Cinnadenia paniculata (Hook. f.) Kosterm., 1973

De Kok, Rogier P. J. & Sengun, Seda, 2020, A revision of Cinnadenia Kosterm. (Lauraceae), Adansonia 42 (4), pp. 105-112 : 108-112

3. Cinnadenia paniculata (Hook. f.) Kosterm.

Adansonia, sér. 2, 13: 223 (1973); Grierson & Long, Flora of Bhutan 1 (2): 39 (1984). — Dodecadenia paniculata Hook.f., Flora of British India 5: 181 (1886); Kanjilal et al., Flora of Assam 4: 93 (1940).

— Type: Bhutan, Griffith 2472 (lecto-, K [ K000778754 ], here designated, Fig. 4 ; isolecto-, BM [ BM000951009 ]).

Tetranthera chartacea var. areolata Meisn. ex Hook. f., Flora of British India 5: 181 (1886).

— Type: Bhutan, Griffith 2472 (holo-, K[K000778754]).

Tetranthera chartacea sensu Meisn. , in DC, Prodromus Systematis Naturalis Regni Vegetabili 15: 186 (1864).

DISTRIBUTION. — North India (Assam, Sikkim) and Bhutan.

ECOLOGY. — Growing along the edges of broad-leaved forest, between 730 and 2000 m elevation. Flowering from October to March; fruiting from April to June.

CONSERVATION ASSESSMENT. — Least Concern. This species is known from an arch shaped area from Sikkim and Bhutan via Northern Assam to Nagaland and Manipur on the border between India and Myanmar. An analysis of the Extent of Occurrence (EOO) gives a IUCN Conservation Assessment of Least Concern, but the Area of Occupancy (AOO) suggests an assessment of Endangered. However, the Lauraceae tend to be under-collected and therefore a greater weight is given to the EOO. Hence, this species must be considered as Least Concern.

ADDITIONAL COLLECTIONS SEEN. — India: Assam, Naga Hills, Kohima, 30.XI.1935, Bar 42 ( K) ; Assam, Howai , King’s collector s.n. ( K, P [ P01749138 , P01749139 ]) ; Manipur, Imphal, XI.1907, Meebold 6726 ( K) ; Manipur, Lingi village, 5. I.1883, Watt 5008 ( K) ; Manipur, 28.X.1885, Clarke 43361 ( K) ; Manipur, 28.X.1885, Clarke 43360 ( K) ; Naga Hills, Reksima, 1.III.1935, Bar 2825 ( K) ; Sikkim, Hooker s.n. ( K [ K000778753 ], P [ P01749136 ]) ; Clarke 27565, Mik , 23.IV.1876 ( K [ K000778752 ]) .

Bhutan. Grierson & Long 4271, Tongsa district, near Pertimi, 3.IV.1982 ( K) ; Grierson & Long 1645, 2 km south of Shamgong , near Pertimi, 5.VI.1979 ( K) .

DESCRIPTION

Trees 10-30 m tall.Twigs slender, 1.6-3.2 mm thick, rounded to slightly angular in cross-section, velutinous when young, glabrescent; terminal leaf bud lanceolate, 2.3-2.5 mm long, apex acute, velutinous. Leaves alternate; leaf blade ovate-lanceolate, 11-22 × 4-7 cm, apex acute to indistinctly acuminate, base broadly cuneate, obtuse or almost rounded, leathery, pinninerved, secondary nerves 9-11 pairs, brochidodromous, tertiary nerves scalariform, thickly to thinly; upper surface glossy, glabrous, midrib sunken, secondary nerves sunken, tertiary nerves conspicuous, forming a distinct pitted surface (hand lens); lower surface pale green, glabrous, midrib and secondary nerves raised, tertiary nerves distinct; petiole half-terete, 9-20 mm long, slender, glabrous. Inflorescences panicles, 3-5 cm long, axillary, sparsely hairy, with 4 or 5 flowers, involucral bracts absent; bracts leaf like; bracteoles unknown, caducous. Flowers unisexual; perianth lobes 6, ovate to orbiculate, 2.6- 3.6 × 2.2-2.6 mm, apex acute, outside sparsely hairy, inside glabrous; stamens 9-12, 1.6-2.4 mm long, sparsely hairy; ovary 1.5-2.5 mm in diameter, glabrous; style c. 1 mm long, stigma peltate. Fruits 1-2 per infructescence, (dried) obovoid, 20-25 × 10-17 mm, apex rounded, glabrous, smooth, pink when cut; cupule fleshy, 4.7-8 mm diameter, margin entire, glabrous; stalk slightly swollen, up to 3 mm thick.

NOTES

Three gatherings are cited in the original description of Dodecadenia paniculata ( Hooker 1886) : J.D. Hooker s.n. from Sikkim, C.B. Clarke 27565 from Mik and Griffith 2472 from Bhutan. The Griffith specimen at K is the only one that has flowers, and as flowers are a more important character for species and genus recognition, it is selected here as the lectotype.

Figures

FIG. 4. — Lectotype of Cinnadenia paniculata (Hook. f.) Kosterm. (K000778754)