Archizelmirinae Rohdendorf, 1962

LUKASHEVICH, ELENA DMITRIEVNA, 2022, Revision of Archizelmira Rohdendorf, 1962 (Diptera, Archizelmiridae), Palaeoentomology 5 (3), pp. 254-261 : 255

publication ID

https://doi.org/ 10.11646/palaeoentomology.5.3.8

publication LSID

lsid:zoobank.org:pub:10969054-B976-4E85-929A-5DED15A29FC4

DOI

https://doi.org/10.5281/zenodo.6828750

persistent identifier

https://treatment.plazi.org/id/821F87D9-3F36-FF9C-85E3-F9FEAF8BFAC8

treatment provided by

Plazi

scientific name

Archizelmirinae Rohdendorf, 1962
status

 

Subfamily Archizelmirinae Rohdendorf, 1962

Composition. Type genus from the Jurassic and the Early Cretaceous of Asia .

Diagnosis. Flagellomeres not differentiated; cervical region not long.Wing with Sc incomplete, extremely short, extending to level of m-cu; R 1, long, extending far beyond M 1+2 furcation; Rs base sloping backwards; R 4+5 apically branched; basal part of M present as non-sclerotized fold; CuP thin, complete; anal lobe of moderate size.

Remarks. Grimaldi et al. (2003) considered the backwards slope of Rs and the very weak development of Sc as the apomorphic characters of Archizelmira . Both characters occur in the new species described below, although the slope of Rs is rather slight. The type genus Archizelmira is separated into a subfamily of its own with two putative apomorphies, viz. the extremely short Sc and the recurrent Rs base.

Some characters have been missed in previous descriptions of the genus. The incomplete CuP (A 1 of authors) has been mentioned as a familial character of Archizelmiridae ( Grimaldi et al., 2003) . Indeed, CuP is often indistinct on compressions due to its thinness, but the present study found a complete CuP in the new species ( Fig. 1 View FIGURE 1 ), as well as in specimens of the type species from both localities ( Fig. 2B, E View FIGURE 2 ) and A. baissa ( Fig. 3E, F View FIGURE 3 ). The basal part of M, present in the new species as a nonsclerotized fold, is recorded for the first time in the family thanks to the excellent preservation of the wing from Kubekovo ( Fig. 1 View FIGURE 1 ). However, such fold is also distinct on the well-preserved wings of A. kazachstanica from Shar Teg ( Fig. 2D, E View FIGURE 2 ). It is visible as well on the photographs accompanying the original description, where it has not been discussed ( Grimaldi et al., 2003: figs 1.2–4). The anal lobe of the wing of Archizelmira is reduced or moderately developed ( Fig. 4 View FIGURE 4 ), whereas it is very strongly developed in the three other genera (‘anal lobe of wing 90 degrees or more’ in Hippa & Vilkamaa, 2005).

Thus the incomplete CuP is not a character of Archizelmiridae but another putative synapomorphy of the Cretaceous clade ( Zelmiarcha + ( Burmazelmira + Archimelzira )). It supplements the diagnostic characters of the Burmazelmirinae subfam. nov. (see below), which also include the reduction of the basal part of M, the strongly pronounced anal lobe, and the characters pointed out by Grimaldi et al. (2003), which were listed above.

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