Asperuginoideae Al-Shehbaz, Hendriks, M.Koch, F.Lens, Lysak, C.D.Bailey, Mumm. & D.A.German, trib. nov. (1: 1), 1982

German, Dmitry A., Hendriks, Kasper P., Koch, Marcus A., Lens, Frederic, Lysak, Martin A., Bailey, C. Donovan, Mummenhoff, Klaus & Al-Shehbaz, Ihsan A., 2023, An updated classification of the Brassicaceae (Cruciferae), PhytoKeys 220, pp. 127-144 : 127

Asperuginoideae Al-Shehbaz, Hendriks, M.Koch, F.Lens, Lysak, C.D.Bailey, Mumm. & D.A.German, trib. nov. (1: 1)

Type.

Asperuginoides Rauschert

Description.

Herbs annual. Trichomes stalked, stellate or substellate, 4-6-rayed, these mixed with glochidate ones on fruit. Multicellular glands absent. Cauline leaves petiolate, not auriculate. Racemes bracteate throughout, usually elongated in fruit, with strongly recurved fruiting pedicels. Flowers actinomorphic; sepals ascending, base of lateral pair not saccate; petals white, claw undifferentiated from blade; filaments slender at base, unappendaged; pollen 3-colpate; ovules 2 per ovary, apical. Fruits dehiscent silicles, suborbicular, latiseptate, unsegmented, wingless, with long-stalked, setose, stiff trichomes glochidiate at apex; septum complete or absent; style distinct; stigma entire. Seeds aseriate, broadly winged; cotyledons accumbent. x = 16.

Distribution.

Afghanistan, Armenia, Iran, Kazakhstan, Kyrgyzstan, Pakistan, Tajikistan, Turkey, Turkmenistan, Uzbekistan.

3c. Chamira

Although the tribe Chamireae was first recognized by Sonder (1846) and later accepted by Schulz (1936), it has not been widely recognized since, and Chamira Thunb. was listed as unplaced in Al-Shehbaz (2012). The findings of Hendriks et al. (2022) agree with those of Mummenhoff et al. (2005), Mandáková et al. (2012), Nikolov et al. (2019), Walden et al. (2020), and Dogan et al. (2021) that Chamira and Heliophila are closely related genera that do not belong to the same tribe, and the former has been used as the outgroup for phylogenetic and genomic studies of the latter. A tribal description comparable to that of other tribes is provided below.