Cephennodes (Cephennodes) paramartensi, Jałoszyński, 2017

Jałoszyński, Paweł, 2017, Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4349 (1), pp. 1-120 : 45-47

Cephennodes (Cephennodes) paramartensi sp. n.

( Figs 97 , 107 , 124–127 , 145 )

Type material. Holotype: NEPAL (Sankhuwasabha District): ♂, "E. NEPAL: KOSI / Forêt N-E Kuwapani / 2250 m, 24.IV.84 / Löbl & Smetana" [white, printed] ( MHNG) . Paratypes (3 exx): 2 ♂♂, "E. NEPAL: KOSI / Forêt N-E Kuwapani / 2350 m, 5.IV.84 / Löbl & Smetana" [white, printed] ; 1 ♂, " NEPAL: Bagmati / Pokhare NE Barah- / bise, 2800m, 3.5.81 / Löbl & Smetana " [white, printed] (paratypes in MHNG and cPJ).

Diagnosis. BL ~ 1.4 mm; body stout, with elytra weakly narrowing behind the broadest site; distal portion of metatibia strongly thickened, with oval subapical porous field much broader than long and with one remarkably long bristle within short setal fringe of apical margin; abdomen in ventral view with sternite II bearing a microserrate carina near its posterior margin and sternite III bearing broadly rounded median lobe; apical portion of median lobe of aedeagus evenly rounded, its longitudinal axis forming about 45°-angle with long axis of aedeagus; apical projection with sinuate distal margin.

Description. Body of male ( Fig. 97 ) brown, with shallow constriction between pronotum and elytra, strongly convex, covered with light brown setae; BL 1.43–1.45 mm (mean 1.43 mm).

Head subtrapezoidal, HL 0.18–0.20 mm (mean (0.18 mm), HW 0.38–0.40 mm (mean 0.38 mm); vertex and frons confluent, evenly and weakly convex; frons with a pair of tiny tubercles; supraantennal tubercles small and weakly elevated; eyes large, strongly convex, coarsely faceted. Punctures on frons and vertex small but distinct, relatively sharply marked, densely and unevenly distributed, sparser at middle and denser on sides, those in middle of frons between eyes separated by spaces 1–2 × as wide as diameters of punctures; setae short, moderately dense, suberect to erect. Antennae relatively short and moderately slender, AnL 0.75–0.80 mm (mean 0.77 mm), pentamerous club indistinctly delimited; antennomere I 1.5 × as long as broad; II slightly shorter and narrower than I, about 1.7 × as long as broad; III slightly narrower and much shorter than II, indistinctly transverse, IV–VI each similar in length and width, each as broad as III but slightly longer, about 1.1 × as long as broad; VII slightly broader and distinctly longer than VI, 1.2 × as long as broad; VIII slightly broader but much shorter than VII, distinctly transverse; IX much larger than VIII, distinctly transverse; X slightly larger than IX, distinctly transverse; XI much broader than X, about as long as IX–X together, slightly less than twice as long as broad.

Pronotum in dorsal view nearly semicircular, broadest slightly in front of middle but very indistinctly narrowing posteriorly, PL 0.45 mm, PW 0.70–0.73 mm (mean 0.71 mm); anterior margin broadly and weakly rounded; lateral margins not serrate, strongly rounded in anterior half, nearly straight in posterior half; posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral carinae narrow and fused with lateral margins; antebasal pits small and very shallow, each located distinctly closer to posterior than to lateral margin of pronotum. Punctures on disc very small and shallow, inconspicuous; setae dense, short, suberect.

Elytra short, oval, broadest between middle and anterior third, EL 0.80 mm, EW 0.75–0.78 mm (mean 0.77 mm), EI 1.03–1.07. Humeral calli weakly elevated; subhumeral lines very narrow but distinct, equal to about 0.4 EL, each developed as sharp stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron large and connected to short, slightly arcuate impression extending posteromesally; apices of elytra separately rounded. Punctures on elytra more distinct than those on pronotum but superficial, diffuse and inconspicuous; setae about as dense and short as those on pronotum but less erect.

Hind wings present, long.

Metaventrite with large and sharply delimited postmesocoxal impressions, median part covered with very fine punctures.

Hind legs with strongly modified tibiae ( Fig. 107 ); in dorsal view lateral (external) tibial margin distinctly recurved and spatulate portion of tibia rapidly narrowed distally; dorsodistal porous field much broader than long; distal fringe of setae with one outstanding long bristle.

Abdomen ( Figs 124–125 ) with sternite II bearing long and microserrate carina along median portion of its posterior margin; sternite III with broadly rounded median lobe ( Fig. 125 ).

Aedeagus ( Figs 126–127 ) moderately stout; AeL 0.18 mm, apical portion of median lobe nearly evenly rounded, its longitudinal axis forming about 45°-angle with the long axis of aedeagus; apical projection with sinuate distal margin; parameres long and slender, not exceeding apex of median lobe, each bearing one apical seta.

Female. Unknown.

Distribution ( Fig. 145 ). Eastern Nepal.

Etymology. The name paramartensi was chosen to emphasize similarity between this species and C. martensi .

Remarks. This species is very similar to C. martensi ; they share a similarly large and stout body and elytra less narrowed behind the broadest site than those in C. dolakhanus . Cephennodes paramartensi can be distinguished from C. martensi on the basis of the antennomere XI distinctly broader than X and distinctly less then twice as long as broad (in C. martensi barely broader than X and distinctly twice as long as broad); the lateral (external) margin of metatibia less recurved; the abdominal sternite II with microserrate carina running along its posterior margin ( C. martensi lacks such a carina, instead the posterior margin of the sternite II is projected posteriorly and forming a very broad subtriangular and rounded lobe, absent in C. paramartensi ); and the aedeagus with the apical portion of median lobe very strongly curved outwards from the long axis of aedeagus (much less so in C. paramartensi ); and the distal margin of apical projection concave distinctly closer to its middle, whereas the concavity in C. martensi is closer to the apex of apical projection. See also remarks for C. monolaminatus .

Figures

FIGURES 94 – 101. Dorsal habitus of Cephennodes (s. str.) dolakhanus s sp. n. (94), C. (s. str.) manangensis sp. n. (95), C. (s. str.) martensi sp. n. (96), C. (s. str.) paramartensi sp. n. (97), C. (s. str.) monolaminatus sp. n. (98), C. (s. str.) thakanus sp. n. (99), C. (s. str.) annapurnaensis sp. n. (100), and C. (s. str.) parbatensis sp. n. (101).

FIGURES 102 – 111. Right metatibia of male in dorsal view. Cephennodes (s. str.) dolakhanus s sp. n. (102, 104), C. (s. str.) manangensis sp. n. (103, 105), C. (s. str.) martensi sp. n. (106), C. (s. str.) paramartensi sp. n. (107), C. (s. str.) monolaminatus sp. n. (108), C. (s. str.) thakanus sp. n. (109), C. (s. str.) annapurnaensis sp. n. (110), and C. (s. str.) parbatensis sp. n. (111). 104 – 111 not to the same scale.

FIGURES 112 – 127. Abdomen of male in right lateral (112, 116, 120, 124) and ventral (113, 117, 121, 125) views and aedeagus in ventral (114, 118, 122, 126) and lateral (115, 119, 123, 127) views. Cephennodes (s. str.) dolakhanus sp. n. (112 – 115), C. (s. str.) manangensis sp. n. (116 – 119), C. (s. str.) martensi sp. n. (120 – 123), C. (s. str.) paramartensi sp. n. (124 – 127). Abbreviations: aml, apex of median lobe; ap, apical projection.

FIGURES 144 – 145. Distribution of Cephennodes (s. str.) dolakhanus sp. n. (144, circle), C. (s. str.) manangensis sp. n. (144, triangle), C. (s. str.) martensi sp. n. (145, circle), C. (s. str.) paramartensi sp. n. (145, triangle).