Bruchomyia Alexander, 1921

Wagner, Rüdiger & Stuckenberg, Brian, 2016, Cladistic analysis of Subfamily Bruchomyiinae (Diptera: Psychodidae), Zootaxa 4092 (2), pp. 151-174 : 161-163

Genus Bruchomyia Alexander, 1921

Bruchomyia Alexander, 1921: 402 .

Type species: Bruchomyia argentina Alexander, 1921 (orig. des.)

Brachomyia: Enderlein, 1936: 108 (incorrect spelling)

Species included: B. almeidai [ Brazil]; B. andina Quate [ Peru]; B. argentina [ Argentina ]; B. brasiliensis [ Brazil]; B. fusca [ Brazil]; B. mineira [ Brazil]; B. peruviana (female) [ Peru]; B. plaumanni [ Brazil]; B. shannoni [ Peru]; B. unicolor [ Brazil].

Diagnosis. Antenna with 28 to 32 antennomeres. Anal vein elongate, almost straight. Males with paired vasa deferentia widening towards testes. Ejaculatory apodeme flared (trumpet-shaped) at base, much longer than aedeagus. Gonocoxites with tubercles on medial surface bearing remarkable clusters of setae, gonostyli bifurcate. Ventral part of male genitalia reduced in size compared to epandrium and cerci. Females with spermatheca ovoid, with a longitudinal slit.

Comments. the diagnostic characters given above can be used to easily distinguish Bruchomyia ; however, the following features are also worth noting: number of antennal flagellomeres increased to 28-32. Antennae 6-8 times longer than head diameter, and about 3 times longer than palps. Ascoids subdiscoidal. Palpus approx. 2 times longer than head, palp segment 5 longer than combined length of segments 1 to 4, palpus without Newstead scales. Wing tip between R3 and R4; M2 originates at the end of second basal cell or more basally. CuA1, CuA2, and a long almost straight cross veins sometimes incomplete (fig. 3a). No lateral hair tufts on abdominal pleurae. Inversion of male genitalia by segments 8 and 9; gonocoxite and gonostyli reduced in size compared with tergite 9. Gonocoxites with tubercle on inner side bearing clusters of setae or spines, gonostylus bifurcate in all species. Ejaculatory apodeme trumpet shaped, strongly increasing in circumference at base and much longer than aedeagus (Bravo & Barrata 2012), parameres vase-shaped (details remain unclear—definitely different from other Neotropical Bruchomyiinae ). Vasa deferentia join to a comparatively long (~ 20% of ejaculatoty apodeme length) ejaculatory duct before entering the aedeagus (fig. 4c). Female spermatheca flat oval with a longitudinal slit (compare Fairchild 1952, figure 55, p. 277).

Bruchomyia is apparently Neotropical but its distribution area appears split. Three species B. andina , B. peruviana , and B. shannoni occur in the Andes in western South America, while the others were mentioned from the eastern part of the continent. No species is yet known from the vast area in between. To interpret relations additional information on species and even more detailed descriptions are needed.