Anthracus hornburgi, Jaeger, 2015

Jaeger, Bernd, 2015, Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina), Linzer biologische Beiträge 47 (2), pp. 1361-1396 : 1371-1373

publication ID	https://doi.org/ 10.5281/zenodo.5193661
DOI	https://doi.org/10.5281/zenodo.5195689
persistent identifier	https://treatment.plazi.org/id/5C580E35-A81B-F11B-FF5E-FDD6FEA1986B
treatment provided by	Felipe
scientific name	Anthracus hornburgi
status	sp. nov.

Treatment

Anthracus hornburgi View in CoL nov.sp. ( Figs 16-18 View Figs 16-18 , 56-61a View Figs 56-61 , 81 View Fig 81 )

Acupalpus annamensis BATES View in CoL : ANDREWES 1947: 8 (specimens from Rangoon partim.)

Acupalpus annamensis BATES View in CoL : LANDIN 1954: 457 (specimens from Rangoon partim.)

Type material: Holotype: ♂ ( MMB) labelled " East Pakistan (EP 22) / Chittagong, Nasirabad / H.S., on light, 18.-22. / leg. Fr. Dvořák X.1970 ", "Moravian museum / Collectio / F. Dvořák" and "HOLOTYPE ♂ / Anthracus View in CoL / hornburgi sp. n. / des. B. Jaeger 2015" [red label].

Paratypes: 2♂♂, 1♀ (cWRA, cJAE) labelled " MYANMAR (Yangon) / Pègū / 60km NNE Yangon / 17°19'N / 96°28'E (lux) / 22.XI.2003 M.Hornburg", "COLL. WRASE / BERLIN". GoogleMaps 2♂♂ ( BMNH, ZIN) labelled " RANGOON / BURMA / 1/12 Malaise", one ♂ additionally with "Acupalpus / annamensis Bat. / H.E. Andrewes det." and "St. Petersburg / Zool. Inst.", and the ♂ other additionally with "Brit. Mus. / 1947-14." 1♂ ( BMNH) labelled "Tharrawady / Burma ", "H.E. Andrewes Coll. / B.M. 1945-97." and "Acupalpus / spec. / annamensis group / det. B. Jaeger 2010". All paratypes additionally with my label "PARATYPE ♂ or ♀ / Anthracus / hornburgi sp. n. / des. B. Jaeger 2015 " [red].

Remarks: One of the paratypes from Pègū is moderately immature with pronotum markedly deformed, and the paratypes from Rangoon are moderately to markedly immature.

Etymology: The species is dedicated to Michael Hornburg, Berlin, Germany, specialist on Jewel beetles who collected some of the paratypes of the species.

Description: General appearance as figured ( Fig. 16 View Figs 16-18 ). Body length 2.9-3.5 mm (HT 3.5); width 1.1-1.4 mm.

Shiny, pronotum slightly, elytra moderately iridescent. Head darker reddish yellow, with clypeus, labrum and mandibles (inner margins and apices blackish) paler, and pronotum paler reddish yellow. Elytra with same ground colour as pronotum, with each elytron having a large dark brown central macula, expanding laterally to interval 7, and leaving base, apex and first interval reddish yellow. Legs and palpi pale yellowish brown, antennae with first two antennomeres yellowish brown, remaining ones very weakly infuscated. Ventral surface more or less uniformly pale reddish brown, with epipleura paler.

Head ( Figs 16-18 View Figs 16-18 ) including eyes 0.81-0.86 times as wide as pronotum, with eyes moderately prominent (head 1.63-1.69 times as wide as head between eyes). Labrum with apical margin almost linear, but somewhat sloped down to the right side. Mandibles medium sized, left mandible moderately sharp at apex, not thickened or truncate. Antennae moderately long, 2.48-2.75 times as long as pronotum and 0.90-0.93 times as long as elytra. Microsculpture on clypeus almost isodiametric, on labrum almost isodiametric to weakly transverse, on frons with very fine and very lightly impressed, on vertex with moderately impressed isodiametric meshes becoming weakly transverse in front of pronotal apical margin.

Pronotum ( Figs 16-18 View Figs 16-18 ) 1.33-1.39 times as wide as long, 1.17-1.23 times as wide as head, widest in second quarter, lateral seta inserted at beginning of second quarter. Apical margin almost rectilinear or very faintly to moderately emarginated, anterior angles narrowly rounded at tips, usually weakly, in one paratype ( Fig 17 View Figs 16-18 ) markedly projecting forward. Sides convex in anterior half and rectilinearly narrowed to posterior angles, which are obtuse and more or less widely rounded. Basal margin almost rectilinear or weakly arcuate medially, and oblique to posterior angles. Lateral furrows moderately wide anteriorly, becoming widened in posterior third, where they are fused with the baso-lateral impressions. The latter medium sized and somewhat oblique, distinctly delimited from the pronotal disc and the somewhat depressed median part of base, fused with basal and lateral margins, the latter somewhat reflexed. Anterior transverse impression obsolete or suggested. Microsculpture on disc with very lightly impressed, moderately transverse meshes becoming more distinct at apical quarter, and at basolateral impressions and lateral furrows with distinct isodiametric meshes.

Elytra ( Fig. 16 View Figs 16-18 ) rather short, 1.51-1.59 times as long as wide, 2.74-2.95 times as long and 1.30-1.41 times as wide as pronotum. Sides moderately widened posteriorly, widest just posterior to the middle. Subapical sinuation very faint. Elytral striae distinctly impressed and impunctate, intervals rather flat, becoming weakly narrowed and weakly convex at apex. Microsculpture on scutellum and around basal pore isodiametric, on elytral intervals rather obsolete, only with traces of very lightly impressed transverse lines.

Metepisterna long, distinctly narrowed posteriad, at inner margin about 1.5-1.6 times longer than wide at basal margin. Prosternum medially with at least 4 and close to apical margin with a row of 6-7 medium long setae (often broken resulting in the insertion points difficult to observe). Prosternal process with 2 distinct setae.

Pro- and mesotarsomeres 2-4 of males distinctly dilated. Protarsomere 4 markedly, mesotarsomere 4 weakly bilobed. Protarsomeres 1-4 and mesotarsomeres 2-4 of males with biseriately arranged adhesive hairs on ventral surface.

Median lobe of aedeagus ( Figs 56-61 View Figs 56-61 ) moderately large, with general appearance as figured. Apical plateau moderately long, somewhat triangular, distinctly narrowed to the narrowly rounded apex (dorsal aspect), the latter with a small hook (lateral aspect). Internal structures composed of 2-3 large teeth in left apical half, 3-4 large to medium sized teeth in right apical half, and an accumulation of 2-4 smaller teeth in basal half.

Comparisons: Anthracus hornburgi nov.sp. occurs sympatrically with A. skalei JAEGER, 2015 and A. spec. 1 (near A. biplagiatus (BOHEMAN) , at Bangladesh, Chittagong, and with A. spec. 2 (near A. annamensis (BATES) at Myanmar, Pègū and Rangoon. It differs from all mentioned taxa by its characteristic median lobe of the aedeagus with apical plateau longer and somewhat triangular, and/or the presence of large subapical teeth in the internal sac (see Figs 56-61 View Figs 56-61 , 74-79 View Figs 74-79 ). From A. spec. 1 it can be also distinguished by the markedly dilated protarsomeres of males, instead of only very lightly dilated, and by the elytra, which are usually shorter relative to pronotal length (EL/PL <2.96, in A. spec. 1> 2.99), and from A. sp. 2 also by the shape of the pronotum which is usually less distinctly narrowed to pronotal base. From A. skalei it differs also by the different proportion of the pronotum (PW/PL> 1.32 in A. hornburgi , <1.33 in A. skalei ). However, there are no reliable characters to differentiate atypical females of all three taxa at this time.

A. hornburgi is also rather similar to A. siamensis nov.sp. from Thailand and Laos. It differs from this species by the external shape and internal structures of the aedeagus (see Figs 56-61 View Figs 56-61 , 63-67 View Figs 62-67 and 69-73 View Figs 68-73 ).

Distribution A. hornburgi nov.sp. is so far known from Myanmar and Bangladesh ( Fig. 81 View Fig 81 )