm, 1948: 94 ( Orochares ; characters; Denmark).

— Horion, 1963: 99 ( Orochares ; Britain; Denmark; Holland; Belgium; France; Switzerland; Italy;

Czechoslovakia; Germany; Austria).

— Lohse, 1964: 50 ( Orochares ; characters; central Europe).

— Pope, 1977: 23 ( Orochares ; Britain).

— Tóth, 1982: 70 ( Orochares ; characters; Hungary).

— Campbell, 1984a: 1237 ( Orochares ; characters; Massachusetts).

— Segers, 1986: 21 ( Orochares ; checklist of species; Belgium).

— Zanetti, 1987: 270 ( Orochares ; characters; Italy).

— Lucht, 1987: 88 ( Orochares ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Thayer, 1993: 286 ( Orochares ; checklist; Europe; eastern USA?).

— Boháč, 1993: 41 ( Orochares ; Slovakia; Czech Republic).

— Zanetti, 1995: 8 ( Orochares ; Italy).

— Downie and Arnett, 1996: 433 ( Orochares ; characters; USA).

— Cibuļskis, 1999: 35 ( Orochares ; list; Latvia).

- DISTRIBUTION: Britain, Denmark, Netherlands, Belgium, Germany, France, Switzerland, Italy, Austria, Hungary, Slovakia, Czech Republic, USA.

calaber Zanetti, 1983: 429 ( Osellia ; Type locality: Montalto, Aspromonte, Reggio di Calabria, 1850 m).

— Zanetti, 1987: 280 ( Osellia ; characters; Italy).

— Thayer, 1993: 286 ( Orochares ; checklist; Calabria).

— Sparacio, 1995: 142 ( Osellia ; characters; notes; Sicily).

— Zanetti, 1995: 8 ( Orochares ; Italy).

- DISTRIBUTION: Italy.

siculus Zanetti, 1983: 434 ( Osellia ; subspecies of calabra [sic]; Type locality: M. Soro, Madonie, Messina, 1840 m).

— Zanetti, 1987: 280 ( Osellia ; subspecies of calabra [sic]; characters; Italy).

— Thayer, 1993: 286 ( Orochares ; subspecies of calaber ; Sicily).

— Zanetti, 1995: 8 ( Orochares ; subspecies of calaber ; Italy).

debilis Sharp, 1889: 472 ( Amphichroum ; Type locality: Yokohama).

— Bernhauer and Schubert, 1910: 70 ( Amphichroum ; catalog).

— Watanabe, 1990: 262 ( Orochares ; characters; Japan).

— Thayer, 1993: 286 ( Orochares ; checklist; Honshu).

- DISTRIBUTION: Japan.

japonicus Cameron, 1933 a: 168 ( Orochares ; Type locality: Japan: Kobe ).

— Thayer, 1993: 286 ( Orochares ; synonym of debilis ).

japonicus Cameron, 1933 , see: debilis Sharp, 1889 .

siculus Zanetti, 1983 , see: calaber Zanetti, 1983 .

suteri Campbell, 1984 a: 1240 ( Orochares ; Type locality: USA: Wisconsin: Kenosha Co.: County Line Woods. Also cited from Illinois).

— Thayer, 1993: 286 ( Orochares ; checklist; Wisconsin; Illinois).

— Downie and Arnett, 1996: 433 ( Orochares ; characters; USA).

- DISTRIBUTION: USA.

tumidus Hatch, 1957: 67 ( Paradeliphrum ; Type locality: Oregon: McMinnville. Also cited from British Columbia ).

— Campbell, 1984a: 1239 ( Orochares ; characters; British Columbia ; Oregon).

— Campbell and Davies, 1991: 91 ( Orochares ; checklist of species; Canada).

— Thayer, 1993: 286 ( Orochares ; checklist; Oregon; British Columbia ).

- DISTRIBUTION: Canada, USA.

villiersi Jarrige, 1971: 488 ( Orochares ; Type locality: Iran: Dacht Argen, 60 km O. de Chiraz).

— Thayer, 1993: 286 ( Orochares ; checklist; Iran).

- DISTRIBUTION: Iran.

Paramannerheimia [1 species; Palaearctic Region]

Paramannerheimia Scheerpeltz, 1976: 8 (species included: janetscheki ). TYPE SPECIES: Paramannerheimia janetscheki Scheerpeltz , fixed by original designation.

janetscheki Scheerpeltz, 1976: 9 ( Paramannerheimia ; Type locality: Nepal: Tarnga [= Thanke] nach Ringmo, 3800–4370 m).

Paratrichodromeus

[2 species; Palaearctic Region]

Paratrichodromeus Zerche, 1992 a: 122 (species included: sukranensis , punctulatus ). TYPE

SPECIES: Paratrichodromeus sukranensis Zerche , fixed by original designation. punctulatus Luze, 1904: 77 ( Geodromicus ; Type locality: Seravschan: Kschut: Artutsch;

Jagnob: Varsaut).

— Bernhauer and Schubert, 1910: 77 ( Geodromicus ; catalog).

— Zerche, 1992a: 123 ( Paratrichodromeus ; Tadzhikistan).

- DISTRIBUTION: Tajikistan, Uzbekistan.

sukranensis Zerche, 1992 a: 124 ( Paratrichodromeus ; Type locality: Asia centr., Tadsch.,

Dschirgatalski Raj., Sukran-Schlucht, 2590 m / Burs-Gletscher-Fuss).

- DISTRIBUTION: Tajikistan.

Pelecomalium

[12 species; Nearctic Region]

Pelecomalium Casey, 1886: 241 (species included: binotatum , modestum ). TYPE SPECIES: Pelecomalium binotatum Casey , fixed by subsequent designation by Blackwelder, 1952: 296.

— Casey, 1894: 412 (notes; key to North American species).

— Bernhauer and Schubert, 1910: 75 (world catalog; 13 species).

— Leng, 1920: 95 (catalog of North American species).

— Hatch, 1957: 49, 55 (characters; key to species of Pacific Northwest).

— Moore, 1966: 48, 49 (characters; notes).

— Moore and Legner, 1975: 206 (catalog; North America north of Mexico).

— Moore and Legner, 1979: 190 (characters; notes).

— Campbell and Davies, 1991: 90 (checklist of species; Alaska and Canada).

— Downie and Arnett, 1996: 432 (characters; species of Northeastern North America ).

— Newton, Thayer, Ashe, and Chandler, 2000: 340 (12 Nearctic species; notes; characters in key).

Heterops Mannerheim, 1843b: 234 1852: 380 (species included: testaceus ; [preoccupied by Blanchard, 1842]; [Note: This name was first published in 1843 as a junior synonym but was made available in a 1852 catalog citation when an available nominal species, “ testaceus Eschscholtz ”, was included. The second included nominal species was unavailable]. TYPE SPECIES: Arpedium testaceum Mannerheim , fixed by monotypy by Mannerheim, 1843b.

— 1852: 380 (catalog; name made available by inclusion of available nominal species).

— Blackwelder, 1952: 185 (type species; synonym of Pelecomalium ).

Pelekyomalium Eichelbaum, 1909: 100 (emendation of Pelecomalium ; synonym of Amphichroum ). TYPE SPECIES: Pelecomalium binotatum Casey , fixed by objective synonymy with Pelecomalium .

— Blackwelder, 1952: 297 (type species; synonym of Pelecomalium ).

alutaceum Casey, 1886: 236 ( Amphichroum ; Type locality: California: Marin Co.).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

binotatum Casey, 1886 , see: testaceum Mannerheim, 1843 .

crassicorne Casey, 1886: 239 ( Amphichroum ; Type locality: California: Siskiyou Co.).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

flavescens Casey, 1894: 414 ( Pelecomalium ; Type locality: California: Lake Tahoe ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

laevicolle LeConte, 1866: 375 ( Amphichroum ; Type locality: Lycoming County, Pennsylvania).

— Fauvel, 1878d: 238 [= 1878e: 74] ( Amphichroum ; characters; Pennsylvania).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

— Downie and Arnett, 1996: 432 ( Pelecomalium ; characters; USA).

- DISTRIBUTION: USA.

modestum Casey, 1886 , see: veterator Casey, 1886 .

nigrum Hatch, 1957 [Note: Nomen nudum], see: testaceum Mannerheim, 1843 .

opaculum Fauvel, 1878d: 241 [= 1878e: 77] ( Amphichroum ; Type locality: Californie, Mariposa ).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

— Hatch, 1957: 56 ( Pelecomalium ; characters; Washington).

- DISTRIBUTION: USA.

pallior Hatch, 1957: 56 ( Pelecomalium ; aberration of opaculum ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; locality cited: Seattle, Washington).

pallidum Casey, 1894: 415 ( Pelecomalium ; Type locality: California: Lake Tahoe; Nevada: Reno ).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

pallior Hatch, 1957 [Note: Nomen nudum], see: opaculum Fauvel, 1878 .

pilosellum Casey, 1886: 237 ( Amphichroum ; Type locality: California: Lake Co.).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

puberulum Fauvel, 1878d: 240 [= 1878e: 76] ( Amphichroum ; Type locality: Californie, San Diego ).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

— Hatch, 1957: 55 ( Pelecomalium ; characters; British Columbia ; Washington; Oregon; Idaho). - DISTRIBUTION: Canada, USA.

ruficeps Motschulsky, 1860 , see: Anthophagus .

scutatum Fauvel, 1878d: 238 [= 1878e: 74] ( Amphichroum ; Type locality: Californie, Mariposa ).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

sparsum Fauvel, 1878d: 237 [= 1878e: 73<†125> ( Amphichroum ; Type locality: Californie, Mariposa ).

— Casey, 1894: 413 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

testaceum Mannerheim, 1843b: 234 ( Arpedium ; [Note: Cited with Eschscholtz as author]; Type locality: insula Sitkha).

— Fauvel, 1878d: 239 [= 1878e: 75] ( Amphichroum ; characters; Alaska; British Columbia ; California).

— Casey, 1894: 413 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

— Van Dyke, 1924: 16 ( Amphichroum ; subgenus Pelecomalium ; Alaska).

— Fall, 1926: 145 ( Pelecomalium ; Alaska).

— Hatch, 1957: 55 ( Pelecomalium ; characters; British Columbia ; Washington; Oregon).

- DISTRIBUTION: Canada, USA.

binotatum Casey, 1886: 242 ( Pelecomalium ; Type locality: California: Marin Co.).

— Casey, 1894: 413 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; cited as binolatum; catalog).

— Hatch, 1957: 55 ( Pelecomalium ; aberration of testaceum ; characters).

nigrum Hatch, 1957: 55 ( Pelecomalium ; aberration of testaceum ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; localities cited: Washington, British Columbia , Washington, Oregon).

veterator Casey, 1886: 238 ( Amphichroum ; Type locality: California: Lake Co.).

— Casey, 1894: 414 ( Pelecomalium ; synonym of modestum ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; catalog).

- DISTRIBUTION: USA.

modestum Casey, 1886: 242 ( Pelecomalium ; Type locality: California: Lake Co.).

— Casey, 1894: 414 ( Pelecomalium ).

— Bernhauer and Schubert, 1910: 75 ( Pelecomalium ; synonym of veterator ).

Philorinum

[9 species; Palaearctic, Ethiopian, and Oriental Regions]

Philorinum Kraatz, 1857d: 965 (species included: humile , nitidulum ). TYPE SPECIES: Omalium humile Erichson , fixed by subsequent designation by Jacquelin du Val, 1868: 80.

— Redtenbacher, 1857: 992 (characters).

— G. Waterhouse, 1858: 31 (catalog of British species).

— Schaum, 1859: 32 (catalog; European species).

— Wencker and Silbermann, 1866: 36 (list of species; collecting notes; France).

— Gemminger and Harold, 1868: 663 (catalog).

— Fauvel, 1871a: 109 [= 1872: 83] (characters).

— Sharp, 1871d: 14 (list of British species).

— Redtenbacher, 1874: 272 (characters).

— Fauvel, 1875a:VI [= 1875b: 208] (catalog).

— Mulsant and Rey, 1880: 153 (characters).

— Fowler, 1888: 408 (notes; cited as Philorhinum ).

— Heyden, Reitter, and Weise, 1891: 119 (list of species of Europe and Caucasus).

— Ganglbauer, 1895: 727 (characters).

— Heyden, Reitter, and Weise, 1906: 132 (list of species of Europe and Caucasus).

— Bernhauer and Schubert, 1910: 61 (world catalog; 3 species).

— Petri, 1912: 54 (list of species of Siebenbürgen).

— Winkler, 1925: 332 (catalog for Palaearctic region).

— Cameron, 1925: 11 (catalog of Indian species).

— Porta, 1926: 28 (characters; key to species of Italy).

— Portevin, 1929: 442 (characters; key to French species).

— Cameron, 1930: 148 (cited as Philorhinum ; characters).

— Joy, 1932: 103 (key to British species).

— Scheerpeltz, 1933: 1052 (world catalog supplement).

— Normand, 1935: 357 (list of species of Tunisia).

— Kloet and Hincks, 1945: 161 (list of British species).

— Tottenham, 1949: 356, 407 (type species: sordidum Stephens = humile Erichson ; list of British species).

— Tottenham, 1954: 28 (characters; British species).

— Kocher, 1958: 94 (checklist of species; Morocco).

— Ferreira, 1962a: 9 (list of Portuguese species).

— Horion, 1963: 96 (list of central European species).

— Lohse, 1964: 48 (characters; key to central European species).

— Scheerpeltz, 1968a: 12 (catalog of Austrian species).

— Steel, 1970a: 16 (larval characters; biological notes).

— Tikhomirova, 1973a: 139 (checklist of species of USSR).

— Pope, 1977: 24 (list of British species).

— Topp, 1978: 308 (larval characters).

— Tóth, 1982: 73 (characters; habitus).

— Zanetti, 1987: 326 (characters; habitus; key to species of Italy).

— Silfverberg, 1992: 22 (list of species of Fennoscandia, Denmark, and the Baltic States).

— Welch, 1993 (ovariole structure and development).

— Boháč, 1993: 41 (in list of species of Czech Republic and Slovakia).

— Zanetti, 1995: 8 (list of species of Italy).

— M. Hansen, Liljehult, Mahler, and Pedersen, 1995: 27 (additional records to Danish list of species).

— M. Hansen, 1996: 95 (list of Danish species).

— M. Hansen, Mahler, Palm, and Pedersen, 1996: 240 (additions to list of Danish species). afghanicum Coiffait, 1982b: 80 ( Philorinum ; Type locality: Afghanistan: Nouristan, Kamdesch, 2200 m).

- DISTRIBUTION: Afghanistan.

africanum Bernhauer, 1912f: 178 ( Philorinum ; Type locality: Usambara).

— Scheerpeltz, 1933: 1052 ( Philorinum ; catalog).

- DISTRIBUTION: Tanzania.

brevipenne Luze, 1910 , see: sordidum Stephens, 1834 .

cadomense Fauvel, 1863 , see: sordidum Stephens, 1834 .

chinense Jarrige, 1948: 39 ( Philorinum , cited as Philorhinum ; Type locality: Yunnan: Pe-Yen- Tsing).

- DISTRIBUTION: China.

floricola Champion, 1920 , see: florivagum Champion, 1923 .

floricola Wollaston, 1864: 602 ( Philorinum ; Type locality: Grand Canary. Teneriffe: above Agua Mansa; Yeod el Alto; Agua Garcia; Taganana. Palma. Hierro).

— Wollaston, 1865: 521 ( Philorinum ; habitat; notes; Canary Islands).

— Fauvel, 1902b: 55 ( Philorinum ; synonym of sordidum ).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; catalog).

— Hernández, Outerelo, and Gamarra, 1994: 187 ( Philorinum ; Canary Islands).

— Assing and Wunderle, 1999: 437 ( Philorinum ; Canary Islands).

- DISTRIBUTION: Canary Islands.

florivagum Champion, 1923: 80 ( Philorhinum ; replacement name for floricola Champion ). — Cameron, 1930: 149 ( Philorinum , cited as Philorhinum ; characters; India).

— Scheerpeltz, 1933: 1052 ( Philorinum ; catalog).

— Scheerpeltz, 1960: 1 ( Philorinum ; Pakistan; Afghanistan).

— Coiffait, 1984: 118 ( Philorinum ).

- DISTRIBUTION: India, Pakistan, Afghanistan.

floricola Champion, 1920: 242 ( Philorhinum ; [preoccupied]; Type locality: W. Almora in Kumaon; Siuni Gad, Garhwal, alt. 4000 ft.).

— Cameron, 1930: 149 ( Philorinum ; synonym of florivagum ).

— Scheerpeltz, 1933: 1052 ( Philorinum ; synonym of florivagum ).

hopffgarteni Eppelsheim, 1892 a: 297 ( Philorinum ; Type locality: Siebenbürgen ; Griechenland, Morea).

— Ganglbauer, 1895: 728 ( Philorinum ; characters; Siebenbürgen; Greece).

— Bernhauer and Schubert, 1910: 61 ( Philorinum ; variety of sordidum ).

— Tóth, 1982: 74 ( Philorinum ; cited as valid species; characters).

- DISTRIBUTION: Romania, Greece.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

Genus

Anthobium

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