Baeocera satana Nakane, 1963

Ogawa, Ryo & Löbl, Ivan, 2013, A revision of the genus Baeocera in Japan, with a new genus of the tribe Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae), Zootaxa 3652 (3), pp. 301-326 : 322

Baeocera satana Nakane, 1963

( Figs. 10 , 19 , 32, 56, 61, 72)

Baeocera satana Nakane, 1963a: 22 ; Nakane, 1963b: 79; Löbl, 1984b: 188; Löbl, 1999: 726; Morimoto, 1985: 258. Eubaeocera satana: Löbl, 1968: 2 .

Diagnosis. Antennomere VII about 1.5 times as long as III; XI about 2.4 times as long as III. Elytral punctuation unevenly sparse and coarse. Lateral portion of metaventrite sparsely and finely punctuate. Parameres asymmetrical; inner sac with sclerites and fine scale-like and denticulate structures. Distal gonocoxite normal and apical portion setose. Bursa copulatrix sclerotized.

Redescription. Body broad-oval, shining. Head, pronotum, frons, tibiae and femora reddish-brown; disc of elytra dark reddish-brown; maxillary palpi, tarsi and antennomeres I–VI yellowish-brown; antennomeres VI–XI dark yellowish-brown. Head, pronotum and elytra sparsely and finely pubescent. Body color on male darker than the female.

Head with interocular distance about twice as long as eye width. Punctuation sparse and fine. Antennomeres I– VI with a few macrosetae, VII–XI covered with some macrosetae; VII about 1.5 times as long as III; XI about 2.4 times as long as III (Fig. 32).

Pronotum wider than long, lateral keel distinct except near basal angles. Punctuation sparse and fine, as on head. Scutellum wider than long, with exposed apex.

Elytra wider than long, widest at basal fourth, lateral margins gradually narrowed apically, minutely serrate at inner part of posterior margin. Elytral punctuation evenly sparse and coarse. Sutural striae extending outwards along basal margin to form basal stria, reaching humeral area and not joined with lateral stria.

Propygidium sparsely and coarsely punctuate; pygidium with 8 macrosetae, sparsely and coarsely punctuate in basal half, finely punctuate from basal half to apex, and gradually narrower toward apex.

Hypomeron impunctate. Mesoventrite sparsely and finely punctuate, as pronotum. Lateral portion of metaventrite sparsely and finely punctuate. Mesepimeron about third as long as wide. Mesocoxa about twice as wide as interspace between mesocoxa; mesocoxal line with row of long basal wrinkles; mesocoxal area narrow. Metepimeron almost as long as wide. Metanepisternum about third as long as wide. Ventrite I coarsely punctuate near base, sparsely and finely punctuate in posterior half, with row of long basal wrinkles and 4 macrosetae; V with 8 macrosetae, sparsely and finely punctuate, and concave at apex.

Meso- and metafemora with microsculpture, sparsely and coarsely punctuate.

Male. Protarsomeres I–III and mesotarsomeres I and II enlarged with tenent setae. Hind tibiae sinuate and bearing spines in basal fifth. Aedeagus about 1.0 mm long; parameres asymmetrical; inner sac with sclerites and fine scale-like and denticulate structures ( Fig. 72 ).

Female. Protarsomeres I–III weakly enlarged without tenent setae. Hind tibiae not sinuate. Distal gonocoxite normal and robust (Fig. 56); vagina membranous, without robust sclerites; bursa copulatrix strongly sclerotized ( Fig. 61 ). Spermathecae undetectable (see Fig. 61 , a pair of spermatheca possibly connected with sclerotized portion of vagina in the middle part).

Measurements (n = 4). Length (PL+EL): 1.91–2.01 mm; width (PW, EW): 1.03–1.10 mm, 1.21–1.25 mm. PL/PW: 0.76–0.79; EL/EW: 0.88–0.94. HW: 0.48–0.50 mm. ID: 0.23 mm.

Approximate ratio of each antennal length (width) as follows (n = 1); 2.0 (0.8): 1.5 (0.7): 1.0 (0.4): 1.1 (0.3): 1.4 (0.3): 1.2 (0.4): 1.5 (0.4): 1.4 (0.6): 1.7 (0.5): 1.7 (0.6): 2.4 (0.8).

Specimens examined. 1Ƥ, Karasuhara, Hyogo Pref., 3. VIII. 1974, T. Takahashi leg.; 2Ƥ, Shiromawari, Uchiko-Town, Ehime Pref., 9. VII. 1995, E. Yamamoto leg.; 13, Tsushima Is., Shushi-gawa River, Kamitsushima- Town, Nagasaki Pref., 16. VII. 2000, T. Kan leg. (All preserved in EUMJ).

Distribution. Japan: Honshu, Shikoku, Kyushu.

Figures

FIGURES 2 – 10. Habitus in dorsal view of the Japanese Baeocera. 2, B. frater; 3, B. myrmidon; 4, B. sordida; 5, B. abnormalis; 6, B. caliginosa; 7, B. curtula; 8, B. nakanei; 9, B. takizawai; 10, B. satana. Scales: 1.0 mm.

FIGURES 11 – 23. Habitus in lateral view of the Japanese Baeocera, and Baeoceroxidium micros (20, 21) and Baeocera falsata (22, 23). 11, B. frater; 12, B. myrmidon; 13, B. sordida; 14, B. abnormalis; 15, B. caliginosa; 16, B. curtula; 17, B. nakanei; 18, B. takizawai; 19, B. satana; 20, 22, dorsal view; 21, 23, lateral view. Scales: 1.0 mm.

FIGURES 68 – 75. Parameres (68, 69) and aedeagus (70 – 75), dorsal views (68 – 74) and lateral view (75). 68, 70, Baeocera nakanei; 69, 71, B. takizawai; 72, B. satana; 73, B. falsata; 74, 75, Baeoceroxidium micros. Scales: 0.2 mm.

FIGURES 58 – 61. Spermathecae and the apertures. 58, Baeocera caliginosa; 59, B. abnormalis; 60, B. nakanei; 61, B. satana. Allows: opening portions of spermathecae on the vagina. Scales: 0.1 mm.