Cimbrochrysa americana, Makarkin & Antell & Archibald, 2022

Makarkin, Vladimir N., Antell, Gwen S. & Archibald, S. Bruce, 2022, A revision of Chrysopidae (Neuroptera) from the late Eocene Florissant Formation Colorado, with description of new species, Zootaxa 5133 (3), pp. 301-345 : 313-315

Cimbrochrysa americana sp. nov.

Figs 8 , 9

Type material. Holotype UCM 33497 (part), deposited in UCM, collected by W.P. Cockerell. A poorly preserved specimen with four well-preserved wings overlapping each other.

Type locality and horizon. USA: Colorado: Teller County: Florissant (Station 14, NW SE NW Section 11, Township 13S, Range 71W); Eocene (late Priabonian; Florissant Formation) .

Etymology. From America, referring to its occurrence in North America.

Diagnosis. May be distinguished from other species of the genus by <10 branches of RP [ C. moleriensis , C. major sp. nov.: ≥10], and more distal position of 1r-m within im (at proximal one-third length) [ C. moleriensis , C. major sp. nov.: more proximal position (from near fork of M to proximal one-fifth length)].

Description. Head mostly poorly preserved; only one eye (diameter ca. 0.4 mm), basal parts of antennae clearly preserved. Thorax fragmentarily preserved. Legs poorly preserved, covered with dense, short setae. Abdomen not preserved.

Forewing 11.9 mm long, 4.3 mm wide (length to width ratio 2.77). Costal space moderately broad, widest rather far proximad origin of RP. Subcostal veinlets simple proximad pterostigmal region, rather widely spaced. Pterostigma indistinct; incorporated veinlets, crossveins not preserved. Distal part of Sc not preserved. Subcostal space relatively narrow; subcostal crossveins not detected. RA ending on margin before wing apex; its veinlets not preserved. RA space moderately broad, with nine crossveins. RP originating relatively far from wing base (at ca. 0.23 of complete length). Anterior trace of RP zigzagged, apically deeply forked, with eight branches, each forked once near margin except RP7–RP8 (left wing) and RP6–RP8 (right wing), which remain simple to margin. Basal crossvein between R, M (1r-m) variable in length (short in right wing, twice that length in left wing), connecting RP, MA within im at proximal one-third length. M dividing to MA, MP slightly distad origin of RP. MA zigzagged, deeply forked at Psc (one branch shallowly forked in left wing); MP zigzagged, deeply forked at Psc. Crossvein 1im moderately long. Im broad (length/width ratio 2.53–2.66), pentagonal. Psm poorly developed, strongly zigzagged. Crossvein 2m-cu (between im, CuA) located nearly in middle of im (slightly in proximal portion). CuA probably with two simple branches. CuA continuing into well-developed Psc, then to outer gradate series of crossveins. CuP deeply forked, its branches widely spaced. c1: c2 length 0.50: 1 in left wing, 0.59: 1 in right wing. A1 arched, simple. A2 strongly arched, simple. Crossvein between A1, A2 relatively short; crossvein between A2, A3 incompletely preserved, oblique. Two gradate series of crossveins nearly parallel; inner series slightly sigmoid, with eight crossveins distad MA; outer series not sigmoid, with seven crossveins distad RP2.

Hind wing ca. 10.6 mm long, ca. 3.8 mm wide. Costal space poorly preserved. Preserved subcostal veinlets simple, widely spaced. Pterostigma indistinct; incorporated veinlets not clearly discernible. Sc fragmentarily discernible. Subcostal space narrow; subcostal crossveins not detected. RA entering margin before wing apex; its veinlets poorly preserved. RA space broad proximally, moderately narrow distally; with eight (right wing) or nine (left wing) crossveins. RP originating relatively far from wing base. Anterior trace of RP zigzagged, apically simple; with eight branches, each forked once near margin except RP7–RP8 (left wing) and RP6–RP8 (right wing), which are simple. Proximal parts of RP, MA fused for long distance. Psm well developed, slightly zigzagged. M dividing to MA, MP distad origin of RP. MA basally crossvein-like, oblique (at obtuse angle); deeply forked at Psc, distal branch rather deeply forked again in both wings. MP deeply forked at Psc. Proximal crossvein between MA, MP (1im) very long. Crossvein 1m-cu slightly oblique; 2m-cu lost. CuA fused with MP for rather short distance; pectinate, with two simple branches arising from CuA proximad fusion with MP. CuP, A1, A2 fragmentarily preserved, apparently simple. Psc poorly developed, formed by MP, MA, basal branches of RP, connecting crossveins; continues to outer gradate series. Two gradate series of crossveins: inner series with eight crossveins distad MA; outer series with eight crossveins distad RP1.

Figures

FIGURE 8. Cimbrochrysa americana sp. nov., holotype UCM 33497 photographed using a polarized filter. Scale bars = 3 mm.

FIGURE 9. Wing venation drawings of Cimbrochrysa americana sp. nov., holotype UCM 33497. A, right forewing; B, left forewing; C, right hind wing; left hind wing. Scale bar = 3 mm (all to scale).