Antithala filipescui, Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 9-12

Antithala filipescui nov. sp.

Figs 3D 1 –D 2 , 3E 1 – E 2 , 3F 1 –F 2 , 4A .

Mitra Sturi nov. form. (Varietät)— Hoernes & Auinger 1880: 90, pl. 10, figs 28–31 only [non Antithala sturi ( Hoernes & Auinger, 1880) ].

Type material. Holotype: NHMW 1876 /0011/0012, SL: 4.3 mm, MD: 1.9 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 10, figs 31a–c), figs 3E 1 –E 2, Fig. 4A . Paratypes: NHMW 2020 /0057/0001, SL: 4.4 mm, MD: 1.9 mm. Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 10, figs 28a–c), figs 3F 1 –F 2 ; NHMW 2020 /0057/0002, SL: 4.9 mm, MD: 2.1 mm. Lăpugiu de Sus ( Romania), figs 3D 1 –D 2 .

Additional material. NHMW 2020 /0057/0003, illustrated in Hoernes & Auinger (1880, pl. 10, figs 29a–c) ; NHMW 2020 /0057/0004, illustrated in Hoernes & Auinger (1880, pl. 10, figs 30a–c); 18 specimens NHMW 2020 /0057/0005; all Lăpugiu de Sus ( Romania) .

Type locality. Lăpugiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, Badenian (Langhian).

Etymology. In honour of Sorin Filipescu (Babeş-Bolyai University, Cluj, Romania), in recognition of his contributions to the Neogene stratigraphy of the Transylvanian Basin.

Diagnosis. Antithala species of small size, with beaded sculpture, four beaded cords on spire whorls, four columellar folds, adapical much stronger than lower three folds.

Description. Shell small, broad biconical with weakly gradate spire. Protoconch (fig. 4A) high conical of 2.7 weakly convex whorls (height: 500 μm, diameter: 460 μm). Teleoconch of 4–4.5 straight sided whorls with periphery at abapical suture; suture distinctly incised. Axial ribs on first teleoconch whorl intersected by weaker spiral grooves forming four spiral rows of subquadrate nodes on later whorls. Nine to ten beaded spiral rows on last whorl accompanied by four spiral cords on fasciole. Spiral groove below subsutural row of nodes slightly more pronounced throughout. Last whorl high; maximum convexity at periphery, weakly convex above, narrowly constricted. Aperture elongate, narrow with narrow anal sinus bordered by broad, low parietal denticle. Columella indistinct, narrow, bearing prominent adapical columellar fold and three weak folds below. Outer lip thickened, with five to six small irregular denticles within, confined to adapical two thirds of aperture. Siphonal canal moderately long, straight with shallow siphonal notch.

Shell measurements and ratios. SL: 4.3–4.9 mm, MD: 1.9–2.1 mm; AA: 44–49°, SL/MD: 2.2–2.4, AL/AW: 5.1–6.7, AH/S: 2.1–2.3.

Discussion. Hoernes & Auinger (1880) discussed this species as ‘variety’ of Mitra sturi ( Hoernes & Auinger, 1880) , but it differs from that species in its stout biconical outline. Moreover, its sculpture is less regular, less tuberculose and the axial ribs are more prominent. The adapical columellar fold of Antithala filipescui nov. sp. is stronger and the abapical folds weaker compared to A. sturi . The higher and narrower spire allows a separation from Antithala claviformis .

The prominent adapical columellar fold distinguishes A. filipescui also from the otherwise similar A. neugeboreni ( Hoernes & Auinger, 1880) . In addition, the periphery of the last whorl of A. filipescui is in a more abapical position compared to A. neugeboreni .

Palaeoenvironment. Unknown; the collections from Lăpugiu de Sus indicate a mixing of coastal-inner neritic and middle neritic elements.

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 3. A1–A2. Antithala claviformis nov. sp., holotype, Coşteiu de Sus (Romania), NHMW 1869/0001/0670. B1–B2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0001. C1–C2. Antithala claviformis nov. sp., paratype, Coşteiu de Sus (Romania), NHMW 2020/0056/0002. D1–D2. Antithala filipescui nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0002. E1–E2. Antithala filipescui nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1876/0011/0012. F1–F2. Antithala filipescui nov. sp. paratype, Lăpugiu de Sus (Romania), NHMW 2020/0057/0001.

FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes & Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes & Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037.