Ceratophyus amdoensis Král & Jákl, 2020

Král, David, Jákl, Stanislav, Lu, Yuanyuan & Bai, Ming, 2020, The genus Ceratophyus Fischer von Waldheim, 1824 (Coleoptera: Geotrupidae) in China, along with description of a new species, Zootaxa 4747 (2), pp. 339-349 : 340-341

Ceratophyus amdoensis Král & Jákl , new species [安多角粪金龟]

( Figs. 1–6 )

Type locality. China, Gansu, 50 km SE of Shandan, 38°22’N 101°20’E, 2300 m a. s. l.

Type material. China, Gansu: Holotype: ♂ ( NMPC), “CHINA–GANSU | 50km SE SHANDAN | 38°22’N, 101°20’E || 15-25-05.2005 H= 2300m | leg/Coll Siniaev [printed]” . Paratypes: 1 ♂ ( NMPC), same data as holotype ; Qinghai: 1 ♂ ( SJCP), “CHINA-Qinghai, Nan Shan | Heimahe, 3.-7.VIII.1994 | 3200-3500 m [printed]” .

Description of holotype (♂). Weakly convex, black, dorsal surface shining; macrosetation of lateral margin of head and pronotum and ventral surface blackish ( Fig. 1 ).

Head ( Figs. 1, 3 ). Mandibles considerably protruding anteriorly, with broadly rounded, bisinuate external outline, smooth. Labrum transverse, broader than long, anterior edge weakly emarginate, surface distinctly punctate. Clypeus triangularly elongated anteriorly and slightly upward, with long, acute triangular apical process. Clypeal horn including its base finely, sparsely punctate. Oblique keels above eyes distinct, reaching to level of anterior outline of eyes; genae elongated in distinct tooth anterolaterally. Clypeus shallowly, sparsely but distinctly punctate, each puncture very fine, separated by about ten puncture diameters. Frons, vertex and occiput except area of clypeal horn impunctate.

Pronotum ( Figs. 1, 3 ) distinctly transverse, broadest at midlength, entirely bordered; lateral margins broadly widened; anterior angles broadly rounded; lateral fovea deeply, coarsely and densely punctate; midline extremely shallow to absent. Punctation of pronotum coarse and dense, disc distinctly doubly punctate. Pronotal horn well developed, straight, narrowly and weakly upturned in lateral aspect, without subapical tooth ventrally.

Scutellum cordate, transverse, impunctate.

Elytra ( Figs. 1, 3 ) distinctly longer than wide, maximum width posterior of middle; surface moderately shiny; between suture and humerus with 7 striae, all of them well impressed along entire length of elytron, stria 5 distinctly shortened before base; intervals convex, almost impunctate; humeral umbone distinct; base of epipleura in area of humeral umbone distinctly narrowed to apex of elytron.

Macropterous.

Legs ( Figs. 1–2 ). Femora unarmed, glabrous, impunctate, with two macrosetaceous transverse carinae. Protibia with 8 external teeth gradually diminishing proximally, apical tooth distinctly bifid, ventromedial edge unarmed; meso- and metatibiae with 3 transverse external carinae.

Ventral surface ( Fig. 2 ). Abdominal ventrites scabrous, coarsely and densely punctate, with short, recumbent, greyish to reddish colored setation.

Male genitalia. Parameres of characteristic shape, as in Figs. 4–6 .

Sexual dimorphism. Female unknown.

Measurements. Total body length: Holotype, 23.0 mm; paratype (Shandan), 18.5 mm, paratype (Heimahe), 21.5 mm.

Variability. In small paratype from Shandan head horn short, more or less straight, and pronotal horn less developed, much shorter than in holotype.

Differential diagnosis. Refer to the species key below.

Etymology. Toponymic; an adjective derived from Amdo, the name of one of three traditional regions of Tibet (nowadays approximately consisting of the Qinghai and western Gansu Provinces, where the new species was collected.

Distribution. China (Gansu, Qinghai); see map on Fig. 19 .

Figures

FIGURE 19. Sketch map of China with marked distribution of Ceratophyus Fischer von Waldheim, 1824 species.

FIGURES 1–3. Ceratophyus amdoensis Král & Jákl, new species, habitus of holotype (♂). 1, dorsal view; 2, ventral view; 3, left lateral view. Not to scale.

FIGURES 4–12. Aedeagus of Ceratophyus Fischer von Waldheim, 1824 species. 4–6, Ceratophyus amdoensis Král & Jákl, new species, holotype; 7–9, C. dauricus (Jekel, 1866) (“Transbaikalia”, NMPC); 10–12, C. kabaki Nikolajev, 2007 (Sichuan, Yading, NMPC). 4, 7, 10, dorsal view; 5, 8, 11, left lateral view; 6, 9, 12, tips, ventral view. Not to scale.