Camchaya loloana var. mukdahanensis H.Koyama, Acta Phytotax. Geobot. 35(1-3): 52. 1984.

Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C., 2014, Revisions and key to the Vernonieae (Compositae) of Thailand, PhytoKeys 37, pp. 25-101 : 53-54

Camchaya loloana var. mukdahanensis H.Koyama, Acta Phytotax. Geobot. 35(1-3): 52. 1984.

Type.

Thailand, Mukdahan, Muang District, Dongman Village, H. Koyama et al. T-30941 (holotype: KYO!). Fig. 6C .

Description.

Annual herbs, 10-70 cm tall. Stems erect, terete, inconspicuously ribbed, scabrous with uniseriate hairs and T-shaped hairs. Leaves alternate, 5-11 by 2-5 cm, ovate or lanceolate, margin serrate or undulate, apex acute or acuminate, base attenuate, chartaceous; both surfaces scabrous with whip-shaped hairs, cylindrical hairs and capitate glands; lateral veins 5-9-paired; petioles up to 15 mm long. Capitulescences terminal and axillary, paniculate. Capitula campanulate, 9-10 mm long, pedunculate. Receptacle convex, 2-2.5 mm in diam., glabrous. Involucres campanulate, in 5-6 series, 7-9 mm long, 4-6 mm in diam. Phyllaries greenish with purple apex, margin with spines up to 0.2 mm long, outer surface arachnoid, glands capitate; the outer lanceolate, apex spinose; the inner ones lanceolate to oblong, apex acuminate. Florets 30-70; corollas funnelform, purple, puberulous, glands capitate; corolla tubes ca. 5.5 mm long; corolla lobes 1.5-2 mm long. Anthers 1.5-2 mm long, apical appendage acute, base rounded. Styles purple, 5-6 mm long, branches ca. 2 mm long. Achenes obovate, 1.3-5 mm long, glabrous, 10-ribbed. Pappus in one series of bristles 1.5-2 mm, absent or deciduous.

Distribution.

Thailand: Nong Khai, Mukdahan, Ubon Ratchathani. Laos.

Specimens examined.

Thailand, Mukdahan, Phu Pha Thoep national park, 16°26.08'N, 104°48.33'E, 22 Oct 2007, S. Bunwong 338 (KKU, US); Phu Pha Thoep national park, 12 Dec 1982, H. Koyama, H. Terao & Th. Wongprasert T-30866 (BKF); Phu Pha Thoep national park, 13 Dec 1982, H. Koyama et al. T-30941 (KYO); Mukdahan, Cham Cha Iee, 13 Dec 1982, H. Koyama, H. Terao & Th. Wongprasert T-30904 (BKF, L); Nong Khai, route number 2186, 16 Dec 1982, H. Koyama, H. Terao & Th. Wongprasert T-31137 (BKF, L); Ubon Ratchathani, Sirinthon Dam, 9 Dec 1982, H. Koyama, H. Terao & Th. Wongprasert T-30662 (BKF, L); Sirinthon Dam, M. Norsangsri 1158 (QBG); Gang Ta na national park, 27 Oct 2007, S. Bunwong 343 (KKU, US),

Diagnostic characters.

Camchaya loloana var. mukdahanensis differs from the typical variety by having smaller capitula, a shorter involucre, and fewer florets.

Ecology.

Rocky area in dipterocarp forest, alt. 250-400 m; flowering August to January.

Vernacular name.

Phu Muang (พู่ม่วง).

Figures

Figure 6. Morphology of Vernonieae in Thailand 2. A Camchaya gracilis B Camchaya loloana C Camchaya loloana var. mukdahanensis D Camchaya pentagona E-F Camchaya spinulifera G Camchaya tenuiflora H-I Camchaya thailandica.