Architipula anglicana ( Tillyard, 1933 ) — 2017

Kopeć, Katarzyna, Krzemiński, Wiesław, Skowron, Karina & Coram, Robert, 2017, The genera Architipula Handlirsch, 1906 and Grimmenia Krzemiński and Zessin, 1990 (Diptera: Limoniidae) from the Lower Jurassic of England, Palaeontologia Electronica 15 (32), pp. 1-7 : 4

Architipula anglicana ( Tillyard, 1933) — comb. nov.

Figure 3

Revised diagnosis. Wing 7.5–10.0 mm long; Sc ending opposite mid-length of R 2+3+4; R 4 about three times as long as R 2+3+4; and slightly longer than Rs; M 1 about three times longer than petiolus; upper part of d-cell approximately twice as long as lower; M 3 markedly undulate.

Redescription. Sc terminates opposite mid-length of R 2 +3+4; cross-vein sc-r situated twice its length before tip of Sc; R 1 ends opposite one third length of R 2 +3 and R 3 combined; r-r ( R 2 ) situated almost twice its length before tip of R 1 ; Rs very long, almost equal to R 4 ; R 4 3.5 times as long as R 2 +3+4 and slightly longer than Rs; d-cell large (about 0.17 the length of the wing), elongate along the upper edge (top of cross-vein m-m shifted away from wing base); M 1 about three times longer than petiolus and slightly longer than the d-cell; M 3 prominently undulate and almost 3.5 times longer than the basal part of d-cell; mc-u in holotype just before bifurcation of M 3+4 into M 3 and M 4, in the remaining specimens located at the fork M 3+4; A 2 relatively long and almost straight, terminally slightly curved to wing margin .

Material examined. Holotype: In. 11298, Alderton Hill from Brodie collection. Other material: I. 11272 also from Alderton Hill; I. 11313, I. 11365, and I. 11398 from Dumbleton, Upper Lias (Lower Jurassic: Toarcian). All specimens are housed in the Natural History Museum, London .

Remarks. Examination of the holotype In. 11298 and additional specimens of Architipula anglicana allowed us to rectify mistakes in the drawing of the holotype by Tillyard (1933, p. 76), which led to his creation of a new genus; for example, in the position of cross-vein m-cu, the addition of cross-veins in the radial sector and the length of Sc. The venation of A. anglicana is, in fact, very similar to that of A. seebachi , notably in the length of vein Sc and the construction of the d-cell, cross-vein m-m situated between M 1+2 and M 3, strongly oblique, its upper part towards the distal part of the wing; M 3 distinctly wavy. It clearly differs in the ratio of the length of the d-cell to the length of the wing, the proportion of M 1 to its petiolus and the location of cross-vein m-cu ( Figures 1-3 ). These differences justify separation at the species, but not genus level, and we have therefore synonymised Liassotipula with Architipula .

Figures

FIGURE 1. Wing venation of Architipula seebachi (Geinitz, 1884). SGWG 122/82, holotype of A. seebachiana Handlirsch, 1906.

FIGURE 2. Wing venation of Architipula seebachi (Geinitz, 1884). SGWG 119/6, holotype.

FIGURE 3. Wing venation of Architipula anglicana (Tillyard, 1933). In. 11298, holotype.