Calendula suffruticosa subsp. boccoyana A.C. Gonç. & P. Silveira, 2023
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publication ID |
https://doi.org/ 10.11646/phytotaxa.605.1.1 |
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persistent identifier |
https://treatment.plazi.org/id/E82AFC0A-1564-CA44-66ED-FB70844DFDA1 |
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treatment provided by |
Plazi |
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scientific name |
Calendula suffruticosa subsp. boccoyana A.C. Gonç. & P. Silveira |
| status |
subsp. nov. |
11a. Calendula suffruticosa subsp. boccoyana A.C. Gonç. & P. Silveira , subsp . nov. Type:— MOROCCO. Al Hoceïma, Bokkoya, Taoussarte, 133 m [35°13’08” N, 4°05’14” W], 14 May 2014, P. Silveira & A.C.R.S. Gonçalves 3340 (holotype RAB!; isotypes AVE!, MARK!). Diagnosis:— Calendula suffruticosa subsp. boccoyana differs from subsp. suffruticosa by the young parts ± white-arachnoid, as opposed to being entirely ± glabrous to glandulose, and by the cymbiform aquenes width (3.2) 4–5.5 (8) mm, as opposed to (6.7) 7.7–10.8
(11.8) mm. Calendula suffruticosa subsp. boccoyana differs from subsp. riffensis by the larger and thicker basal leaves which are
(6.2) 8.5–9 (9.5) × (1.8) 2.1–2.8 (3) cm (as opposed to <8.3 × 2.3 cm) and 1.7–2.2 mm thick (as opposed to <0.6 mm thick).
Description:––Perennial herbs. Stems (17) 24–39 (50) cm long, prostrate to decumbent, with both glandulose and white-arachnoid pubescence, the latter predominant in younger parts. Basal leaves (6.2) 7.4–9.3 (9.5) × (1.2) 1.3–1.8 cm, (0.3) 0.4–0.6 (0.8) mm thick, oblanceolate to spatulate, apex obtuse, or sometimes ± acute, margins sub-entire to sinuate-dentate, with both glandulose and white-arachnoid pubescence, the latter predominant in younger parts. Capitula solitary, (2) 2.2–2.8 (3.2) cm diameter. Outer achenes rostrate (10) 13–19.5 (24) × (1) 1.5–2 (2.3) mm, straight or slightly curved, without dorsal spines, sometimes with one tooth at the base and/or at the apex; middle achenes bialate (13.8) 14.4–17.5 (18) × (3) 3.3–8 (8.7) mm, with a rostrum (3.3) 5–9.3 (10) mm; most frequently cymbiform (4.7) 5.8–8.3 (8.8) × (3.2) 4–5.5 (8) mm, with ventral wings surpassing the lateral ones; inner achenes vermiculate-alate (5) 5.2–5.3 × (2.2) 2.7–3.7 (4.8) mm, and vermiculate-exalate (3) 3.8–5 (5.5) × (1.2) 1.4–2 (2.2) mm, both circular. Figure 17 View FIGURE 17 .
Habitat and distribution:— Limestone rocks/cliffs, at an elevation of ± 133 m. It grows in a semi-arid Mediterranean bio-climate. Endemic to Morocco with a restricted distribution in the coastal mountains of the National Park of Al Hoceïma (Bokkoya) to El-Yebha .
Conservation status:—Endangered (EN). This taxon is under numerous threats, especially due to climate change (in some places rise in sea level, extreme weather events) and drought. The subspecies is rare and local with a small-restricted range; it is known from two populations located in hillside slopes with difficult access. The number of mature individuals is estimated to be <50 in one of the populations, based on field observations, but accurate data for other localities in coastal cliffs of difficult access are lacking. The AOO is 8 km 2, which qualifies the taxon for EN. Therefore, we propose its conservation status as EN B2a.
Chromosome number:— 2n = 32.
Genome size:—3.17 ± 0.06 pg.
Eponymy:—For Bokkoya in Morocco where it occurs.
Additional collections:— MOROCCO. Tanger-Tetouan-Al Hoceïma : El-Jebha (El-Yebha), 90m [35°12’39” N, 4°39’46” W], 11 June 2012, P GoogleMaps . Silveira & A. C. R. S . Gonçalves 3266 ( AVE!, RAB!); Al Hoceima, El-Yebha, 400 m, [35°12’36” N, 4°39’50” W], 9 April 1994, J. M GoogleMaps . Montserrat , B . Valdés s.n. ( BC!); 400 m, [35°12’36” N, 4°39’50” W], 9 April 1994, J. M GoogleMaps . Montserrat , B . Valdés 2948 94 ( SEV!); Ras Sidi-el-Ahbed (Bokkoya), [35°11’ N, 4°07’ W], 16 April 1929, P GoogleMaps . Font Quer s.n. ( BC!); [35°11’ N, 4°07’ W], 7 March 1929, P GoogleMaps . Font Quer s.n. ( BC!) .
| P |
Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants |
| A |
Harvard University - Arnold Arboretum |
| C |
University of Copenhagen |
| R |
Departamento de Geologia, Universidad de Chile |
| S |
Department of Botany, Swedish Museum of Natural History |
| AVE |
Universidade de Aveiro |
| RAB |
Institut Scientifique |
| J |
University of the Witwatersrand |
| M |
Botanische Staatssammlung München |
| B |
Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet |
| BC |
Institut Botànic de Barcelona |
| SEV |
Universidad de Sevilla |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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