Asytesta alexandriae Setliff, 2012

Setliff, Gregory P., 2012, 3462, Zootaxa 3462, pp. 1-125 : 38-53

Asytesta compressipes species group

Asytesta alexandriae Setliff , new species

( Figures 131–132, 191)

Diagnosis. The mirrored L-shaped maculae on either side of the dorsomedian vitta on the pronotal disk will distinguish this species from its congeners. This species may be confused with A. woodlarkiana new species, which has a broken transverse line at the middle of the pronotal disk that transects the dorsomedian vitta, but A. woodlarkiana also has large irregularly distributed white maculae covering its dorsum and has a darker background color.

Description. Measurements (n = 4): body length 7.1–7.8 mm (mean 7.5 mm), body width 3.2–3.5 mm (mean 3.3 mm), pronotal length 2.8–3.0 mm (mean 3.0 mm), elytral length 4.3–4.8 mm (mean 4.5 mm), rostral length 2.1–3.0 mm (mean 3.0 mm). Body elongate-oval, 2.1–2.3 X longer than broad; integument dark reddish-brown, antennae and tarsi light reddish-brown. Sparsely covered with small, light tan to dark brown scales, with more densely distributed white scales in vittae and maculae. Vertex of head, pleura, legs, and venter more densely clothed in white to tan scales, white scales denser on tibia; vestiture on legs interspersed with dense, white, hairlike setae; male with suberect, hair-like setae along ventral margin of protibia, setae longer than elsewhere on tibia; tarsi clothed with sparse, white, hair-like setae. Pronotum uniformly dark; disk with dorsolongitudinal vitta from apex to base, 2 pairs of irregular maculae on either side of medial vitta, anterior pair at cervical constriction smaller, sublateral, posterior pair behind middle larger, transverse, anterior and posterior maculae joined by faint scales, forming L-shaped marks; anterolateral region of pronotum with distinct, well defined ring of white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring extending slightly beyond middle of procoxa. Elytra with patchy white scales evenly distributed on disk, entire anterior margin with fascia of white scales, reaching epipleural margin; epipleural margin lined with dense white scales from base to slightly past middle; intervals 5 with broken, longitudinal vittae on declivity. Mesosternum and metasternum covered with dense white scales except transverse glabrous line at suture between sclerites, ventrites without scales except for sparse, evenly distributed, hair-like setae on ventrite 1 and 5 and lateral margins of 2–4, glabrous in middle. Scales fine (indistinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum slightly shorter than or subequal to pronotal length, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1–1.2 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, diameter subequal to punctures at base of elytra. Elytra 1.3–1.4 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs moderately long, hind femora exceeding elytral apices by 1/3 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, obtuse, tooth-like in male, broader, gradually tapering to apex in female (as in Figures 48–49); dorsal margin strongly convex (as in Figures 51–52). Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct suprauncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 7–8 minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct under high magnification.

Female terminalia. As in generic description.

Material examined. Holotype ♂, " New Guinea: NE, Feramin; 150– 120 m, May 23–31, 1959 / W. W. Brandt, Collector, Bishop" ( BPBM) . Paratypes: 1 ♂, 1 ♀, " New Guinea: NE, Feramin; 150– 120m, June 15–18, 1959 / W. W. Brandt, Collector, Bishop" ( BPBM) ; 1 ♀, " New Guinea: NE, Feramin; 150– 120m, June 7–14, 1959 / W. W. Brandt, Collector, Bishop" ( BPBM) .

Distribution. Papua New Guinea: Sandaun Province: Feramin.

Etymology. This species is named for my daughter, Alexandria, who was born during the course of this study.

Asytesta arachnopus Heller

( Figures 56–57, 137–138, 191)

Asytesta arachnopus Heller, 1895: 16 . Lectotype, here designated, in SMTD. Labeled “New Guinea, 8926/ Coll. J. Faust, Aukauf 1900/ Type/ Staatl., Museum fur, Tierkunde Dresden.” Type locality: Papua New Guinea: Sattelberg.

Diagnosis. Asytesta arachnopus is most closely allied with A. gestroi Heller in vestiture, markings (viz., a pair of diverging sublateral vittae that do not reach the posterior margin of the pronotum), size, and leg morphology. In A. arachnopus , however, the vittae on the fifth elytral intervals do not reach the humeri and in A. gestroi the humeri are covered by large, irregular, humeral maculae spanning intervals 3–7.

Redescription. Measurements (n =14): body length 6.6–9.1 mm (mean 7.8 mm), body width 3.1–4.0 mm (mean 3.6 mm), pronotal length 2.6–3.6 mm (mean 3.1 mm), elytral length 4.0– 5.5 mm (mean 4.7 mm), rostral length 2.0–3.0 mm (mean 2.5 mm). Body elongate-oval, 2.0–2.4 X longer than broad; integument dark brown to black, shiny, antennae and tarsi dark reddish-brown. Clothed in sparsely distributed, dark brown scales; integument not obscured by squamae except by more densely distributed white scales in vittae, in sharp contrast to dark background squamae. Pleura, legs, and venter more densely clothed in white to tan scales, white scales denser on tibia; rim of mesosternal receptacle sparsely covered with fine white scales; vestiture on legs interspersed with white, hair-like setae; dorsal edge of all tibiae lined with white scales; tarsi clothed with dense, white, hair-like setae. Pronotal disk with narrow dorsolongitudinal vitta from behind cervical constriction to base; pair of sublateral longitudinal vittae from anterior margin to just before base, slightly divergent past cervical constriction; sometimes with short, faint, longitudinal vittae on sides from anterior margin to midline, absent in most specimens; anterolateral region of pronotum with distinct, well defined ring of white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytra with broad sutural stripe of white scales from just posteriad of midline to apex; pair of similarly colored longitudinal vittae on fifth intervals from humeri or slightly behind humeri to subapex; epipleural margin with white scales from slightly behind base to apex. Mesosternum and metasternum clothed in sparse white to tan scales, ventrites mostly glabrous, with sparse lightbrown squamae on ventrite 1 and 5, ventrites 2–4 more or less glabrous. All scales very fine to minute (indistinct under low magnification), appressed; white setae distinct, suberect to decumbent.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.0–1.3 X broader than long, broadest slightly before base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.2–1.5 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by about 1/2 their length; mesofemora exceeding elytral apices. Profemora bidentate or unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange reduced in males, present, gradually tapering to apex in female; dorsal margin strongly convex ( Figures 56–57). Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with reduced supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin strongly emarginate at middle, wing binding patch distinct, with two rows of 6–7 minute plectral tubercles reaching posterior margin, tubercles distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 6–8 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible only under high magnification. Tergite VIII slightly longer and apex narrower than in A. humeralis (see Figure 89). Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no internal sclerites visible.

Material examined. Lectotype (here designated): ♂, "New Guinea, 8926/ Coll. J. Faust, Aukauf 1900/ Type/ Staatl., Museum fur, Tierkunde Dresden" ( SMTD) . Paralectotypes (here designated): 1 ♀, "Typus/ New Guinea 8926/ Staatl., Museum fur, Tierkunde Dresden" ( SMTD) ; 2 ♂♂, 2 ♀♀, "New Guinea, 8926/ Paratype / Staatl., Museum fur, Tierkunde Dresden" ( SMTD) ; 1 ♀, " K. Wilhelmsland / Cotypus/ Asytesta arachnopus m., Determined K. M. Heller " ( SMTD) ; Non-type material: 1 ♀, " Asytesta arachnopus m., Det. K. M. Heller 1911/ D.N. Guinea / Coll. Bennigsen / Coll. DEI Müncheberg " ( DEI) ; 1 ♂, " Asytesta arachnopus m., Det. K. M. Heller / Coll. Kraatz / Coll. DEI. Mucheberg " ( DEI) ; 1 ♂, " Asytesta arachnopus m., Det. K. M. Heller 08/ Simbang, D. N. Guinea / Coll. v. Bennigsen / Coll. DEI. Mucheberg " ( DEI) ; 1 ♂, " D. Neu-Guinea., Wahnes, Franklin Müller/ Asytesta arachnopus m 1895, K. M. Heller Det. 1920/ Coll. DEI. Mucheberg " ( DEI) ; 1 ♀, "D. N. Guinea / Coll.v. Bennigsen / Coll. DEI. Mucheberg " ( DEI) .

Distribution. Papua New Guinea: Morobe Province: Huon Peninsula: Sattelberg [type locality], Simbang; Madang Province: Madang.

Remarks. Two additional cotypes with identical labels as the lectotype were also seen by me at MNHN but were not borrowed for examination. These specimens are also considered paralectotypes. I have selected a male specimen from Faust’s collection in SMTD as the lectotype to fix the name to that specimen. It is one of two specimens that are labeled “type”. All other paralectotypes originally were labeled with either cotype or paratype labels.

Asytesta cheesmanae Setliff , new species

( Figures 149–150, 191)

Diagnosis. This handsome species includes some of the largest specimens (11.5 mm) recorded for the genus approaching and exceeding the usually much larger body sizes of the other crowned weevil genera. It has an elongate body, with dark background vestiture and distinct vittae on the dorsum. It is closely related to A. compressipes and A. eudyasmoides ; however, neither of these species, nor any other Asytesta species except A. gestroi have the large oblong humeral maculae of A. cheesmanae . Asytesta gestroi differs in having white sublateral vittae on the pronotum whereas A. cheesmanae has burnt-orange lateral vittae on the pronotum.

Description. Measurements (n =9): body length 6.8–11.5 mm (mean 7.9 mm), body width 3.0– 4.8 mm (mean 3.4 mm), pronotal length 2.8–4.8 mm (mean 3.2 mm), elytral length 4.0– 6.7 mm (mean 4.8 mm), rostral length 2.0– 4.4 mm (mean 2.3 mm). Body elongate-oval, 2.3–2.4 X longer than broad; integument dark brown to black, antennae and tarsi dark reddish-brown. Sparsely covered with small, light tan to dark brown scales, with more densely distributed white to burnt-orange scales in vittae and maculae. Vertex of head, pleura, legs, and venter more densely clothed in white to tan scales, white scales denser on tibia; rim of mesosternal receptacle sparsely covered with fine white scales; vestiture on legs interspersed with white, hair-like setae; dorsal edge of all tibiae lined with yellowish-white scales; tarsi clothed with sparse, white, hair-like setae. Pronotal disk with narrow dorsolongitudinal vitta from apex to base, pair of longitudinal vittae on sides, anterolateral region of pronotum with distinct, well defined ring of white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring extending beyond middle of procoxa to basal pronotal margin. Elytra with pair of large, white, oblong maculae between intervals 3–6 in basal 1/4 of elytra; broad sutural stripe of tan to orange scales from just posteriad of scutellum to apex and pair of similarly colored longitudinal vittae on fifth intervals, joining with white humeral maculae; epipleural margin with dense tan to orange scales from base to slightly past middle. Mesosternum and metasternum covered with sparse white scales, ventrites clothed in sparse brown squamae on ventrite 1 and 5, ventrites 2–4 more or less glabrous. Scales very fine (indistinct under low magnification), appressed.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum slightly shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.0–1.1 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one yellowish-white to brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.3–1.5 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by nearly 1/2 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange almost to completely obsolete in males, present, gradually tapering to apex in female; dorsal margin strongly convex (as in Figures 51–52). Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with reduced supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin strongly emarginate at middle, wing binding patch distinct, with two rows of 6–8 minute plectral tubercles reaching posterior margin, tubercles distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 8–11 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible only under high magnification. Tergite VIII slightly longer and apex narrower than in A. humeralis (see Figure 89). Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no internal sclerites visible.

Material examined. Holotype: ♂, " Dutch New Guinea:, Japen I., R. Manai – Undei, 500ft. x. 1938, L. E. Cheesman, B.M.1938-593" ( BMNH) . Paratypes: 1 ♂, label data same as holotype ( GPSC) ; 1 ♂, label data same as holotype [specimen disarticulated] ( UMSP) ; 4 ♂♂, 2 ♀♀, " Dutch New Guinea:, Japen I., Mt. Baduri, 1000ft. viii. 1938, L. E. Cheesman, B.M.1938-593" ( BMNH) .

Distribution. Indonesia: Yapen Island.

Etymology. This species is named in honor of L. E. Cheesman, intrepid explorer in the Papuan region and collector of the type series.

Asytesta compressipes (Chevrolat)

( Figures 51–52, 68, 151–152, 191)

Arachnopus compressipes Chevrolat, 1877: 189 . Transferred to Asytesta by Heller, 1908: 23, see also Heller, 1933: 19. Holotype not examined.

Cyamobolus trivittatus Pascoe, 1885: 274 , synonymy by Heller, 1908: 23. Holotype in BMNH, labeled “N.G, Andai/ Cyamobolus trivittatus , det. Pasc/ Type, H.T.” Type locality: Andai.

Asytesta trivittata (Pascoe) ; transferred to Asytesta by Heller, 1895: 15.

Asytesta trivirgata Pascoe , error in Heller, 1933: 19.

Diagnosis. This is a large species ranging up to 10.5 mm in length with an elongate body and immaculate, brown background vestiture. The only marking on the pronotum is a white dorsomedian longitudinal vitta. The elytra have similarly colored, complete, longitudinal vittae on the fifth intervals that reach the apices. This species is closely related to A. cheesmanae new species and A. eudyasmoides . However, A. cheesmanae has large oblong humeral maculae that interrupt the vittae on the fifth intervals and (usually) sublateral vittae on the pronotal disk. Asytesta eudyasmoides lacks vittae on the pronotum and elytra and instead has small humeral rectangular maculae.

Redescription. (Based on holotype and paratypes of Cyamobolus trivittatus ). Habitus ( Figures 151–152). Measurements (n = 10): body length 7.3–10.5 mm (mean 9.0 mm), body width 3.0– 4.4 mm (mean 3.8 mm), pronotal length 2.8–4.3 mm (mean 3.5 mm), elytral length 4.5–6.2 mm (mean 5.5mm), rostral length 2.7–3.6 mm (mean 3.2). Body elongate-oval, 2.2–2.5 X longer than broad; integument dark reddish-brown, antennae and tarsi light reddish-brown. Densely covered with small, fuscous scales, becoming brown on elytral disk; white dorsomedian vittae on pronotal disk and vittae on elytral intervals 5 from humeri to apex in sharp contrast to background squamae. Frons, base of rostrum, pleura, meso- and metasternites clothed with dingy yellowish-white to tan scales; coxae and dorsal portion of middle and hind femora with yellowish-white to tan scales interspersed with tan, hair-like setae, dorsal edge of all tibiae lined with yellowish-white scales; tarsi covered with sparse, white, hair-like setae, rim of mesosternal receptacle sparsely covered with fine tan scales. Scales very fine (indistinct at low magnification), appressed.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes ( Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, median longitudinal carinae absent. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum subequal in length to pronotum or slightly shorter than pronotum, parallel sided in basal 2/3, slightly broadened in apical 1/3, with small median interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.0–1.2 X broader than long, broadest at base, subtruncate, median area weakly produced. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/ 2 diameter of puncture, each bearing one yellowish-white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.3–1.5 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity ( Figure 68); fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by nearly 1/2 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange obsolete in males, present, gradually tapering to apex in female ( Figures 51–52); dorsal margin strongly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with reduced supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 8 minute plectral tubercles reaching posterior margin, tubercles distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 7–9 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible only under high magnification. Tergite VIII slightly longer and apex narrower than in A. humeralis (see Figure 89). Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no internal sclerites visible.

Material examined. Holotype for Cyamobolus trivittata Pascoe : ♂, "N.G, Andai/ Cyamobolus trivittatus , det. Pasc / Type, H.T" ( BMNH) . Paratype for C. trivittatus : 1 ♂, "Nuova Guinea, Andai, Ag. 79, L. M. D. Albertis / typus/ trivitattus, Pascoe/ Cyamobolus trivittatus typus! Pascoe" ( MSNG) . Non-type material: 2 ♂♂, "N. Guinea, Andai XII, Coll. Bruijn 1875" ( MSNG) ; 2 ♂♂, "Nuova Guinea, Andai, Ag. 79, L. M. D. Albertis " ( MSNG) ; 2 ♂♂, "N. Guinea, Andai XII, Coll. Bruijn 1875" ( MSNG) ; 1 ♂, "Andai, N. Guinea, Doherty/ Asytesta trivittata, Det. K. M. Heller 1912 " ( MSNG) ; 1 ♂, "N. Guinea, Coll. Heller / Asytesta trivittata, Det. K. M. Heller 1912 " ( MSNG) ; 1 ♀, "[illegible]/ Coll. J. Faust, Ankauf 1900" ( MSNG) ; 1 ♂, "[illegible]/ Coll. J. Faust, Ankauf 1900" ( SMTD) ; 2 ♀♀, "Museum Paris, Nouv. Guinee, Baie de Geelvink , Rafray & Maindron, 1878" ( MNHN) ; 1 ♀, "N. Guinea, Andai VI, Beccari 1875" ( SMTD) ; 1 ♀, "N. Guinea, Badui / [illegible]/ Coll. J. Faust, Ankauf 1900" ( SMTD) ; 1 ♂, " Neu-Guinea , [illegible], ex coll. Fruhstorfer / Coll. Kraatz / Asytesta trivittata, det. K. M. Heller " ( DEI) ; 2 ♂♂, "Nov. Guinee / Heller/ Asytesta trivittata, det. K. M. Heller 1922 " ( USNM) ; 1 ♀, "Nov. Guinee / Heller/ Asytesta trivittata, det. K. M. Heller 1922 / Collection Bovie, thru, Buchannan" ( USNM) ; 1 ♂, "Wasior DMP logging, camp, Km 27, Wombu, S 03˚01' E 134˚28', 200 m, 10.I.2001 / Irian Jaya , Manokwari Prov., Wandammen Bay, leg. A. Riedel " ( ARC) ; 1 ♂, " Salawatti Isl. , Solol, 0–350m, 6-7.XI./ Irian Jaya, Sorong Prov., leg. A. Riedel " ( ARC) .

Distribution. West Papua: Manokwari Province: Andai [type locality], Wandammen Bay; Cenderawasih Bay (= Geelvink Bay); Sorong Province: Salawatti Island.

Remarks. The following notes refer to “ Cyamobolus trivittata ” synonymized above. Pascoe (1885: 274) did not mention the number of specimens in the type series. Several other specimens with identical data labels as the paratype are in the MSNG collection and may also be paratypes. One additional specimen from Andai was also seen by me in the BMNH collection but was not borrowed for examination.

Asytesta concolora Setliff , new species

( Figures 153–154, 191)

Diagnosis. This species is closely related to A. cheesmanae new species, A. compressipes , and A. eudyasmoides in its elongate body and dark immaculate background vestiture. Unlike these three species, A. concolora lacks any indication of white maculae or vittae anywhere on the body.

Description. Measurements (n = 3): body length 9.1–9.6 mm (mean 9.3 mm), body width 3.5–4.0 mm (mean 3.9 mm), pronotal length 3.7–3.8 mm (mean 3.8 mm), elytral length 5.3–5.8 mm (mean 5.6 mm), rostral length 3.0– 3.2 mm (mean 3.1 mm). Body elongate-oval, 2.3–2.6 X longer than broad; integument black, antennae and tarsi dark reddish-brown. Densely covered with small, fuscous scales, becoming brown on elytral disk; body lacking white vittae and maculae. Vertex of head, pleura, legs and venter similarly clothed with dense appressed fuscous, except frons and base of rostrum with pale scales; coxae and dorsal portion of middle and hind femora with white scales interspersed with white, hair-like setae, dorsal edge of all tibiae lined with bright white scales; tarsi covered with sparse, white, hair-like setae, rim of mesosternal receptacle covered with fine white scales. Scales very fine (indistinct under low magnification), appressed.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes weakly convex, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum slightly shorter than pronotal length, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum slightly longer than broad to 1.1 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, diameter subequal to punctures at base of elytra. Elytra 1.4–1.5 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by nearly 1/2 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange obsolete in males, present, gradually tapering to apex in female (as in Figures 51–52); dorsal margin strongly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with reduced supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Material examined. Holotype: ♂, " Wandammen Bay, Wondiwoi Mts., Wasior , 250–600m, 4.I.2001 / Irian Jaya, Manokwari Prov. , leg. A. Riedel " ( MZB) . Paratypes: 1 ♂, 1 ♀, label data same as holotype ( ARC) .

Distribution. West Papua: Manokwari Province: Wasior.

Etymology. The specific epithet refers to the uniformly dark body without white maculae or vittae on the pronotum and elytra in this species that occur in all of its close congeners.

Asytesta eudyasmoides Heller

( Figures 155–156, 191)

Asytesta eudyasmoides Heller, 1914a: 655 . Lectotype, here designated, in SMTD. Labeled “ Z. Neiuw Guinea, Lorantz 1909- 10, Bivak eiland, I.10/ Typus!, A.eudyasmoides .” Type locality: Biak Island .

Diagnosis. This species is closely related to A. cheesmanae new species, A. compressipes , and A. concolora new species based upon its elongate body form and dark immaculate background vestiture. Unlike these species, A. eudyasmoides has small, white, rectangular maculae on the elytral humeri. This species also superficially resembles A. humeralis due to its dark background squamae and white humeral maculae; however, A. eudyasmoides is more elongate narrow and has unidentate profemora (bidentate in A. humeralis ).

Redescription. Measurements (n = 2): body length 8.1–8.7 mm (mean 8.4 mm), body width 3.8–3.9 mm (mean 3.9 mm), pronotal length 3.1–3.5 mm (mean 3.3 mm), elytral length 5.0– 5.2 mm (mean 5.1 mm), rostral length 3 mm. Body elongate-oval, 2.1–2.2 X longer than broad; integument dark reddish-brown, antennae and tarsi light reddish-brown. Densely covered with small, fuscous scales, becoming brown on elytral disk; humeri with narrow rectangular maculae of yellowish-white scales on intervals 5–7, maculae extends posteriad a short distance in some specimens on fifth intervals, thin line of yellowish-white scales on sides connecting humeral maculae to epipleural margin, in sharp contrast to dark background squamae. Head, including base of rostrum, pleura, meso- and metasternites clothed with dingy yellowish-white to tan scales; coxae and dorsal portion of middle and hind femora with yellowish-white to tan scales interspersed with white, hair-like setae, dorsal edge of all tibiae lined with yellowish-white scales; tarsi covered with sparse, white, hair-like setae, rim of mesosternal receptacle covered with fine white scales. Scales very fine (indistinct under low magnification), appressed.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum subequal in length to pronotum or slightly shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Clypeus emarginate. Antennal club oval.

Thorax. Pronotum 1.1–1.2 X broader than long, broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by nearly 1/2 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange obsolete in males (as in Figure 51–52); dorsal margin strongly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with reduced supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 8 minute plectral tubercles reaching posterior margin, tubercles distinct under high magnification.

Female terminalia. Female not known.

Material examined. Lectotype (here designated): ♂, " Z. Neiuw Guinea, Lorantz 1909-10, Bivak eiland, I.10/ Typus!, A.eudyasmoides " ( SMTD) . Non-type material: 1 ♂, " Z. Neiuw Guinea, Lorantz 1909-10, Bivak, XI.09.750 m" ( ZMAN) .

Distribution. Indonseia: Biak Island.

Remarks. A lectotype is selected from Heller’s collection to fix the name to a single specimen to ensure stability of nomenclature.

Asytesta fayae Setliff , new species

( Figures 157–158, 192)

Diagnosis. This species is most similar to A. versuta but is differentiated by the patch of white scales on the sides of the pronotum from the anterior margin to the base that is broader near the head and rapidly convergent towards the pronotal base where it meets ninth intervals on the elytra. Each elytron bears a broad white vittae from the base to near the apex. Additionally, the male holotype lacks any indication of a tooth-like flange on the ventral margin of the protibia. These flanges are always present in A. versuta and its allies.

Description. Measurements (n = 1): body length 7.1 mm, body width 3.1 mm, pronotal length 3.0 mm, elytral length 4.1 mm, rostral length 2.5 mm. Body elongate-oval, 2.3 X longer than broad; integument reddish-brown to dark brown, antennae and tarsi light reddish-brown. Clothed in densely distributed, patchy, tan to dark brown scales; integument not obscured by squamae except in patches on elytra and pronotum and by more densely distributed yellowish-white scales in vittae, in sharp contrast to dark background squamae. Vertex of head, frons, base of rostrum, legs, and pleura more densely clothed in cream to brown scales; rim of mesosternal receptacle lined with fine yellowish-white scales; vestiture on legs evenly interspersed with white, hair-like setae, slightly denser along dorsal edge; male with suberect, hair-like setae along ventral margin of protibia, setae longer than elsewhere on tibia; tarsi clothed with dense, white, hair-like setae. Pronotal disk with dorsolongitudinal vitta from apex to base; sides with large irregular patch of white scales from anterior margin to base, broader at apex, convergent towards base, meeting intervals 9 of elytra; anterolateral region of pronotum with distinct, well defined ring of white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring extending beyond middle of procoxa to basal pronotal margin. Elytra with faint sutural stripe of yellowish-white scales from midline to apex; pair of white, longitudinal vittae on intervals 5 and 9 originating at humeri, two vittae joining at subapex, vittae not broadened apically, restricted to intervals; epipleural margin with white scales from base to slightly before apex; humeral and epipleural vittae connected by short fasciae along elytral base, fascia not reaching dorsum. Mesosternum and metasternum clothed in tan to brown scales, ventrites mostly without squamae, sparsely distributed light-brown squamae on ventrite 1 and 5, ventrites 2–4 more or less glabrous. Scales fine to (indistinct under low magnification), appressed; white setae distinct, suberect to decumbent.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 104), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Clypeus truncate. Antennal club oval.

Thorax. pronotum as broad as long, broadest slightly before base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one yellowish-white to brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by about 1/2 their length; mesofemora exceeding elytral apices. Protibia strongly compressed laterally, ventral margin with flange obsolete in male (as in Figure 51); dorsal margin moderately convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. As in generic description.

Female terminalia. Female is not known.

Material examined. Holotype: ♂, " H.C. Siebers, Kei Eil. 1922, Gn. Daab IV, ± 300 m / Asytesta sp. " ( DEI).

Distribution. Indonesia: Maluku province. Kei Islands.

Remarks. The unique male specimen lacks any indication of a protibial flange. Females may possess protibial flanges similar to A. compressipes , where flanges are present in females but not males.

Etymology. This epithet is dedicated to the memory of Faye R. Setliff.

Asytesta gestroi Heller

( Figures 161–162, 191)

Asytesta gestroi Heller, 1895: 15 . Lectotype, here designated, in SMTD. Labeled “ K. Wilhelmland, 8576/ Type.” Type locality: Papua New Guinea: Sattelberg.

Diagnosis. Asytesta gestroi is most closely allied with A. arachnopus Heller in vestiture, markings, size, and leg morphology. Asytesta arachnopus , however, differs from this species in lacking large, irregular, humeral maculae spanning intervals 3–7 and the dorsolongitudinal vitta on the pronotum of A. arachnopus is nearly obsolete. Asytesta cheesmanae new species also has large oblong humeral maculae but these are not connected to the vittae on intervals 5 as in A. gestroi . This species further differs from A. gestroi by the lateral pronotal vittae and elytral vittae comprised primarily of burnt orange scales. In contrast, A. gestroi has bright white sublateral vittae on the pronotum.

Redescription. Measurements (n =6): body length 7.0– 8.5 mm (mean 7.8 mm), body width 3.1–3.9 mm (mean 3.5 mm), pronotal length 2.9–3.5 mm (mean 3.1 mm), elytral length 4.1–5.0 mm (mean 4.7 mm), rostral length 2.5–2.8 mm (mean 2.6 mm). Body elongate-oval, 2.1–2.4 X longer than broad; integument dark brown to black, shiny, antennae and tarsi dark reddish-brown. Clothed in sparsely distributed, dark brown scales; integument not obscured by squamae except by more densely distributed white scales in vittae, in sharp contrast to dark background squamae. Frons, base of rostrum, pleura, scutellum, and legs more densely clothed in white scales, white scales denser on tibia; rim of mesosternal receptacle with fine white scales on inner margin; vestiture on legs interspersed with white, hair-like setae; dorsal edge of all tibiae lined with white scales; tarsi clothed with dense, white, hair-like setae. Pronotal disk with faint, very narrow dorsolongitudinal vitta from behind cervical constriction to base, not more than 2–3 scales wide; pair of broader sublateral longitudinal vittae from anterior margin to just before base, slightly divergent past cervical constriction; sometimes with short, faint, lateral longitudinal vittae from anterior margin to midline, absent in most specimens; anterolateral region of pronotum with distinct, well defined, oblique, vittae from lower margin of postocular lobe to just above middle of procoxae ( Figure 161); dorsal portion of ring not extending beyond middle of procoxa. Elytra with broad sutural stripe of white scales from midline to apex; pair of similarly colored longitudinal vittae on intervals 5 from humeri or slightly behind humeri to subapex; humeri with large patch of white scales covering space between intervals 3–7; epipleural margin with dense white scales from base to slightly before apex. Mesosternum and metasternum clothed in sparse white to tan scales, ventrites mostly without squamae, sparsely distributed light-brown squamae on ventrite 1 and 5, ventrites 2–4 more or less glabrous. Scales very fine to minute (indistinct under low magnification), appressed; white setae distinct, suberect to decumbent.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.0–1.2 X broader than long, broadest slightly base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.3–1.4 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by about 1/2 their length; mesofemora exceeding elytral apices. Profemora bidentate or unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange reduced in males, present, gradually tapering to apex in female; dorsal margin strongly convex (as in Figures 56–57). Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with reduced supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male and female terminalia. As in generic description.

Material examined. Lectotype (here designated): 1 ♀, " K. Wilhelmland, 8576/ Type" ( SMTD) . Paralectotypes (here designated): 2 ♂♂, "K. Wilhelmland , 8576/ Paratype " ( SMTD) ; 1 ♂, " K. Wilhelmland, 8576/ Cotypus/ Gestroi, Hell. / D. Heller, 1895 " ( MSNG) ; 1 ♀, "K. Wilhelmland, 8576/ gestroi, Heller / Coll. J. Faust, Ankauf 1900/ typus" ( SMTD) ; 1 ♀, "K. Wilhelmland, 8576/ Paratype " ( SMTD) . Non-type material: 1 ♂, 1 ♀, " D. Neu-Guinea, Wahnes Franklin-Müller/ Heller det." ( DEI) .

Distribution. Papua New Guinea: Morobe Province: Sattleberg.

Remarks. Two additional cotypes in the MNHN with the same label data as the lectotype were also located but were not borrowed for examination. These specimens are to be considered paralectotypes. The lectotype serves to fix the species name to the specimen identified above to ensure the stability of nomenclature.

Asytesta sejuncta Faust

( Figures 175–176, 192)

Asytesta sejuncta Faust, 1898: 164 [key], 167 [description]. Lectotype, here designated, in SMTD. Labeled “N. Guinea S.E., Moroka, 1300m., Loria VII-IX 93/ sejuncta, Faust / Coll. J. Faust Ankauf 1900/ Type.” Type locality: Moroka.

Diagnosis. The arrangement of whitish vittae and macula will serve to distinguish this species from its congeners. The pronotal disk in this species has a dorsolongitudinal vitta and a faint pair of incomplete longitudinal vittae on the sides that do not reach the base. The elytra have a faint sutural stripe from slightly behind the midline to the apex and a pair of white, longitudinal vittae on the fifth intervals. It is most closely related to A. versuta and A. fayae new species. Asytesta versuta differs in the immaculate background vestiture and sharply contrasting and unbroken dorsal and lateral vittae. Asytesta fayae differs by the broad irregular patch of white scales on the sides of the pronotum originating on the anterior margin and reaching the base and is broader near the head and convergent towards the pronotal base, meeting intervals 9 of elytra. Asytesta fayae also bears a broad white vitta from the elytral base to near the apex and males apparently lack a tooth-like flange on the ventral margin of the protibia.

Redescription. Measurements (n =2): body length 5.7–5.8 mm (mean 5.8 mm), body width 2.8–3.0 mm (mean 2.9 mm), pronotal length 2.5 mm, elytral length 3.2–3.3 mm (mean 3.3 mm), rostral length 2.2–2.3 mm (mean 2.3 mm).Body elongate-oval, 2.0 X longer than broad; integument reddish-brown to dark brown, antennae and tarsi light reddish-brown. Clothed in densely distributed, patchy, tan to dark brown scales; integument not obscured by squamae except in patches on elytra and pronotum and by more densely distributed yellowish-white scales in vittae, in sharp contrast to dark background squamae. Vertex of head, frons, base of rostrum, legs, and pleura more densely clothed in cream to tan scales; rim of mesosternal receptacle lined with fine yellowish-white scales; vestiture on legs evenly interspersed with white, hair-like setae, slightly denser along dorsal edge; male with suberect, hair-like setae along ventral margin of protibia, setae longer than elsewhere on tibia; tarsi clothed with dense, white, hair-like setae. Pronotal disk with dorsolongitudinal vitta from apex to base; pair of incomplete, lateral, longitudinal vittae from near anterior margin terminating before base, with small, irregular patches or maculae of cream to tan scales dispersed on sublateral and lateral margins; anterolateral region of pronotum with distinct, well defined ring of white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring extending beyond middle of procoxa to basal pronotal margin. Elytra with faint sutural stripe of yellowish-white scales from midline to apex; pair of white, longitudinal vittae on intervals 5 from humeri to subapex, vittae not broadened apically, restricted to intervals 5; epipleural margin with white scales from base to apex; humeral and epipleural vittae connected by short fasciae along elytral base, fascia not reaching dorsum. Mesosternum and metasternum clothed in white to tan scales, ventrites mostly without squamae, sparsely distributed light-brown squamae on ventrite 1 and 5, ventrites 2–4 more or less glabrous. Scales fine (indistinct under low magnification), appressed; white setae distinct, suberect to decumbent.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1 X broader than long, broadest slightly before base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one yellowish-white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.1 –1.2 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by about 1/2 their length; mesofemora exceeding elytral apices. Protibia strongly compressed laterally, ventral margin with flange distinct, obtuse, tooth-like in male, broader, gradually tapering to apex in female (as in Figures 48–49); dorsal margin strongly convex (as in Figure 51–52). Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin strongly emarginate at middle, wing binding patch distinct, with two rows of 6–7 minute plectral tubercles reaching posterior margin, tubercles distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 7–8 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible only under high magnification. Tergite VIII slightly longer and apex narrower than in A. humeralis (see Figure 89). Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no internal sclerites visible.

Material examined. Lectotype (here designated): ♂, "N. Guinea S.E., Moroka , 1300m., Loria VII-IX 93/ sejuncta, Faust / Coll. J. Faust Ankauf 1900/ Type" ( SMTD) . Paralectotype (here designated): 1 ♀, "N. Guinea S.E., Moroka , 1300m., Loria VII-IX 93/ 216/ Asytesta sejuncta , type" ( MSNG) .

Distribution. Papua New Guinea: Central Province: Moroka.

Remarks. A lectotype is designated for this species to ensure stability of nomenclature. One additional specimen was also seen by me in the MNHN collection with the same data labels as the lectotype but was not borrowed for examination. This specimen may also belong to the original type series.

Asytesta versuta Faust

( Figures 183–184)

Asytesta versuta Faust, 1898: 163 [key], 166 [description]. Holotype in SMTD, labeled “ versuta, Faust / N. Guinea Staudin. / Coll. J. Faust, Ankauf 1900/ Type.” Type locality: New Guinea (more specific locality information not provided).

Diagnosis. This species is superficially similar to A. gestroi and A. arachnopus ; however, neither species possess well developed lateral vittae on the pronotum and instead have well developed sublateral vittae. Asytesta versuta is more appropriately allied with A. sejuncta and A. fayae new species, from which it is distinguished by immaculate background vestiture, and sharply contrasting and unbroken dorsomedian and lateral vittae on the pronotum. Asytesta fayae also differs from this species by the irregular patch of white scales on the sides of the pronotum from the anterior margin to the base that is broader near the head and convergent towards the pronotal base. This band of scales becomes broad white vittae on intervals 9 of elytra spanning the area from the base to near the apex and males of that species apparently lack a tooth-like flange on the ventral margin of the protibia.

Redescription. Measurements (n =1): body length 8.2 mm, body width 3.8 mm, pronotal length 3.5 mm, elytral length 4.7 mm, rostral length 2.5 mm. Body elongate-oval, 2.2 X longer than broad; integument reddishbrown to dark brown, antennae and tarsi light reddish-brown. Clothed in sparsely distributed, dark tan to brown scales; integument not obscured by squamae except by more densely distributed yellowish-white scales in vittae, in sharp contrast to dark background squamae. Frons, base of rostrum, and pleura more densely clothed in white scales, white scales denser on tibia; rim of mesosternal receptacle with fine white scales; vestiture on legs evenly interspersed with white, hair-like setae, slightly denser along dorsal edge; male with suberect, hair-like setae along ventral margin of protibia, setae longer than elsewhere on tibia; tarsi clothed with dense, white, hair-like setae. Pronotal disk with dorsolongitudinal vitta from apex to base; pair of lateral longitudinal vittae from near anterior margin to base; anterolateral region of pronotum with distinct, broad, well defined, oblique, vittae from lower margin of postocular lobe to just above middle of procoxae ( Figure 183); dorsal portion of ring not extending beyond middle of procoxa. Elytra with broad sutural stripe of yellowish-white scales from midline to apex; pair of white longitudinal vittae on fifth intervals from humeri to subapex, vittae broader in apical 1/3 of elytra, occupying space between intervals 4–6; epipleural margin with white scales from base to apex; humeral and epipleural vittae connected by short fasciae along elytral base, fascia not reaching dorsum. Mesosternum and metasternum clothed in sparse white to tan scales, ventrites mostly without squamae, sparsely distributed light-brown squamae on ventrite 1 and 5, ventrites 2–4 more or less glabrous. Scales very fine to minute (indistinct under low magnification), appressed; white setae distinct, suberect to decumbent.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1 X broader than long, broadest slightly before base, subtruncate, medial area weakly produced. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one white, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, minute, smaller than punctures at base of elytra. Elytra 1.2 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, slightly elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by about 1/2 their length; mesofemora exceeding elytral apices. Protibia strongly compressed laterally, ventral margin with flange distinct, obtuse, tooth-like in male (as in Figures 48–49); dorsal margin strongly convex. Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. As in generic description.

Female terminalia. Female is not known.

Material examined. Holotype: ♂, " versuta, Faust / N. Guinea Staudin. [Staudinger?]/ Coll. J. Faust, Ankauf 1900/ Type" ( SMTD).

Distribution. New Guinea (specific locality not provided).

Remarks. Faust (1898) suggested that this species may be the male of A. gestroi , of which Faust only had females. I examined a series of male A. gestroi and determined that the two species are distinct (see diagnosis). Faust (loc. cite) indicated that this species was based on a single specimen, thus designation of a lectotype is not necessary.

Asytesta woodlarkiana Setliff , new species

( Figures 187–188, 191)

Diagnosis. The scattered, large white maculae (blotches) that are randomly distributed all over the otherwise black dorsum in combination with a complete dorsomedian longitudinal vitta and a broken transverse line transecting the dorsomedian vitta on the pronotum uniquely distinguish this species from its congeners. One other species, A. alexandriae new species may be confused with this species due to the similar markings on its pronotal disk; however, A. alexandriae has lighter brown vestiture and the mottling is less developed. Also, the transverse line on the pronotal disk is reduced to a pair of maculae in A. alexandriae that are joined to an apical pair of maculae by a fine line of pale scales, forming an L-shape on either side of the midline.

Description. Measurements (n = 7): body length 5.4–7.7 mm (mean 6.4 mm), body width 2.8–3.8 mm (mean 3.1 mm), pronotal length 2.2–3.4 mm (mean 2.6 mm), elytral length 3.2–4.5 mm (mean 3.8 mm), rostral length 1.9–2.3 mm (mean 2.1 mm). Body oval, 1.9–2.2 X longer than broad; integument dark reddish-brown, antennae and tarsi light reddish-brown. Sparsely covered with small, dark brown to fuscous scales, with more densely distributed dirty white scales in vittae and maculae. Frons, pleura, coxae, basal portion of femora, and venter more densely clothed in white to tan scales, tibia with black background scales; vestiture on femora and extreme base of tibia interspersed with white, hair-like setae; tibia otherwise immaculate; male with suberect, hair-like setae along ventral margin of protibia, setae longer than elsewhere on legs; tarsi clothed with sparse, white, hair-like setae. Dorsum dark, heavily mottled with large, irregular, circular maculae; maculae less densely distributed on sides. Pronotal disk with dorsolongitudinal vitta from apex to base; pair of short, curved, transverse maculae on either side of medial vitta at midline, not reaching sides; short, lateral longitudinal vittae from anterior margin to slightly before midline; anterolateral region of pronotum with distinct, well defined ring of white scales enclosing area from lateral margin of prosternal canal to anterior margin of procoxae, with darker background scales in center (as in Figure 104); dorsal portion of ring not extending beyond middle of procoxa. Elytra with thin fascia spanning entire anterior margin in most specimens, margin between intervals 5 bare in some specimens, fasciae reaching epipleural margin; humeri with short, rectangular, maculae on intervals 5 and 6; macula on intervals 5 longer than macula on intervals 6; intervals 5 with broken, longitudinal vitta on declivity; faint stripe on suture from slightly before midline to apex; epipleural margin lined with dense white scales from base to slightly past middle, epipleural vittae distinctly thicker in posterior half. Mesosternum and metasternum covered with dense white scales except transverse glabrous line at suture between sclerites, ventrites without scales except for sparse, evenly distributed, hair-like setae on ventrite 1 and 5 and lateral margins of 2–4, glabrous in middle. Scales fine (indistinct under low magnification), appressed to suberect.

Head. Visible in dorsal view, with semicircular, crown-like carinae on vertex above eyes (as in Figure 102), interrupted at back by glabrous V-shaped area where head articulates on pronotum, medial longitudinal carinae absent. Eyes flat, large, subcordate, posterior margin truncate; interocular distance narrower than subbasal width of rostrum. Rostrum shorter than pronotum, parallel sided to just before apex, slightly broadened at apex, with small medial interocular pit at base often obscured by squamae. Basal 1/3 of male rostrum rugose, coarsely punctate to apex, with suprascrobal carinae on lateral margins straight, not produced or forming lateral tooth-like process. Basal 1/3 of female rostrum less rugose than male, apical 2/3 smooth, finely punctate to apex. Clypeus truncate. Antennal club oval.

Thorax. Pronotum 1.1–1.3 X broader than long, broadest at broadest at base; medial area weakly produced, posterior margin subtruncate. Pronotal disk evenly convex at base, weakly rugose with small rasp-like, setose granules; granules evenly distributed; punctures not obscured by vestiture, shallow, evenly distributed, spaces between punctures less than 1/2 diameter of puncture, each bearing one brown, hair-like seta. Prosternal canal deep, lateral walls vertical; mesosternal receptacle very shallow, nearly obliterated, lateral margins weakly produced, posterior margin broadly open. Scutellum visible, diameter subequal to punctures at base of elytra. Elytra 1.1–1.3 X longer than broad, anterior margin truncate, declivity gradually declining to apex; punctures as on pronotum, basalmost punctures of each stria at anterior margin like all remaining punctures. Humeri not flattened or produced, subcontiguous with posterolateral margin of pronotum. Intervals convex, granulate. Granulate prominence on third intervals weakly elevated slightly before middle of elytra; prominence raised slightly above adjacent intervals, not distorting outline of dorsal contour in lateral view, terminating gradually at declivity; fourth intervals granulate, weakly produced, not elevated above fifth intervals, all other intervals not produced, lacking granulate prominences. Hind wing fully developed.

Legs elongate, hind femora exceeding elytral apices by slightly less than 1/2 their length; mesofemora exceeding elytral apices. Profemora unidentate; profemoral teeth not larger than teeth on meso- and metafemora; teeth simple, distal margin entire. Protibia strongly compressed laterally, ventral margin with flange distinct, obtuse, tooth-like in male, broader, gradually tapering to apex in female (as in Figures 48–49); dorsal margin strongly convex (as in Figures 51–52). Dorsal margin of middle and hind tibiae similarly convex. Apex of protibia with distinct supra-uncal process; uncus and premucro well developed, broadly separated with small flange situated between, supporting terminal setal brush. Tarsomere 2 short, trapezoidal, strongly flattened (as in Figure 64).

Abdomen. Ventrite 1 weakly distended, as long as or longer than remaining ventrites combined, on slightly lower plane than remaining ventrites, not overlapping ventrite 2; ventrite 2 partially folded into declivity of ventrite 1, slightly longer than ventrite 3 and 4. Intercoxal process on ventrite 1 broader than length of ventrite 1.

Male terminalia. Tergite VII broader than long, anterior margin convex, posterior margin emarginate at middle, wing binding patch distinct, with two rows of 6–8 minute plectral tubercles reaching posterior margin, tubercles greatly reduced, distinct under high magnification.

Female terminalia. Tergite VII longer than broad, anterior and posterior margins broadly convex, wing binding patch distinct, with two rows of 7–8 minute plectral tubercles; plectral rows not reaching posterior margin; tubercles visible minute, distinct under high magnification. Bursa more or less indistinct from vagina; vagina with walls weakly sclerotized; no internal sclerites visible.

Material examined. Holotype: ♂, " New Guinea: Papua , Woodlark I. (Murua), Kulumadau Hill, Mar. 23–30, 1957 / W. W. Brandt, Collector, Bishop" ( BPBM).

Paratypes: 2 ♂♂, " New Guinea: Papua, Woodlark I. (Murua), Kulumadau Hill, Jan 28–30, 1957 / W. W. Brandt, Collector, Bishop" ( BPBM) ; 1 ♂, " New Guinea: Papua, Woodlark I. (Murua), Kulumadau Hill, Feb. 3, 1957 / W. W. Brandt, Collector, Bishop" ( GPSC) ; 1 ♂, " New Guinea: Papua, Woodlark I. (Murua), Kulumadau Hill, Feb. 25, 1957 / W. W. Brandt, Collector, Bishop" ( BPBM) ; 1 ♀, " New Guinea: Papua, Woodlark I. (Murua), Kulumadau Hill, Mar 4–9 1957 / W. W. Brandt, Collector, Bishop" ( BPBM) .

Distribution. Papua New Guinea: Milne Bay Province: Woodlark Island (= Muyua Islands).

Etymology. The species epithet is based on the type locality.

Remarks. This species is difficult to assign to a species group since it may be considered intermediate between the elongate and long-legged form typical of the A. compressipes group and the more compact body form of the A. gazella group. Additional specimens are required help resolve the enigmatic placement of this species.