Chilicola (Chilioediscelis) patagonica Toro and Moldenke

Willis, Erin & Packer, Laurence, 2008, Revision and phylogenetic analysis of Chilioediscelis (Hymenoptera: Colletidae) with descriptions of three new species, Zootaxa 1762 (1), pp. 29-52 : 36-37

Chilicola (Chilioediscelis) patagonica Toro and Moldenke

Diagnosis: Males can be differentiated from all other species except C. andina by the colouration of their legs (See diagnosis for C. andina on how to separate these two species from all others and each other). Females of the species can be distinguished by their unique metasomal colouration: T1 mostly dark brown, remaining terga red, suffused with brown on sides and across disc of T2. Other species with a mostly red metasoma, C. penai and C. sardonyx , have brown colouration restricted to the sides on T1 and T2; while those with more reduced red have dark colouration extending across at least T1-T3.

Description: Male: Length 5.4mm, forewing length 4.2mm, head width 1.6mm, intertegular span 1.3mm.

Colouration: As in C. andina except yellow as follows, labrum either above or entirely (varies among individuals), clypeus mostly, maculation on lower paraocular area to just above dorsal margin of clypeus (present along dorsolateral margin of the clypeus), dorsoapical spot on all femora, anterior surface of pro- and mesotibia, most of metatibia (at most a small dark spot on outer surface and carina brown), pro- and mesobasitarsus entirely, metabasitarsus basally.

Pubescence: As in C. andina except hairs of sterna sparser, not noticeably longer on S1 and S2 (= 2MOD).

Surface Sculpture: As in C. andina except as follows, gena striate close to mandible base, punctures on gena and hypostomal area deeper and somewhat denser (i=1-2d); mesoscutum with punctures deeper and slightly sparser (i=1–4d); punctures on mesopleuron denser and deeper (i=1–2d), noticeably sparser above level of scrobe; dorsal area of propodeum reticulate basally and striate apically.

Structure: As in C. andina except as follows: Head: Labrum not concave dorsally. Apex of clypeus more strongly concave medially in apical view. Frontal line not raised above surface of frons, difficult to detect among striae. Vertex concave in frontal view. Ratio of pedicel:F1:F2:F3 (8:7:6:7). Flagellomeres broad, length only slightly greater than breadth. Malar space shorter, length:basal depth of mandible (3:7). Gena narrower, ratio of width:eye (9:17).

Mesosoma: Ratio of scutellum:metanotum:propodeum (12:5:7). Scrobal groove entire. Ventral surface of metatrochanter acutely angulate in lateral view. Metafemur not so swollen, length to breadth (41:22). Metatibia more swollen length to breadth (45:15). Metabasitarsus shorter, less than 5 times as long as greatest depth (27:6). Ratio of length of stigma to marginal cell on wing margin (15:23). Stigmal margin in marginal cell convex.

Metasoma: S7 with ventral lobe elongate, outer margin irregular and bearing uneven row of long setae; dorsal lobe slightly posteriorly directed, narrower than that of C. andina . S8 as in C. andina except apicolateral margins of apical lobe more broadly rounded. Genital capsule broadened towards apical 0.33 of gonocoxite. Penis valve with one very large, membranous lobe and one comparatively narrow one.

Description: Female: Length 5.6mm, forewing length 4.2mm, head width 1.6 mm, intertegular span 1.3mm.

Colouration: As in C. andina except pro- and mesotibia only yellow basally. T1 mostly dark brown, remaining terga red, suffused with brown on sides and across disc of T2, antecostal areas black, those of T3- T5 visible through previous tergum and appearing as narrow black transverse bands.

Pubescence: As in C. andina except: apical hair bands only evident on T1 & T2.

Surface Sculpture: As in male except as follows: punctures finer, punctures on gena and hypostomal area denser (i=d); facial striae stronger; mesoscutum with punctures sparser (i=1–6d); mesopleuron with punctures not noticeably sparser above scrobe than below.

Structure: As in male except as follows: Head: Eyes less convergent below, UOD:LOD (36:32). OOC:IOC, (9:11). Scape longer and narrower (15:3); ratio of pedicel:F1:F2:F3 (7:6:4:5). Ratio of widths of gena:eye (7:15).

Mesosoma: As in C. andina except: more robust, length to breadth (80:60). Hind leg unmodified. Metabasitarsus 6 times longer than greatest depth (30:5).

Metasoma: As in C. andina except: apical impressed areas more extensive, more than 0.33 X as long as corresponding tergum.

Material Studied: Holotype, allotype and one paratype of each sex ARGENTINA, Santa Cruz, Los Antiguos, 21.xi.1966 Irwin & Schlinger. Two males and one female, CHILE, Magallanes, Lag. Azul, xii.1978 W. Sielfeld. Two females ARGENTINA, Santa Cruz, Lago Belgrano, i.1978 E. Maury. One male and one female, ARGENTINA, Santa Cruz, 25km S of Los Antiguos S46.42.654 W071.40.422, 653m, 22.xi.2003 L. Packer. The types and Sielfeld’s specimens are at CASC; the remaining material is in PBYU.